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Understanding Species Diversity

This document discusses methods for measuring species diversity in ecological communities. It defines species richness as the number of different species and species abundance as the relative abundance of species. Common diversity indices that combine both richness and abundance include the Shannon-Wiener Index and Simpson's Index. The Shannon-Wiener Index ranges from 0 to 5 and is relatively easy to calculate but not always sensitive to actual differences between sites. Simpson's Index measures dominance and gives the probability that any two randomly selected individuals will belong to different species. Similarity indices can also be used to compare species composition between communities under different management over time.

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Goutam Das
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475 views3 pages

Understanding Species Diversity

This document discusses methods for measuring species diversity in ecological communities. It defines species richness as the number of different species and species abundance as the relative abundance of species. Common diversity indices that combine both richness and abundance include the Shannon-Wiener Index and Simpson's Index. The Shannon-Wiener Index ranges from 0 to 5 and is relatively easy to calculate but not always sensitive to actual differences between sites. Simpson's Index measures dominance and gives the probability that any two randomly selected individuals will belong to different species. Similarity indices can also be used to compare species composition between communities under different management over time.

Uploaded by

Goutam Das
Copyright
© © All Rights Reserved
We take content rights seriously. If you suspect this is your content, claim it here.
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Download as DOCX, PDF, TXT or read online on Scribd
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Measuring Species Diversity

1. Definition.
a. Difficult to define because defining consists of two distinct components:
i. Species Richness = Variety of species or the number of different species (or
genera, families, etc).
ii. Species Abundance = Relative abundance of species
b. Species Richness = an index based on the number of species
i. Numerical species richness = number of species per specified number of
individual
ii. Species density = number of species per unit are
iii. Simple and easy to calculate and therefore intuitively appealing.
iv. However, because it does not account for relative abundance, it is often not
sensitive to environment disturbance.
c. Species Abundance or Evenness = Describes relative abundance of species.

2. Benefits of Diverse Range Plant Community:


a. A mixture of plants provides forage for a variety of insect and vertibratel species
(biodiversity)
b. A mixture of plants will contain some plants that can survive drought, insect plagues,
and/or disease outbreaks so that the site will have some soil protection/forage/etc. in
those years
c. A mixture of plants contains a variety of genetic material that may be useful in long-
term survival.
d. The community benefits from a mixture of plants (soils improve with nitrogen fixers,
deep rooted plants bring nutrients up from soil layers below other plants roots, some
species work together so that both can survive – commensalism)
e. Healthy diverse plant communities generally have all niches filled and are
theoretically less likely to be invaded by noxious or opportunistic introduced species.

Disadvantages of Diverse Composition in the Range Plant Community:

f. It becomes more difficult to manage because different species of plants have different
grazing tolerances and different rates of phonological development.
g. Not all plant species are desirable for all foraging species.
h. Diverse communities are often a sign of fragmented or somewhat degraded site where
much of species richness is contributed by disturbance species.

Diversity Indices – that combine both richness and abundance

i. Shannon – Wiener Index (H’)

i.Most commonly used index in ecological studies


ii.Values range from 0 to 5, usually ranging from 1.5 to 3.5

iii.Calculated: H´= - ∑[ (
iv.Where ni=number of individuals or mount (e.g.biomass) of each species (the ith
specices ) and N=total number of individuals (or amount) for the site,and In= the natural log of
the number.

v.Advantages and Disadvantages:

1) Relatively easy to calculate

2) Fairly sensitive to actual site differences

3) There are several instances where H’ is similar between sites even though sits are
different.

j.Simpson’s Index(λ)-

i. λ is a measure of dominance therefore,(1- λ) measures species diversity.

ii.Gives the probability that any two individuals drawn at random from an infinitely
large community belong the different species.

n(n-1)
iii..Calculate:λ=∑
N(N-1)

iv.When ni =number of individuals or amount of each species (i.e. the number of


individuals of the ith species) and N=total number of individuals for the site.

v.Advantages and Disadvantages

1) Less sensitive to species richness and heavily weighted towards the most
abundant species.
2) Generally less sensitive than Shannon-Weiner H’

3.Similarly Indices-measure similarity between communities based on species composition.

i. Useful in comparing communities under different management.


ii. Can also be used to compare composition changes over time(e.g. determine how
similar the communities is to what is was 10 years ago…if data exist)
iii. Similarly indices overcome some short comings such as:
1) When communities are very different comparing diversity indices may not be
meaningful
2) Diversity indices often don’t change for sites with similar species but different
proportion.

iv.Similarity Index-

1) Values ranges from 0-1 with the higher value suggesting greater similarity

2) Calculate: Sim= 2∑ nc
∑n1+∑n2
3) Where nc =the common species between sites ;n1= the species of site 1 and n2

4) When cover is being used, similarity is simply ∑ cover in common for each spp.

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