Thierry Bourgoin
Muséum National d'Histoire Naturelle, Zoology, Faculty Member
A synthesis of planthopper suprafamilial and familial classification is provided and discussed, solving some nomenclatural issues for the group, and adopting an enriched presentation with a clear distinction between the taxon name and its... more
A synthesis of planthopper suprafamilial and familial classification is provided and discussed, solving some nomenclatural issues for the group, and adopting an enriched presentation with a clear distinction between the taxon name and its concept.
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Gedanochila museisucini gen. et sp. nov. is described, based on inclusions in the Eocene Baltic amber. A morphological phylogenetic analysis supports the placement of Gedanochila gen. nov. into the tribe Achilini. Definition, content and... more
Gedanochila museisucini gen. et sp. nov. is described, based on inclusions in the Eocene Baltic amber. A morphological phylogenetic analysis supports the placement of Gedanochila gen. nov. into the tribe Achilini. Definition, content and subdivisions of the tribe as well as position of extinct taxa placed within are briefly discussed.
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Abstract A new species, Dorytocus jiaxiaoae Song, Szwedo & Bourgoin sp. nov., from the mid-Cretaceous Burmese amber is described and illustrated. This species represents the first record of the adult of the extinct family Dorytocidae.... more
Abstract A new species, Dorytocus jiaxiaoae Song, Szwedo & Bourgoin sp. nov., from the mid-Cretaceous Burmese amber is described and illustrated. This species represents the first record of the adult of the extinct family Dorytocidae. The supplementary description for the fourth-instar nymph of Dorytocus ornithorhynchus Emeljanov & Shcherbakov is provided. The morphological peculiarities of the new fossil species are briefly discussed, and the diagnostic characters of the family diagnosis are suggested and improved is now based on the combination of both adult and nymph characters.
Research Interests: Zoology, Geography, Geology, Biology, Fossil record, and 4 moreCretaceous, Hemiptera, Nymph, and Burmese
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A digital resource, which aims to become a comprehensive catalogue of all known species of organisms on Earth.
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Two new genera of the tribe Sarimini are described from south of China. They are Sarimissus gen. nov. with a new species S. maculifrons sp. nov. from Hainan province and Duplexissus gen. nov. with a new species D. punctatulus sp. nov.... more
Two new genera of the tribe Sarimini are described from south of China. They are Sarimissus gen. nov. with a new species S. maculifrons sp. nov. from Hainan province and Duplexissus gen. nov. with a new species D. punctatulus sp. nov. from Yunnan province. The photos of habitus and the male genitalia of the new species are provided. The biology, distribution and origin of the tribe Sarimini are discussed.
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To our present knowledge Auchenorrhyncha are represented in Reunion island by 102 species described : 66 Fulgoromorpha and 36 Cicadomorpha. Inmost of the undisturbed biotopes studied, Fulgoromorpha show more significant diversity than... more
To our present knowledge Auchenorrhyncha are represented in Reunion island by 102 species described : 66 Fulgoromorpha and 36 Cicadomorpha. Inmost of the undisturbed biotopes studied, Fulgoromorpha show more significant diversity than Cicadomorpha. Except for Delphacidae and Flatidae which feed primarily on exotic plants, the other families of the Fulgoromorpha are characterised by a strong proportion of endemic species mainly dependent on indigenous plants. Our research shows preferential trophic relation between some Fulgoromorpha families or species and particular plant taxa. This is the case with Derbidae which are all associated with indigenous Monocotyledones. Achilidae and a new endemic species of Meenoplidae attached to the new genus Insulisia Bonfils,1998, seem to be monophagous on some indigenous plants. Cixiidae and other families are mostly polyphagous but linked with indigenous plants. These preferential associations between endemic Fulgoromorpha and indigenous plants allow to provide informations on the structure and evolution of ecosystems in Reunion island. With the objective of maintaining the original character of Reunion biodiversity, these studies would allow to direct conservation decisions for this endangered insular biotope. Lastly, these results also open new fields of research : comparative studies of the evolution of Mascarene islands environments, as well as modalities of Mascarenes colonization by Auchenorrhyncha. (Resume d'auteur)
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An endemic karst dwelling gecko from western Madagascar is commonly encountered in caves. Copulation was observed during daytime and we discuss two conflicting or complementary hypotheses about its cave occurrence during daytime, trophic... more
An endemic karst dwelling gecko from western Madagascar is commonly encountered in caves. Copulation was observed during daytime and we discuss two conflicting or complementary hypotheses about its cave occurrence during daytime, trophic hypothesis and predation avoidance hypothesis.
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A new planthoppers genus and species of CixiidaeOecleini, Coframalaxius bletteryigen. nov. sp. nov. newly discovered in a cave near Nice in southern France, is described. Molecular analysis confirms the morphology-based classification of... more
A new planthoppers genus and species of CixiidaeOecleini, Coframalaxius bletteryigen. nov. sp. nov. newly discovered in a cave near Nice in southern France, is described. Molecular analysis confirms the morphology-based classification of Coframalaxius as sister to Trigonocranus within the Oecleni. Several morphological characters are further discussed. A double-grasping coxo-femoral and femoro-tibial system is regarded as apomorphic for the oecline taxa and would allow the nymph to firmly grab the roots and rootlets on which it feeds or use to progress in the soil. Wing vein patterns are discussed in the Cixiidae: 1) for the forewings, Oecleini belong to the trifid type of the anterior MP branch, leading to the reinterpretation of some recently described Neotropical species, 2) for the hindwing, four connection types (U-, V-, Y- and I-types) between MP and CuA are described. Oecleini belongs to I-type with a complete fusion of MP3+4 with CuA1. Although the area where the cave is loc...
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Cixiidae are small strictly phytophagous hemipteran insects worldwide distributed. Ecology and systematics of Chinese fauna remains poorly investigated. For instance, does their distribution follows the patterns of biogeogaphical... more
Cixiidae are small strictly phytophagous hemipteran insects worldwide distributed. Ecology and systematics of Chinese fauna remains poorly investigated. For instance, does their distribution follows the patterns of biogeogaphical distribution established for their host plants or other related-taxa because they are all obligatory phytophagous taxa? Do they follow the usual distributional Chinese realms and boundaries already recognized? Which zoogeographical Chinese regions and connections between them do they depict. To investigate these issues, we provide here a referenced and comprehensive checklist of the 250 cixiid species currently reported from China (77 new records), with their precise distribution at the regional level. In the 8 Chinese main zoogeographical regions usually recognized and 2 adjacent areas, we analyzed further their diversity at the tribal, generic, and specific levels using a non-metric multidimensional scaling and an unweighted pairwise group analysis using ...
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Abstract The genera of Tropiduchidae tribe Paricanini are discussed and their key to identification of the genera is given and key characters illustrated. The new genus Lukabales gen. nov. with a new species Lukabales ecarinatus sp. nov.... more
Abstract The genera of Tropiduchidae tribe Paricanini are discussed and their key to identification of the genera is given and key characters illustrated. The new genus Lukabales gen. nov. with a new species Lukabales ecarinatus sp. nov. is described based on specimens collected in montane areas of Cambodia and Thailand.
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Background Cixiidae are small strictly phytophagous hemipteran insects worldwide distributed. Ecology and systematics of Chinese fauna remains poorly investigated. For instance, does their distribution follows the patterns of... more
Background Cixiidae are small strictly phytophagous hemipteran insects worldwide distributed. Ecology and systematics of Chinese fauna remains poorly investigated. For instance, does their distribution follows the patterns of biogeogaphical distribution established for their host plants or other related-taxa because they are all obligatory phytophagous taxa? Do they follow the usual distributional Chinese realms and boundaries already recognized? Which zoogeographical Chinese regions and connections between them do they depict. To investigate these issues, we provide here a referenced and comprehensive checklist of the 250 cixiid species currently reported from China (77 new records), with their precise distribution at the regional level. In the 8 Chinese main zoogeographical regions usually recognized and 2 adjacent areas, we analyzed further their diversity at the tribal, generic, and specific levels using a non-metric multidimensional scaling and an unweighted pairwise group anal...
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A new Burmese amber genus Maculixiusgen. nov. with its type species Maculixius jiewenaesp. nov. is described in the planthopper family Cixiidae. This new genus is unique in Burmese Cixiidae by its forewing venation, with Pcu and A1... more
A new Burmese amber genus Maculixiusgen. nov. with its type species Maculixius jiewenaesp. nov. is described in the planthopper family Cixiidae. This new genus is unique in Burmese Cixiidae by its forewing venation, with Pcu and A1 merging in the middle of clavus, the late bifurcation of ScP+R slightly after this level, and the early CuA forking well before this level. Although the head capsule is missing, it belongs to the recently described tribe Acrotiarini, based on the pentacarinated mesonotum and the distinctly arched RA on forewing with cell C1 wider submedially than apically. Morphological characteristics of Acrotiarini are discussed, and a key to identification of Acrotiarini genera is provided. The new taxon broadens the knowledge of the tribe, and it underlines the already great diversity of the family in the Cretaceous.
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Prace prezentuje seznam polokřidlých (Hemiptera) nalezených v cervnu 2015 během terenni exkurze pořadane v ramci mezinarodni konference 7th European Hemiptera Congress ve Stýrsku (Rakousko). Celkem bylo nalezeno 144 druhů křisů... more
Prace prezentuje seznam polokřidlých (Hemiptera) nalezených v cervnu 2015 během terenni exkurze pořadane v ramci mezinarodni konference 7th European Hemiptera Congress ve Stýrsku (Rakousko). Celkem bylo nalezeno 144 druhů křisů (Auchenorrhyncha), 143 druhů plostic (Heteroptera), 13 druhů mer (Psylloidea) a 2 druhy msic (Aphididae). Druhy Ribautodelphax imitans (Delphacidae), Eurhadina saageri (Cicadellidae), Notonecta maculata (Notonectidae), Notonecta meridionalis (Notonectidae) a Polymerus cognatus (Miridae) představuji nove nalezy pro Stýrsko.
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Two new species of the genus Saigona Matsumura, 1910, S. baiseensis Zheng & Chen sp. nov. and S. maculata Zheng & Chen sp. nov., from China (Guanxi) are described and illustrated. A revised identification key to the 16 species of Saigona... more
Two new species of the genus Saigona Matsumura, 1910, S. baiseensis Zheng & Chen sp. nov. and S. maculata Zheng & Chen sp. nov., from China (Guanxi) are described and illustrated. A revised identification key to the 16 species of Saigona is provided. 15 species of the genus are known from China only.
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Abstract A new family Yetkhatidae Song, Szwedo et Bourgoin fam. nov. from mid-Cretaceous amber of Myanmar is described. It is characterized by the tegmina with a triangular basal cell and in having the apex of the clavus closed, the... more
Abstract A new family Yetkhatidae Song, Szwedo et Bourgoin fam. nov. from mid-Cretaceous amber of Myanmar is described. It is characterized by the tegmina with a triangular basal cell and in having the apex of the clavus closed, the female genitalia with robust endogonocoxal processes, the head capsule with trigones and foliated frontal margins, ridged lora (visible frontally) and by intermediate carinae on the mesonotum. Two new monotypic genera are described in this new family: Yetkhata Song, Szwedo & Bourgoin gen. nov., with Yetkhata jiangershii Song, Szwedo & Bourgoin sp. nov. and Parwaina Song, Szwedo & Bourgoin gen. nov. with Parwaina liuyei Song, Szwedo & Bourgoin sp. nov. A key to recognition of these genera is provided, and their morphological characters and disparity are briefly discussed.
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To explore the phylogenetic basis of the current classification of Dictyopharidae, especially the relationships among the tribes of Dictyopharinae, we present the first cladistic analysis of this family based on 146 morphological... more
To explore the phylogenetic basis of the current classification of Dictyopharidae, especially the relationships among the tribes of Dictyopharinae, we present the first cladistic analysis of this family based on 146 morphological characters of adults. Our analysis includes 104 of 125 recognized genera and subgenera within 12 extant tribes of Dictyopharinae, plus nine genera representing all four tribes of Orgeriinae. The results of this study support Dictyopharidae as a monophyletic group with Aluntiini sister to the remaining Dictyopharidae, but do not support Orgeriinae as sister to Dictyopharinae. Seven major lineages – Aluntiini, Arjunini, Hastini, (Taosini + Lappidini) + Nersiini, a Xenochasma+ complex (including ‘Orgeriinae’), Orthopagini, and Dictyopharini – are recovered in Dictyopharidae. The Xenochasma+ complex is proposed here and includes Xenochasma clade + (Scoloptini + ((Fernandea clade + Phylloscelini) + (Rancodini + [Capenini + Orgeriinae]))). Within this complex, some genera are of uncertain tribal placement, and the Orgeriinae are retained as a subfamily until a molecular phylogeny can confirm the results of this paper. The implied weighting analysis supports the monophyly of most tribes of Dictyopharinae (except Taosini), the sister‐group relationships of (Taosini + Lappidini) with Nersiini, and Orthopagini + Dictyopharini, and the current tribal classification for Dictyopharinae. Most Dictyopharinae genera fit their respective tribal affiliation, but some proposed changes are that Pharodictyon, Paramisia, Dictyopharoides s.s., Chondrophana, Sicoris, Chondrodire, and Tupala are provisionally placed in Hastini; Pukuakanga is moved into Nersiini; Sinodictya and Raphiophora are transferred into Orthopagini; and Chiltana, Litocras, and Viridophara are placed in Dictyopharini.
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The first part of this paper provides a historical review of the classification of the family Tettigometridae, including the description of a new tribe, Plesiometrini trib. nov. to accommodate three Afrotropical genera of the subfamily... more
The first part of this paper provides a historical review of the classification of the family Tettigometridae, including the description of a new tribe, Plesiometrini trib. nov. to accommodate three Afrotropical genera of the subfamily Tettigometrinae. The name Nototettigometra Muir 1924, is proposed to replace Hilda Kirkaldy 1900, homonym of Hilda Hörnes Auinger 1884, and corresponding new combinations are given, including Nototettigometrinae nom. subst. to replace Hildinae Fennah 1952. An identification key to suprageneric taxa of Tettigometridae including both male and female characters is provided. In the second part, 18 tettigometrid species are recorded from Iran. A new species, Tettigometra (Tettigometra) parihana sp. nov. is described. Tettigometra (Metroplaca) longicornis and Tettigometra (Tettigometra) impressifrons are reported as new records for the Iranian fauna. An identification key and distribution maps are provided for the Iran fauna.
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A morphological phylogenetic analysis of the tribe Tropiduchini (Hemiptera: Tropiduchidae) is provided for 21 genera among the 26 recognized, including the new genus Oechalinella Wang gen.n., with a new species Oechalinella bifasciata... more
A morphological phylogenetic analysis of the tribe Tropiduchini (Hemiptera: Tropiduchidae) is provided for 21 genera among the 26 recognized, including the new genus Oechalinella Wang gen.n., with a new species Oechalinella bifasciata Wang sp.n. Monophyly of the tribe is well supported by synapomorphies derived from the male genitalia. The strict consensus supports two subclades: ((Montrouzierana + Thymbra) + Thaumantia+) as sister to the clade (Leptovanua + (Vanua + Varma+) + Daradacella+). In the first subclade, the clade (Montrouzierana + Thymbra), distributed in New Caledonia and Papua New Guinea, is well supported by three synapomorphies based on the tegmina, while the Thaumantia+ group is characterized by the conformation of the anal tube and male gonostyli, and distributed from Malesia to Papuasia and in Central Africa. The second subclade is largely distributed from China to Malesia, Papuasia, Southwestern and the Northwestern Pacific islands. The position of the genus Leptovanua remains uncertain. The Varma+ clade is recovered in all analyses. Implied weighting analysis placed the Varma+ lineage as sister to all other genera forming a new group (Leptovanua+ + ((Montrouzierana + Thymbra) + Thaumantia+)). Fennah's definition of Tropiduchini is reviewed according to the phylogenetic results and an identification key to all genera is provided with a referenced synopsis of their distribution. Globally, Tropiduchini exhibits an inter‐tropical latitudinal distribution and a paradoxical bimodal longitudinal distribution pattern in continental Africa, South‐east Asia and the Pacific islands. Distribution data concern nearly 80% of insular taxa. The phylogenetical results suggest that: (i) the tribe originates from continental China; (ii) it evolved mainly from Papuasia by stepping‐stone colonization events, promoting subsequent insular speciation; and (iii) its evolution has mainly been directed by dispersion versus vicariance.
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Three new Oriental Issidae genera and two new species identified by a previous molecular phylogeny analysis are described. Two new genera belong to the tribe Kodaianellini: Kodaianellissus gen. nov., type species: K. intorqueus sp. nov.... more
Three new Oriental Issidae genera and two new species identified by a previous molecular phylogeny analysis are described. Two new genera belong to the tribe Kodaianellini: Kodaianellissus gen. nov., type species: K. intorqueus sp. nov. from China and Tetricissus gen. nov., type species: T. philo (Fennah, 1978) from Vietnam. Another new genus representing Sarimini: Longieusarima gen. nov., type species: L. lunulia sp. nov. is described from China.
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Abstract. A new species of the planthopper taxon Bennini (Hemiptera: Fulgoromorpha: Cixiidae) was discovered in the wide interstitial voids of a granitic blockfield in Vietnam. A key to the species of the Vietnam-endemic Bennini genus... more
Abstract. A new species of the planthopper taxon Bennini (Hemiptera: Fulgoromorpha: Cixiidae) was discovered in the wide interstitial voids of a granitic blockfield in Vietnam. A key to the species of the Vietnam-endemic Bennini genus Sanghabenna is provided and Sanghabenna florenciana sp. nov. is described and illustrated. Molecular data including 18S in part and CO1 barcode sequences of the species, as well as for the first time a detailed description of nymphal morphology of a Bennini are provided. According to the ecological parameters of its habitat and its morphological configuration, Sanghabenna florenciana sp. nov. is classified as an exapted eutroglophile taxon.
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A molecular phylogeny of the planthopper family Issidae (Hemiptera, Fulgoroidea) is provided using both Maximum likelihood and Bayesian inference analyses. The phylogeny is based on 18S, two parts of 28S, COXI and Cytb genes from 50... more
A molecular phylogeny of the planthopper family Issidae (Hemiptera, Fulgoroidea) is provided using both Maximum likelihood and Bayesian inference analyses. The phylogeny is based on 18S, two parts of 28S, COXI and Cytb genes from 50 genera and 79 ingroup species (including 8 species recently excluded from Issidae). As with the only few previous studies, an important taxonomic impediment is observed with the sampling; however for the first time, all analyses depict several fully supported lineages, which challenge the recent proposed "modern classification" of the family. It also highlights a strong coherence between these lineages and their respective geographical distribution. All previously excluded taxa are confirmed as not being part of the Issidae as recently defined which monophyly is confirmed. Accordingly, a new classification of the family is proposed with 3 subfamilies and 7 tribes as follows. Neotropical issid Thioniini in Thioniinae stat. rev. is re-established...
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Abstract. Euxaldar jehucal Fennah, 1978 is redescribed. Euxaldar lenis sp. nov. is described from Southern Vietnam. Sexual dimorphism and asymmetry in male genitalia are recorded for the genus Euxaldar Fennah, 1978.
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One Issidae specimen stored in Paris museum historical collections is reported as holotype of Issus analis Brullé, 1833. From the original description, which is confirmed by study of this specimen, the species is moved to the genus... more
One Issidae specimen stored in Paris museum historical collections is reported as holotype of Issus analis Brullé, 1833. From the original description, which is confirmed by study of this specimen, the species is moved to the genus Zopherisca Emeljanov, 2001 under a new combination Zopherisca analis (Brullé, 1833), comb. n. Date of description is discussed and modified from 1832 to 1833 accordingly. Unfortunately being a female as type specimen, the species remains quite enigmatic until some molecular analsysis could be undertaken on this old material.
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This is the first substrate-borne communication record of the troglobitic planthopper Typhlobrixia namorokensis in the caves of the Ambovonomby network, Namoroka Tsingy, Madagascar. A supplementary morphological description of the male... more
This is the first substrate-borne communication record of the troglobitic planthopper Typhlobrixia namorokensis in the caves of the Ambovonomby network, Namoroka Tsingy, Madagascar. A supplementary morphological description of the male genitalia is provided and, for the first time, female genitalia of T. namorokensis are characterized. Molecular data were obtained for T. namorokensis; molecular analysis also reveals the presence of another (yet unidentified) cavernicolous cixiid in the Antsifotra caves network. The evolutionary origin of T. namorokensis is discussed. Its adaptation to caves and its behaviour allows its identification as a true troglobiont. Its vibrational signal structure is compared to the calls of other cave-dwelling cixiids.
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Does the distribution of the Hemiptera planthoppers Cixiidae, follow the patterns of biogeogaphical distribution already well established for their host plants or other taxa because they are all obligatory phytophagous taxa? Are their... more
Does the distribution of the Hemiptera planthoppers Cixiidae, follow the patterns of biogeogaphical distribution already well established for their host plants or other taxa because they are all obligatory phytophagous taxa? Are their realms and boundaries recognized? What are their zoogeographical regions and usual connections? To investigate these issues, we provide here a referenced and comprehensive checklist of the 253 cixiid species currently reported from China, with their precise distribution at the regional level. Seventy seven of these species are new records for China. In the 8 Chinese main zoogeographical regions usually recognized and 2 adjacent areas, we analyzed further their diversity at the tribal, generic and specific levels using a non-metric multidimensional scaling and an unweighted pairwise group analysis using an arithmetic mean cluster analyses. The observed distribution patterns have shown that an intercalary Sino-Japanese realm is present between the Palaea...