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JP5677839B2 - 突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物 - Google Patents

突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物 Download PDF

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JP5677839B2
JP5677839B2 JP2010515401A JP2010515401A JP5677839B2 JP 5677839 B2 JP5677839 B2 JP 5677839B2 JP 2010515401 A JP2010515401 A JP 2010515401A JP 2010515401 A JP2010515401 A JP 2010515401A JP 5677839 B2 JP5677839 B2 JP 5677839B2
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fatb
seq
nucleic acid
mutant
plant
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JP2010532667A5 (fr
JP2010532667A (ja
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ラハ,ベンヤミン
デン・ブール,バルト
ランベルト,バルト
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Bayer CropScience NV
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    • C12Q1/6888Nucleic acid products used in the analysis of nucleic acids, e.g. primers or probes for detection or identification of organisms
    • C12Q1/6895Nucleic acid products used in the analysis of nucleic acids, e.g. primers or probes for detection or identification of organisms for plants, fungi or algae
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    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01HNEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
    • A01H1/00Processes for modifying genotypes ; Plants characterised by associated natural traits
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    • AHUMAN NECESSITIES
    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
    • A01HNEW PLANTS OR NON-TRANSGENIC PROCESSES FOR OBTAINING THEM; PLANT REPRODUCTION BY TISSUE CULTURE TECHNIQUES
    • A01H5/00Angiosperms, i.e. flowering plants, characterised by their plant parts; Angiosperms characterised otherwise than by their botanic taxonomy
    • A01H5/10Seeds
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    • A01AGRICULTURE; FORESTRY; ANIMAL HUSBANDRY; HUNTING; TRAPPING; FISHING
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    • C12N15/63Introduction of foreign genetic material using vectors; Vectors; Use of hosts therefor; Regulation of expression
    • C12N15/79Vectors or expression systems specially adapted for eukaryotic hosts
    • C12N15/82Vectors or expression systems specially adapted for eukaryotic hosts for plant cells, e.g. plant artificial chromosomes (PACs)
    • C12N15/8241Phenotypically and genetically modified plants via recombinant DNA technology
    • C12N15/8242Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits
    • C12N15/8243Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits involving biosynthetic or metabolic pathways, i.e. metabolic engineering, e.g. nicotine, caffeine
    • C12N15/8247Phenotypically and genetically modified plants via recombinant DNA technology with non-agronomic quality (output) traits, e.g. for industrial processing; Value added, non-agronomic traits involving biosynthetic or metabolic pathways, i.e. metabolic engineering, e.g. nicotine, caffeine involving modified lipid metabolism, e.g. seed oil composition
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    • C12N9/14Hydrolases (3)
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  • Measuring Or Testing Involving Enzymes Or Micro-Organisms (AREA)
  • Breeding Of Plants And Reproduction By Means Of Culturing (AREA)
  • Enzymes And Modification Thereof (AREA)
  • Coloring Foods And Improving Nutritive Qualities (AREA)
  • Micro-Organisms Or Cultivation Processes Thereof (AREA)
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JP2010515401A 2007-07-09 2008-07-07 突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物 Expired - Fee Related JP5677839B2 (ja)

Applications Claiming Priority (5)

Application Number Priority Date Filing Date Title
EP07075568.1 2007-07-09
EP07075568 2007-07-09
US95894507P 2007-07-10 2007-07-10
US60/958,945 2007-07-10
PCT/EP2008/005551 WO2009007091A2 (fr) 2007-07-09 2008-07-07 Plante de la famille brassica comprenant des allèles mutants de l'acyle gras-acp thioestérase

Publications (3)

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JP2010532667A JP2010532667A (ja) 2010-10-14
JP2010532667A5 JP2010532667A5 (fr) 2011-08-25
JP5677839B2 true JP5677839B2 (ja) 2015-02-25

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JP2010515401A Expired - Fee Related JP5677839B2 (ja) 2007-07-09 2008-07-07 突然変異脂肪酸アシル−acpチオエステラーゼ対立遺伝子を含むアブラナ属植物

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US (2) US8981180B2 (fr)
EP (1) EP2167667B1 (fr)
JP (1) JP5677839B2 (fr)
CN (1) CN101688220B (fr)
AR (1) AR067480A1 (fr)
AU (1) AU2008274551B2 (fr)
CA (2) CA2692687C (fr)
DK (1) DK2167667T3 (fr)
EA (1) EA030862B1 (fr)
ES (1) ES2545083T3 (fr)
PL (1) PL2167667T3 (fr)
SI (1) SI2167667T1 (fr)
UA (1) UA99471C2 (fr)
WO (1) WO2009007091A2 (fr)
ZA (2) ZA200908886B (fr)

Families Citing this family (41)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
JP4786544B2 (ja) 2003-11-21 2011-10-05 ジボダン エス エー N置換p−メンタンカルボキサミド類
AR078740A1 (es) 2009-10-22 2011-11-30 Dow Agrosciences Llc Proteinas con dedos de zinc disenadas por ingenieria que tienen como objetivo genes vegetales comprendidos en la biosintesis de los acidos grasos
SI2501804T1 (sl) 2009-11-20 2016-09-30 Bayer Cropscience Nv Rastline Brassice, ki vsebujejo mutirane FAD3 alele
WO2011075716A1 (fr) 2009-12-18 2011-06-23 Cargill, Incorporated Plantes du genre brassica donnant des huiles à faible teneur totale en acides gras saturés
EP2525658B1 (fr) 2010-01-22 2017-03-01 Bayer Intellectual Property GmbH Combinaisons d'agents actifs acaricides et/ou insecticides
MX345064B (es) * 2010-05-12 2017-01-16 Agrigenetics Inc Uso de marcadores específicos de gen midrib-3 café en maíz para introgresión de rasgo.
US9695434B2 (en) * 2010-05-25 2017-07-04 Cargill, Incorporated Brassica plants yielding oils with a low alpha linolenic acid content
CA2837011C (fr) * 2010-05-25 2023-01-17 Cargill Incorporated Plantes du genre brassica donnant des huiles a faible teneur en acide alpha-linolenique
CN103282473A (zh) * 2010-11-03 2013-09-04 索拉兹米公司 具有降低倾点的微生物油、从其中产生的介电流体、以及相关方法
MX2013005258A (es) 2010-11-15 2013-07-05 Bayer Ip Gmbh N-aril pirazol(tio)carboxamidas.
WO2013020985A1 (fr) 2011-08-10 2013-02-14 Bayer Intellectual Property Gmbh Combinaisons de composés actifs comprenant des dérivés spécifiques d'acide tétramique
WO2014006162A1 (fr) 2012-07-06 2014-01-09 Bayer Cropscience Nv Plantes du genre brassica ayant une composition d'huile de graines modifiée
IN2014DN11215A (fr) 2012-07-06 2015-10-02 Bayer Cropscience Nv
CA2878287A1 (fr) 2012-07-06 2014-01-09 Bayer Cropscience Nv Sequences de genes rod1 provenant du soja et leurs utilisations
CN104995174A (zh) 2012-12-19 2015-10-21 拜耳作物科学股份公司 二氟甲基-烟酰-四氢萘基胺
US9567615B2 (en) 2013-01-29 2017-02-14 Terravia Holdings, Inc. Variant thioesterases and methods of use
US9816079B2 (en) 2013-01-29 2017-11-14 Terravia Holdings, Inc. Variant thioesterases and methods of use
KR20150128770A (ko) * 2013-03-15 2015-11-18 솔라짐, 인코포레이티드 티오에스테라아제 및 맞춤형 오일 생산용 세포
US9783836B2 (en) 2013-03-15 2017-10-10 Terravia Holdings, Inc. Thioesterases and cells for production of tailored oils
US9290749B2 (en) 2013-03-15 2016-03-22 Solazyme, Inc. Thioesterases and cells for production of tailored oils
BR112017001404A2 (pt) 2014-07-24 2017-11-21 Terravia Holdings Inc tioesterases variantes e métodos de utilização
EP3194582A2 (fr) 2014-09-18 2017-07-26 TerraVia Holdings, Inc. Acyl-acp thioestérases et leurs mutants
US10494642B2 (en) 2015-04-28 2019-12-03 Basf Agricultural Solutions Seed, Us Llc Brassica plants with modified seed oil composition
WO2017205665A1 (fr) * 2016-05-25 2017-11-30 Cargill, Incorporated Nucléases modifiées pour générer des mutants de délétion dans des plantes
CN109640679A (zh) 2016-08-12 2019-04-16 嘉吉公司 特种低饱和物菜籽油
US10440915B2 (en) 2016-09-01 2019-10-15 Cargill, Incorporated Canola cultivar 15RH0611
US10440916B2 (en) 2016-09-06 2019-10-15 Cargill, Incorporated Canola cultivar 15RH0612
US10440909B2 (en) 2016-09-06 2019-10-15 Cargill, Incorporated Canola cultivar 15RH0613
TWI873082B (zh) * 2017-08-17 2025-02-21 日商莎卡達種子股份有限公司 甘藍(Brassica oleracea)植物之異型檢出法
US20220033835A1 (en) * 2018-09-14 2022-02-03 Chuanxin Sun Rice plant material resistant against biotic stress
CN113166776A (zh) * 2019-01-04 2021-07-23 嘉吉公司 用于在植物中产生突变的工程化核酸酶
CA3047768A1 (fr) 2019-06-21 2020-12-21 BASF Agricultural Solutions Seed US LLC Variete de canola hybride 7cn0425
US20230062896A1 (en) * 2020-01-31 2023-03-02 Calyxt, Inc. Increase of saturated fat in soybean
US11997966B2 (en) 2020-04-24 2024-06-04 BASF Agricultural Solutions Seed US LLC Canola hybrid variety 7CN0020
US11672218B2 (en) 2020-04-24 2023-06-13 BASF Agricultural Solutions Seed US LLC Canola hybrid variety 7CN0065
US11997965B2 (en) 2020-04-24 2024-06-04 BASF Agricultural Solutions Seed US LLC Canola hybrid variety 8CN0001
US20240292798A1 (en) 2020-12-21 2024-09-05 BASF Agricultural Solutions Seed US LLC Brassica napus plants comprising an improved fertility restorer
TW202342756A (zh) 2022-03-01 2023-11-01 美商巴斯夫農業解決方案種子美國有限責任公司 Cas12a切口酶
CN114958880B (zh) * 2022-04-06 2023-08-29 吉林大学 大豆脂酰-酰基载体蛋白硫酯酶GmFATA2基因及应用
US20250243482A1 (en) 2023-08-31 2025-07-31 BASF Agricultural Solutions Seed US LLC Class 2 type v crispr-cas prime editing
WO2025083165A1 (fr) 2023-10-19 2025-04-24 Basf Agricultural Solutions Us Llc Plantes brassica napus présentant une résistance à la maladie du pied noir améliorée

Family Cites Families (16)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
EP0476093B1 (fr) * 1990-04-04 1997-05-07 Pioneer Hi-Bred International, Inc. Production de graines de colza ameliorees presentant une teneur reduite en acides gras satures
US5344771A (en) 1990-04-26 1994-09-06 Calgene, Inc. Plant thiosterases
US5512482A (en) 1990-04-26 1996-04-30 Calgene, Inc. Plant thioesterases
DK0563191T4 (da) 1990-12-20 2000-07-17 Du Pont Nucleotidsekvenser af sojabønne-acyl-ACP-thioesterase-gener
US5945585A (en) 1990-12-20 1999-08-31 E. I. Du Pont De Nemours And Company Specific for palmitoyl, stearoyl and oleoyl-alp thioesters nucleic acid fragments encoding acyl-acp thiosesterase enzymes and the use of these fragments in altering plant oil composition
US5455167A (en) 1991-05-21 1995-10-03 Calgene Inc. Medium-chain thioesterases in plants
DK0534858T4 (da) 1991-09-24 2005-08-08 Keygene Nv Selektiv restriktionsfragmentamplifikation: en general fremgangsmåde til DNA-fingeraftryk-dannelse
US5850022A (en) 1992-10-30 1998-12-15 Calgene, Inc. Production of myristate in plant cells
CA2176137A1 (fr) 1993-11-10 1995-05-18 Alois Toni Voelker Sequences de thioesterases vegetales actives sur les substrats acyl-(proteine vectrice d'acyle)
JPH10505237A (ja) 1994-08-31 1998-05-26 イー・アイ・デユポン・ドウ・ヌムール・アンド・カンパニー カノラ及びダイズのパルミトイル−acpチオエステラーゼ遺伝子のヌクレオチド配列、並びにダイズ及びカノラ植物の油の脂肪酸量の調節におけるこれらの使用
US5625130A (en) * 1995-03-07 1997-04-29 Pioneer Hi-Bred International, Inc. Oilseed Brassica bearing an endogenous oil wherein the levels of oleic, alpha-linolenic, and saturated fatty acids are simultaneously provided in an atypical highly beneficial distribution via genetic control
US5985557A (en) 1996-01-24 1999-11-16 Third Wave Technologies, Inc. Invasive cleavage of nucleic acids
US7067722B2 (en) * 1999-08-26 2006-06-27 Monsanto Technology Llc Nucleic acid sequences and methods of use for the production of plants with modified polyunsaturated fatty acids
US6706950B2 (en) 2000-07-25 2004-03-16 Calgene Llc Nucleic acid sequences encoding β-ketoacyl-ACP synthase and uses thereof
CA2490771A1 (fr) * 2002-06-21 2003-12-31 Monsanto Technology Llc Sequences d'acides nucleiques associees a la thioesterase et methodes d'utilisation pour la production de plantes presentant une teneur en acides gras modifiee
AU2005294298C1 (en) 2004-10-08 2017-03-09 Corteva Agriscience Llc Oil and seeds with reduced saturate levels of fatty acids

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EA030862B1 (ru) 2018-10-31
EA201070114A1 (ru) 2010-08-30
AR067480A1 (es) 2009-10-14
US8981180B2 (en) 2015-03-17
ES2545083T3 (es) 2015-09-08
CA2692687C (fr) 2021-10-26
AU2008274551B2 (en) 2014-05-29
EP2167667B1 (fr) 2015-05-20
ZA200908886B (en) 2011-02-23
WO2009007091A2 (fr) 2009-01-15
CA3128584A1 (fr) 2009-01-15
US10837066B2 (en) 2020-11-17
AU2008274551A1 (en) 2009-01-15
CN101688220B (zh) 2017-04-05
ZA201009002B (en) 2011-07-27
CA2692687A1 (fr) 2009-01-15
CN101688220A (zh) 2010-03-31
UA99471C2 (ru) 2012-08-27
EP2167667A2 (fr) 2010-03-31
WO2009007091A3 (fr) 2009-07-23
SI2167667T1 (sl) 2016-03-31
US20110145944A1 (en) 2011-06-16
PL2167667T3 (pl) 2015-10-30
DK2167667T3 (en) 2015-07-27
JP2010532667A (ja) 2010-10-14
US20150232951A1 (en) 2015-08-20

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