Egyptian Journal of Aquatic Biology & Fisheries
Zoology Department, Faculty of Science,
Ain Shams University, Cairo, Egypt.
ISSN 1110 – 6131
Vol. 24(7): 441 – 452 (2020)
www.ejabf.journals.ekb.eg
An evaluation for the exploitation level of Egyptian Marine Fisheries
Shimaa I. Maiyza 1*; Sahar F. Mehanna2; Ibrahim A. El-karyoney3
1.National Institute of Oceanography & Fisheries (NIOF), Red Sea branch, Egypt.
2. National Institute of Oceanography & Fisheries (NIOF), Suez and Aqaba Gulfs branch, Egypt.
3. National Institute of Oceanography and Fisheries (NIOF), Alexandria branch, Egypt.
*
Corresponding Author: shemo_iam@yahoo.com
__________________________________________________________________________
ARTICLE INFO
ABSTRACT
Article History:
Received: April 26, 2020
Accepted: Nov. 1, 2020
Online: Nov. 3, 2020
_______________
Egyptian marine fisheries contribute to the economy of the country
where it provides an important source for food and employment. There are
many challenges that affected the sustainability of marine fish stocks from
which overfishing due to the increasing fishing effort. In the present study,
annual catch and effort data were analyzed to estimate the maximum
Keywords:
sustainable yield (MSY) and the corresponding level of fishing effort (fmax)
Fish Stock assessment,
for both the Mediterranean & Red seas, Egypt. According to the Schaefer
Surplus Production
model (1954), the value of MSY was estimated at 73.0*103 tons for
Models,
Mediterranean sea fisheries, and at 63.1*103 tons for Red sea fisheries. These
Overfishing,
values of MSY were obtained at fishing effort (fmax) of 3.5*103 &
Egyptian marine fisheries, 1.6*103 standard boats in Mediterranean & Red seas respectively. The results
Maximum Sustainable
indicated that the optimum exploitation in Egyptian marine fisheries required
Yield.
an urgent reduction in fishing effort. These results confirmed the
overexploitation situation of the marine fish stocks in Egypt.
INTRODUCTION
The marine fishery resource provides important source for food and has been
traditionally exploited for centuries in Egypt. With the rapid increase in the human
population over the years, the demand for fish has increased considerably. During the last
years there has been a growing awareness among the users about the need to conserve
and manage marine fisheries resources based on data collected in a scientific manner so
as to ensure proper utilization and maximum sustainable yield (MSY). Egypt has a long
coastline on both Mediterranea (1100 Km) and Red seas (1080 Km). Egyptian marine
fisheries (Fig. 1) has a rich and diverse commercial species varied from demersals,
semipelagics, benthopelagics and pelagics. Over exploitation on the Egyptian coastal
resources has caused decline of many fish, mollusc and crustacean stocks. Stock
assessment of fishery is important for its effective management and the effective
management strategy which based on a sound scientific basis can help to obtain
sustainable yield every year (Mehanna and El-Gammal, 2007; Osman, 2015; El-Mahdy,
2017; Ali, 2017; Hassanein, 2017; Mehanna, 2020; Mohamed et al., 2020). The state of a
442
Shimaa I. Maiyza et al., 2020
fishery can be assessed by many mathematical models. Catch and effort statistics
constitute the minimum requirements for the assessment of a fishery. Effort and catch
per-unit effort statistics provide indices that are related to important quantities, such as
the fishing mortality and the density of the exploited stock (Gulland, 1983). Surplus
production models (SPMs) also known as biomass dynamics models (BDMs) are
frequently used to evaluate MSY and to study the harvested population. These models
need catch and effort data which is valued when age structured data are absent (Haddon,
2011). Surplus production models are used to determine the optimum level of effort
which produces the maximum yield that can be sustained without affecting the long-term
productivity of the stock, the so-called maximum sustainable yield (MSY). A number of
studies were undertaken to estimate MSY of the marine fishery resources in Egypt based
on SPMs (Mehanna, 2002, 2005, 2009; Mehanna and El-Gammal, 2007; Mehanna and
Haggag, 2010; Mehanna et al., 2020).
The present study analysed the catch and effort data of Mediterranean and Red
seas fish stocks to estimate the maximum sustainable yield (MSY) and the corresponding
level of fishing effort (fmax) using Schaefer model (1954). The results of Schaefer (1954)
model provide baseline information on maximum sustainable yield for the Egyptian
marine waters, which could be useful for any future fishery assessment.
MATERIALS AND METHODS
A time series of catch and effort data for the period from 1995 to 2016 (22 years)
were collected from two sources; the Central Agency of Public Mobilization and
Fisheries Statistics (CAPMS) and General Authority for Fish Resources Development
(GAFRD). Fishing effort is expressed by the number of operational fishing boats in the
Egyptian marine waters of both Mediterranean and Red Sea. As the Egyptian marine
fisheries is of multi-fleet nature, the fishing boats of different characteristics were
standardized.
The surplus production model of Schaefer (1954) was applied in this study to the
per annum total catch in tons and standardized number of fishing units. This model
considered the stock as one big unit of biomass.
Schaefer model (1954) is one of Surplus production models, which can be used
to estimate the maximum sustainable yield (MSY) and the corresponding level of fishing
effort (fmax), expresses the catch per unit effort (CPUE) as a function of effort as follows:
Y /f = a + b f
Where: Y: yield (catch in weight),
f: effort,
and Y/f: yield (weighted catch) per unit of effort.
By fitting the effort against catch per unit effort via ordinary least square method (OLS),
the obtained parameters a and b can used to estimate the MSY and fMSY using the
following formulae:
MSY = - a2 / 4b
fmsy = - a / 2b
443
An evaluation for the exploitation level of Egyptian Marine Fisheries
The slope (b) will be negative if the catch per unit effort (Y/f) decrease with the
increase of effort (f) and vise versa.
Fig.1. Egyptian Marine fisheries (Mediterranean and Red Seas coasts)
RESULTS AND DISCUSSION
1. Egyptian marine fisheries
Egypt drives its fish production from two resources; natural resources (marine and
inland waters), and artificial resources or aquaculture. Egyptian Marine fisheries include
two seas, Mediterranean sea in the north and Red sea in east of Egypt (Figs. 1&2).
Egyptian Marine fisheries one of the most important sources for fish production in
Egypt, and represented the largest area compared to other fish resources. The actual area
of these fisheries are approximately about 11.2*106 acre during the period of study
(1995-2016), which represented about 84.8% of natural fisheries area and 80.5% of
Egyptian fisheries area. While exploitable area of these fisheries is only about 31.8%,
30.2% from natural fisheries area and Egyptian fisheries area consequently (Table 1).
The Mediterranean Sea fisheries area varies between actual and exploitable areas
which are about 6.8*106 and 3.3*106 acre, whereas the Red Sea fisheries these areas are
4.4*106 acre and 0.9*106 acre for actual and exploitable areas respectively. As the area of
Mediterranean Sea fisheries is greater than that of Red Sea fisheries, its contribution to
the Egyptian natural fisheries is higher (Fig. 2). On the other hand, the coast length of
Egyptian Marine fisheries is approximately about 2.4*103 Km, divided to about 1.1*103
444
Shimaa I. Maiyza et al., 2020
Km (46%) for Mediterranean Sea fisheries and up to 1.3*103 Km (54%) for Red sea
fisheries with its two Gulfs (Table 1).
Table 1. Relative actual and exploitable area of Egyptian marine fisheries during 19952016.
Items
Egyptian Marine fisheries
Mediterranean Sea
Red Sea
The total
A
E
A
E
A
E
6.8
3.3
4.4
0.9
11.2
4.2
Area (acre*106)
Egyptian marine
60.7
78.6
39.3
21.4
fisheries area
(%)
Natural fisheries
51.5
25.0
33.3
6.8
area (%)
Egyptian
48.9
23.7
31.6
6.5
fisheries area
(%)
Coast length
1100
1300
(Km)
Egyptian marine
fisheries coast
46
54
length
(%)
A: Actual area for fishing
E: Exploitable area for catching.
Source: Maiyza, 2015 and Google Earth program.
Natural
fisheries
Egyptian
fisheries
13.2
13.9
100.0
100.0
ــــــــــ
ــــــــــ
84.8
31.8
100.0
ــــــــــ
80.5
30.2
ــــــــــ
100.0
2400
ــــــــــ
ــــــــــ
100.0
ــــــــــ
ــــــــــ
Fig. 2. Contribution of Egyptian marine fisheries in fish production during 2018
From the above illustrated table and graph, Red Sea fisheries is longer in coast
length and smaller in its area than in Mediterranean Sea fisheries because of Red Sea
coast has many meanders compared to Mediterranean Sea. Consequently, the
contribution of Mediterranean fisheries to the fish production in Egypt is more than the
contribution of Red Sea fisheries.
445
An evaluation for the exploitation level of Egyptian Marine Fisheries
It is worth mentioning that the marine fisheries in Egypt were contributed by 25%
of total fish production in 80’s, severly decreased to only 5% in 2018 (Mehanna, 2020;
GAFRD, 2018).
2. Catch and effort statistics
The total fish catch in the Egyptian Marine fisheries during the period of the study
is about 119.3*103 tons, which represents about 31.37%, 13.3%, from Natural fisheries
production, and total Egyptian fish production consequently, (Table 2). The relative
importance of the Mediterranean Sea fisheries is higher than The Red Sea fisheries in its
contribution to the Egyptian Marine fisheries, natural fisheries and Egyptian fisheries in
its fish production and area.
Table 2. Catch in Egyptian marine fisheries during 1995-2016.
Marine fisheries
(seas)
Mediterranean Sea
Red Sea
Egyptian marine fisheries
Natural fisheries
Total Egyptian fish
production (tons*103)
64.4
54.9
119.3
380.2
% from
Egyptian
Marine
fisheries
54.0
46.0
100.0
ـــــــ
897.1
ـــــــ
Average
Production
(tons*103)
% from
Natural fisheries
production
% from
total Egyptian fish
production
16.9
14.4
31.3
100.0
7.2
6.1
13.3
42.4
ـــــــ
100.0
Source: CAPMAS, 2016 and GAFRD, 2016.
There are 10 governorates shared the Egyptian marine fisheries, 7 governorates
are lie on Mediterranean Sea and shared its fisheries; arranged according to its
productivity as Damietta, Beheira, Port Said, Alexandria, Kafr El-shiekh, North Sinai and
Matrouh. Whereas three governorates are shared the Red Sea fisheries, which are Suez,
Red Sea and South Sinai (Table 3).
Table 3. Catch and effort of different governorates shared the Egyptian marine fisheries
during 1995 – 2016.
Red Sea
Mediterranean Sea
Egyptian Marine fisheries
Damietta
Beheira
Port Said
Alexandria
Kafr El-shiekh
North Sinai
Matrouh
Total
% of Totals
Suez
Red Sea
South Sinai
total
% of Totals
Totals
Production
(tons*103)
19.5
13.0
12.2
11.6
5.8
2.9
0.4
65.4
53.8%
29.1
26.1
1.0
56.2
46.2%
121.6
Source: CAPMAS, 2016 and GAFRD, 2016.
%
30.0
19.8
18.6
17.7
8.9
4.4
0.6
100.0
51.8
46.4
1.8
100.0
Motor
boat
830
617
408
702
199
175
10
2941
70.5%
550
602
79
1231
29.5%
4172
Effort
Total
Sailing
boat
(boat)
%
242
1072
22.9
293
910
19.5
494
902
19.3
395
1097
23.4
216
415
8.9
54
229
4.9
41
51
1.1
1735
4676
100.0
66.7%
69.0%
341
891
42.5
456
1058
50.5
68
147
7.0
865
2096
100.0
33.3%
31.0%
2600
6772
446
Shimaa I. Maiyza et al., 2020
Fish catch of Egyptian marine fisheries during 1995-2016 is about 119.3*103 tons
with percentage contribution of 54% for Mediterranean fisheries and 46% from Egyptian
marine fisheries. Generally there is a decreasing trend in the catch of marine fisheries
during the period of study reflecting the decreasing of productivity and relative
abundance of these fisheries (Fig. 3).
Fig. 3. Trend of fish production from Egyptian marine fisheries during 1995–2016
The fishing effort in Mediterranean and Red Sea fisheries represented by fishing
boats are contributed about 69% and 31% of fishing effort in Egyptian marine fisheries
respectively, which is estimated at about 6.8*103 boats during the period of the study.
It is noted that the fishing effort represented by the number of fishing vessels in
Mediterranean Sea fisheries is higher than Red Sea fisheries; also motor boats are bigger
than sailing boats in Egyptian marine fisheries in general (Table 3).
3. Egyptian marine fisheries management
Fish stock assessment is very important process because it determines: (1)
fisheries productivity, (2) the impact of fishing process on fisheries, (3) the effect of
changing fishing rates. So it is essential to ensure the fisheries management and optimum
exploitation (Musick & Bonfil, 2005) by:
- Identify the level of exploitation that gives maximum weighted production in the long
term from under study fisheries.
- Fish stocks assessment studies must be direct to fishermen and decision-makers in
fisheries management, because these studies answer to number of questions: when and
where? What catch? , as well as what is size supposed to catch? and how to fish (fishing
methods and tools).
To manage Egyptian marine fisheries and to evaluate its current exploitation
situation, the maximum sustainable yield (MSY) and the corresponding level of fishing
effort (fmax) for catch are estimated using the surplus production models. These models
deal with the entire stock, the entire fishing effort and the total yield obtained from the
447
An evaluation for the exploitation level of Egyptian Marine Fisheries
stock, without details about growth and mortality parameters or the effect of mesh size
on the age of fish capture .
To apply Schaefer model on Egyptian marine fisheries, the catch and catch per
unit fishing effort expressed as ton/standard fishing unit during 22 years (1995-2016) are
used.
3.1. MSY and fmax for Mediterranean Sea fisheries
Schaefer model has been applied on Mediterranean sea fisheries during the
period from 1995 to 2016 to estimate the maximum sustainable yield (MSY) and the
corresponding level of fishing effort (fmax) for fish catch (Figs.4 & 5). The estimated
MSY was 73.0*103 ton at fishing effort of 3.5*103 standard boats. These results
confirmed the over exploitation situation in the Egyptian Mediterranean fisheries (Table
4) and coincide with the previous studies dealing with the assessing the fish stocks in
Mediterranean (eg. Mehanna, 2009, 2014, 2019a&b; Mehanna and Haggag, 2010&2011;
Mehanna et al., 2011; El-Serafy et al., 2015; Hassanein, 2017).
tons
25.0
Y/F = 41.536 - 0.0059 f
R² = 0.6456
CPUE
20.0
15.0
10.0
5.0
0.0
4000
4500
5000
Effort
f
5500
6000
boa
Fig. 4. Relationship between CPUE and effort in Mediterranean Sea fisheries during 1995-2016
Fig. 5. Yield Curve of Mediterranean Sea fisheries according to Schaefer Model
448
Shimaa I. Maiyza et al., 2020
Table 4. MSY and fmax estimated via Schaefer model of Egyptian Marine fisheries during
1995-2016.
Mediterranean
Sea
Egyptian marine
fisheries
Items
Results
Schaefer Model
MSY
(tons*103)
fmax
(boats)
Overproduction %increasing
Y = 41.536 f ± 0.0059 f2
73.0
3500
Effort
%drcreasing
25.1
Red Sea
Schaefer Model
MSY
(tons*103)
Fmax
(boats)
Overproduction %increasing
Effort
13.6
Y = 76.21 ± 0.0234 f2
63.1
1628
14.9
%drcreasing
22.3
3.2. MSY and fmax for Red Sea fisheries
Schaefer model has been applied on Red Sea fisheries also to estimate the
maximum sustainable yield (MSY) and the corresponding level of fishing effort (fmax)
(Figs. 6 & 7). It is obvious that the value of MSY is 63.1*103 ton, whereas the value of
fmax is 1.6*103 boats. The overfishing situation was observed (Table 4) and agreed with
all previous studies dealing with tstock assessment of Red Sea fisheries (eg. Mehanna and
El-Gammal, 2007; Mehanna, 2005 to 2020; Tesfamichael and Mehanna, 2012; Osman,
2015; Mohamed, 2016; El-Mahdy, 2017; Mehanna et al., 2016, 2017, 2018, 2019;
Mohamed et al., 2020).
CPUE
ton/boat
50.0
45.0
40.0
35.0
30.0
25.0
20.0
15.0
10.0
5.0
0.0
1500
Y/f = 76.21-0.0234 f
R² = 0.6697
boat
1700
1900
2100
Effort
f
2300
2500
Fig. 6. Relationship between CPUE and effort in Red sea fisheries during 1995-2016.
449
An evaluation for the exploitation level of Egyptian Marine Fisheries
Fig. 7. Yield Curve of Red sea fisheries according to Schaefer Model
In spite of the large area of marine fisheries in Egypt, these fisheries are not
exploited rationally due to the decrease of relative abundance of its fish stocks as a result
of different reasons such as pollution (domestic and industrial sewages), over fishing and
illegal fishing, habitat degradation, human activities on coastal areas, in addition to
climate changes. These challenges especially the increasing of fishing effort had affected
the sustainability of fish production from Egyptian marine fisheries (Mediterranean Sea
& Red Sea fisheries). This situation was proved by applying Schaefer model (1954)
where the estimated optimum effort was less than the actual one in both seas.
The results (Figs. 5 & 7; Table 4) showed that a maximum sustainable yield of
about 73.0 and 63.1 thousand ton could be obtained from Mediterranean and Red seas
fisheries, respectively. The estimated optimum fishing effort was 3500 and 1656 standard
fishing unit for Mediterranean and Red seas fisheries, respectively. This means that, the
present level of fishing effort of the two seas fisheries should be decreased to obtain
MSY, while for the catch it will be increased by about 13.6% and 14.9% of its current
value by applying the optimum number of vessels.
CONCLUSION
In conclusion, based on the obtained results, the Egyptian marine fisheries are in
situation of over fishing. Although, there are some regulatory measures to manage these
fisheries like closed season for three months in Red Sea and 45 days in Mediterranean,
prohibition any new fishing licenses and any improvements on the fishing boats but these
regulations not enforced and not sufficient to conserve our fisheries. So, further studies
based on the analytical models to assess and manage the different fish stocks exploited by
different fishing gears are required. Also, make a data base about the Egyptian fisheries
containing good records for fishery statistics to facilitate the evaluation and managing
450
Shimaa I. Maiyza et al., 2020
these resources is urgent need. As well as, a multi fleet-multi species assessment models
should be used in the case of our marine fisheries.
REFERENCES
Ali, A.G. (2017). An Analytical Economic Study of The Egyptian Red-Sea Fisheries,
Ph.D., Faculty of Agriculture, Alexandria University, Egypt, .
CAPMAS (2016). The Central Agency for Public Mobilization and Statistics: Year-book
of fishery statistics (1995-1999), Cairo, Egypt.
El-Karashili, A. F. (2010). Production Planning and Management - Marine
Industries, Books Sector, Ministry of Education, Rosalieouf Press, Cairo, Egypt
El–Kholei, A. (2008), Is there overexploitation in sardine, mullet and sole catch in the
Egyptian Mediterranean Sea fisheries? A supply production model approach,
Alexandria Journal Agriculture Research, 53 (2): 1-10.
El-Serafy, S. S.; El-Gammal, F. I.; Mehanna, S. F.; Abdel-Hamid, N. H. and Farrag,
E.F.E. (2015). Age, Growth and Reproduction of the Tub Gurnard, Chelidonichthys
lucerna (Linnaeus, 1758) from the Egyptian Mediterranean waters off, Alexandria.
International Journal of Fisheries and Aquatic Sciences 4(1): 13-20.
El-Mahdy, S. M. (2017). Biological studies, population dynamics and stock assessment
of Acanthopagrus bifasciatus from the Red sea, Egypt. PhD Thesis, Assiut Univ.
GAFRD (1995-2018). The Genreal Authority for Fish Resources Development: Yearbook of fishery statistics (1995-2018), Cairo, Egypt.
Gulland, J. A. (1983). Fish stock assessment: a manual of basic methods. Chichester,
UK,Wiley Inter science,FAO/Wiley series onfood and agriculture, pp 1–223.
Haddon, M. (2011). Modeling and quantitative methods in fisheries. Second edition,
Chapman & Hall/CRC press, London 449pp.
Hassanein, E. M. (2017). Biological studies, stock assessment and fisheries management
of some species of family Mullidae from the Mediterranean Sea at Alexandria. PhD
Thesis, Fac. Sci. Assiut University.
Maiyza, SH. I. (2015). An Economic study of Fish Production In Alexandria,
Mediterranean Sea Fisheries, Ph.D., Faculty of Agriculture, Alexandria University,
Egypt, pp.164-167.
Mehanna, S. F. (2005). Population dynamics of the areolate grouper Epinephelus
areolatus from the Egyptian sector of Red Sea. 12th International Conference of
Union of Arab Biologists, El-Hodeida University, Yemen.
Mehanna, S. F. (2009). Growth, Mortality and Spawning Stock Biomass of striped red
mullet Mullus surmuletus, in the Egyptian Mediterranean waters. Med. Mar. Sci.,
10 (2): 5-17.
Mehanna, S. F. (2011). Population dynamics and management of snubnose emperor
Lethrinus bungus (L. borbonicus) from the Foul Bay, Red Sea. INOC-XI
International Symposium 2011, Bogor, Indonesia: pp.121-129.
Mehanna, S. F. (2014). Population dynamical parameters of the Atlantic lizardfish
(Synodus saurus) from the Mediterranean waters of Egypt. International Congress
“Estuaries & Coastal Protected Areas, ECPA; 2014 4-6 November 2014, Turkey.
451
An evaluation for the exploitation level of Egyptian Marine Fisheries
Mehanna, S. F. (2019a). An overview on fish production in Egypt and how to achieve its
sustainability. 3rdInternational Conference for Women in Science, 12-14 March
2019.
Mehanna, S. F. (2019b). Stock assessment of European hake, Murleccius murleccius
from the Egyptian waters of Mediterranean Sea. Expert group meeting on Hake,
Rome, 2-7 December 2019.
Mehanna, S. F. (2020). Challenges faced the small scale fisheries and its sustainable
development. ICAR- Central Marine Fisheries Research Institute, Research Centre
Mangalore, 7-10 January 2020
Mehanna, S. F. and El-Gammal, F. I. (2007). Gulf of Suez fisheries: current status,
assessment and management. J. King Abdulaziz University, Mar. Sci.,18: 3-18.
Mehanna, S. F. and Haggag, H. M. (2010). Port Said Fisheries: current status,
assessment and management. 3rd International conference on Fisheries and
Aquaculture, 29 November-1December, Cairo, Egypt. www.cabdirect.com.
Mehanna, S. F. and Haggag, H. M. (2011). Maximum sustainable yield of the flatfishes
(Family: Soleidae) from Port Said waters, Mediterranean sea, Egypt. Egypt. J.
Aquat. Biol. & Fish., Vol. 15, No. 3:285- 293.
Mehanna S. F.; Fattouh, S. A.; Farrag, E. F. and El-Sherbeny, A. S. (2011). Spatio–
temporal variations in the distribution and abundance of demersal fishes in the
Egyptian Mediterranean waters. 4th International conference on Fisheries and
Aquaculture, 3-5 October, Cairo, Egypt.
Mehanna, S.F.; Mahmoud, U. and Mohammed, A. S. (2016). Fishery status of
Carangoides bajad and Caranx melampygus (Family Carangidae) from Shalateen
Fishing Area, Red Sea, Egypt, based on yield per recruit (Y/R) analysis. Egypt. J.
Aquat. Biol. & Fish., 20 (2): 61 – 68.
Mehanna, S. F.; Osman, A. M.; Farrag, M. M. and Osman, Y. A. (2017). Age and
growth of three common species of goatfish exploited by artisanal fishery in
Hurghada fishing area, Egypt. Journal of Applied Ichthyology, 1-5.
Mehanna, S. F.; Mohammed, A. S.; Mohsen, S. and Abdel-Maksoud, Y. (2018).
Stock assessment and management of rabbitfish Siganus rivulatus from the
Southern Red Sea, Egypt.2nd International conference, Faculty of Science, Ain
Shams University “Sustainable Innovations and Sustainable Development SISD,
23-26 October, 2018.
Mehanna, S. F.; Osman, Y. A.; Khalil, M. T. and Hassan, A. (2019). Age and growth,
mortality and exploitation ratio of Epinephelus summana (Forsskål, 1775) and
Cephalopholis argus (Schneider, 1801) from the Egyptian Red Sea coast, Hurghada
fishing area. Egyptian Journal of Aquatic Biology & Fisheries, 23(4): 65-75. 115.
Mohammad, A. S. (2016). Biological studies, stock assessment and fisheries
management of Carangoides bajad (Forsskal, 1775) and Caranx melampygus
(Cuvier, 1833) from the Red Sea, Egypt. PhD Thesis, Fac. Sci., Assiut University.
Mohamed, A. S.; Mehanna, S. F.; Osman, Y. A. and El-Mahdy, S. M. (2020). Age,
growth and population parameters of the spiny squirrelfish, Sargocentron
spiniferum (Forsskål, 1775) from Shalateen fishing area, Red Sea, Egypt. Egyptian
Journal of Aquatic Biology & Fisheries, 24 (2): 469 – 480.
452
Shimaa I. Maiyza et al., 2020
Musick, J.A. and Bonfil, R. (2005). Management Techniques for Elasmobranch
Fisheries. Food and Agriculture Organization of United Nations, FAO
Fisheries Technical Paper, No. 474, Rome, FAO, 251 pp.
Osman, Y. A. (2015). Population Dynamics and Stock Assessment of the most important
species of Family Scaridae (parrot) in Hurghada, Red Sea. MSc. Thesis, Fishery and
Aquaculture Technology, Faculty of Science, Port Said University.
Schaefer, M. B. (1954). Some Aspects of the Dynamic Of Population Important To The
Management Of The Commercial Marine Fisheries, Bulletin of the Inter-American
Tropical Tuna Commission, 1(2): 27–56.
Tesfamichael, D. and Mehanna, S. F. (2012). Red Sea fisheries of Egypt: Heavy
investment and their consequences. In: Tesfamichael, D. and Pauly, D. (eds.) Catch
reconstruction for the Red Sea large marine ecosystem by countries (950 – 2010).
Fisheries Centre Research Reports, 20 (1), Vancouver.