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Psilophyta To Sphenophyta

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The key takeaways are that seedless vascular plants include ferns, club mosses and horsetails. They transport water through vascular tissue and reproduce via spores rather than seeds. Most are homosporous but some are heterosporous.

The main characteristics of seedless vascular plants are that they have true roots, stems and leaves but reproduce via spores rather than seeds. They transport water through vascular tissue and are limited to moist environments as their sperm require water.

Lycophyta, or club mosses, have needle-like or scale-like leaves and reproduce sexually underground. Sphenophyta, or horsetails, have jointed stems and leaves arranged in whorls. They reproduce similarly to lycophytes but horsetails were once larger tree-sized plants.

Seedless Vascular Plants

Comparing Spore-Bearing Vascular Plants

Characteristics Lycophyta Sphenophyta Pterophyta


Common Club Mosses Horsetails Ferns
Organisms
Water By vascular By vascular By vascular
transportation tissue tissue tissue

Structure Look like miniature True leaves, Creeping or


pine stems, and roots underground
trees; scalelike rhizomes (stems);
leaves fronts (leaves);
some have no
roots or leaves
Seedless vascular plants
General Characteristics

• Seedless vascular plants include ferns, whisk


ferns, club mosses, and horsetails.
• The plants do not produce seeds so, like
bryophytes, they are dispersed (spread) by
windblown spores.
General Characteristics
• Most species are homosporous, meaning they
produce only one type of spore.
• The gametophyte and sporophyte are
independent.
• They can produce a separate male or female
gametophyte or bisexual gametophytes, the
condition is known as homospory
General Characteristics
• Some are heterosporous, can produce two
types of spores megaspore and microspore
• Macrosporangia produces megaspore and
microsporangia produces microspore.
• Megaspore produces female gametophytes
and microspore produces male gametophytes
Seedless vascular plants

• They are vascular plants and therefore have


true roots, stems, and leaves.
• The sperm are flagellated and require water
for reproduction. These plants are therefore
limited to moist areas.
Seedless vascular plants
• Many of the seedless vascular plants were
once tree-sized.
• During the carboniferous period (near the end
of the Paleozoic), these plants were so
abundant that in some areas, their remains
accumulated faster than they decomposed.
These accumulations produced our fossil
fuels.
Seedless vascular plants

• The earliest known vascular plants had a


pattern of branching that increased the
number of sporangia. 
• Leaves of later plants probably evolved from
webbing between the branches.
Phylumm Psilophyta
• Consits of 142 species
• Most species is
commonly known as
Whisk ferns
• Group consisting of
two extant genera,
Psilotum and
Tmesipteris, in one
family, Psilotaceae.
Phylumm Psilophyta
• These plants are
primitive in
structure: Psilotum
lacks both roots
and leaves and is Rhynia
structurally similar - one of earliest
to the fossil genus vascular plants
(ca. 400 million years
Rhynia. ago)
- lacked roots
Phylumm Psilophyta
• Recent molecular systematic studies suggest
that the family is actually related to primitive
ferns.
• Tmesipteris has a more complex morpholgy in
that it has structures on the aerial shoot that
are foliar.
Phylumm Psilophyta
• Both Psilotum and
Tmesipteris have
compound sporangia
called synangia. In the
case of Psilotum these
are three-parted.
Phylumm Psilophyta
Phylumm Psilophyta
• While Psilotum lacks true
leaves, it possesses leaf-
like extentions of the
stem called enations.
• Because these lack
vasculature, they are not
considered leaves.
However, in Psilotum
complanatum, a vascular
trace occurs below the
enations. The foliar
structures of Tmesipteris
are vascularized.
Phylumm Psilophyta
• The gametophytes of
both genera are non-
photosynthetic and live
in association with a
fungus.
• In the case of Psilotum,
the gametophyte of
certain strains produce
vascular tissue.
Phylumm Psilophyta
• While both genera have
aerial branches arising
from stems embedded
in its substrate, they
both lack roots. The
rhizomes are infected
with mycorrhizae.
Phylum Lycophyta
• Ancient Lycophytes
• Appeared 390 million years ago
(mya)
• Grew to 30 m tall
• Extremely abundant due to the Lycopodium
moist warm environment
• Most died out 280 may due to a
new drier cooler environment
• include Lycopodium and
Selaginella

Selaginella
Phylum Lycophyta
Modern Lycophytes
• Much smaller than ancestors
• Grow close to the ground
• Found mainly in moist/damp forests
• Can be found in deserts and mountains though
• AKA the Club mosses and Spike mosses
• because they look like the moss gametophytes—
but they are NOT MOSSES!
• Sporophyte generation is dominant
Lycophyta Leaves and
reproduction?
• Leaves protect the reproductive cells
• Leaves occur in spirals, whorls, pairs
• Leaves form clusters called STROBILUS at the end
of stems.
Lycophyte Reproduction/Life Cycle:

• Sporangium burst and release spores


• Prothallus: gametophytes formed from
spores; relatively small; lives in or on the soil;
form both archegonia and antheridia;
• In some lycophytes, 2 types of spores form
Small spores: become male prothallus
– Form antheridium
Lycophyte Reproduction/Life Cycle:
• Large spores: become female prothallus
– Form archegonium
• Sperm from the antheridiumswim through a
film of water on the prothallus to the egg in
an archegonium and fertilize the egg.
• Then a sporophyte plant grows from the
zygote.
Lycopodium
• genus of clubmosses, also known
as ground pines, in the family
Lycopodiaceae, a family of fern-
allies (see Pteridophyta).

• They are flowerless, vascular,


terrestrial or epiphytic plants, with
widely-branched, erect, prostrate
or creeping stems, with small,
simple, needle-like or scale-like
leaves that cover the stem and
branches thickly.
Lycopodium
• The fertile leaves are arranged
in cone-like strobili.
• Specialized leaves (sporophylls)
bear reniform spore-cases (
sporangia) in the axils, which
contain spores of one kind only.
These club-shaped capsules give
the genus its name.
Lycopodium
• Lycopods reproduce sexually by spores.
The plant has an underground sexual
phase that produces gametes, and this
alternates in the life cycle with the
spore-producing plant.
• The prothallium developed from the
spore is a subterranean mass of tissue of
considerable size and bears both the
male and female organs (antheridium
and archegonia).
• However, it is more common that they
are distributed vegetatively through
above or below ground rhizomes.
Lycopodium life cycle
Selaginella
• Spike Mosses
• Especially abundant in tropics.
• Branch more freely than ground
pines.
• Leaves have a ligule on upper
surface.
• Produce two different kinds of spores
and gametophytes (heterospory).
Selaginella life cycle
Strobili
Selaginella

Cross section of the Selaginella stem


Selaginella

Cross section of strobili


Selaginella

Microsporangium and Megasporangia


Selaginella

Cross section of strobili


Phylum Sphenophyta
• Horsetails or Scouring rushes
• Look like horsetails and contain silica
that helped to scour dishes and
utensils.
• Ancient members were tree sized,
today, they grow only to a maximum of
1 m.
Equisetum
• Sphenophytes have jointed stems
Phylum Sphenophyta
• Leaves form strobillus at the tips
of some stems
• Most grow in marshes, stream
banks; damp soil
• Some grow in fields, roadsides; in
drier areas
• Reproduction very similar to
lycophytes.
Phylum Sphenophyta
• The plant is essentially stem, it has a rhizome
which puts out adventitious roots.
• The leaves are a whorl of non-photosynthetic
scales at each node. Some species produce lots of
feathery branches.
• Their cell walls contain silica which makes the
stems coarse textured, and led to their use as a
natural scouring pad for cook ware.
• Spores are produced in strobili and although the
plant is homosporous the gametophytes are
unisexual.
Phylum Sphenophyta
Equisetum life cycle
Thank you

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