Gorgonops

Gorgonops; Temporal range: Lopingian (Wuchiapingian), 260–254 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	G. whaitsi skull (specimen 5537)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Synapsida
Clade:	Therapsida
Clade:	†Gorgonopsia
Family:	†Gorgonopsidae
Subfamily:	†Gorgonopsinae
Genus:	†Gorgonops; Owen, 1876
Type species
	†Gorgonops torvus; Owen, 1876
Species
	†G. torvus Owen, 1876; †G. whaitsi Broom, 1912; †G. longifrons Haughton, 1915;

Gorgonops (from Greek: Γοργών 'Gorgon' and ὤψ 'eye, face', literally 'Gorgon eye' or 'Gorgon face') is an extinct genus of gorgonopsian therapsid, of which it is the type genus. Gorgonops lived during the Late Permian (Wuchiapingian), about 260–254 million years ago in what is now South Africa.

History of discovery

The holotype of the type species, Gorgonops torvus, was one of the first therapsids discovered. It was described by Richard Owen, who also coined the name "Dinosauria" on the basis of the first known dinosaur fossils.^[1] G. torvus was also used as the type for the family Gorgonopsidae, which was described by Richard Lydekker in 1890.^[2] Five years later, in 1895, Harry Govier Seeley used this genus to establish the larger clade of Gorgonopsia.^[3] In later years, a large number of further species and genera were designated, though many of these were later determined to be synonyms.

Gorgonops is known from the Tropidostoma and most of the Cistecephalus Assemblage Zones.^[4]

Description

Restoration of G. whaitsi

Gorgonops was a medium-sized gorgonopsian, with a skull length of 22–35 centimetres (8.7–13.8 in), depending on the species. They ranged from 1.2–2 metres (3.9–6.6 ft) long from nose to tail. Gorgonops would have been one of the key predators across southern Africa during the Late Permian. Because the canines were so large, they would have had little trouble in penetrating the tough hides of some of the herbivores of the time, particularly pareiasaurs such as Pareiasaurus. Aside from the teeth, one of the key predatory advantages that Gorgonops had over prey was its semi-erect gait, compared to the sprawling gait exhibited by most prey animals of the time. Aside from allowing for more energy efficient locomotion, this allowed Gorgonops to travel at relatively high speeds.

Skull

Gorgonops compared to a human.

Relative to body size, Gorgonops had a deep skull with a triangular profile when viewed from above. Perhaps the most distinctive features were two enlarged canine teeth that were so big (12 centimetres (4.7 in) long) they almost protruded beyond the lower jaw. To help protect these teeth, the lower jaws grew in such a shape so that the anterior (front) portion was thicker than the posterior (rear) portion. This form would have protected the enlarged canine teeth from accidental damage, and was similar in bone function to the flanges of bone of sabre-toothed cats in the Cenozoic.

Species

Gorgonops torvus (Owen, 1876)

G. torvus

The type species. The holotype is an incomplete and flattened skull, allegedly found at Mildenhall's farm (Xlu Xlu), on the Queen's Road south of Fort Beaufort, in the Eastern Cape of South Africa. A number of other specimens have been found since, all from the Tropidostoma and/or Cistecephalus Assemblage Zone(s). This was a medium-sized therapsid, with a skull about 22 cm in length. It is distinguished from other species by a longer snout, and other details of the bones of the skull. Originally considered rather simple, it is actually (according to Sigogneau-Russell) a rather specialised member of the group.^[5]

Gorgonops whaitsi (Broom, 1912)

Head of G. whaitsi

Larger than G. torvus, with the rear of the skull wider, and other details of proportion. Originally the type species of Scymnognathus. Despite being known from a large number of specimens from the Karoo Basin, Beaufort West (Tropidostoma/Cistecephalus Assemblage Zone), the species remains poorly known. Watson and Romer placed Gorgonops and Scymnognathus in two different families, while Sigogneau-Russell placed the two species in the same genus, and considers G. whaitsi a more primitive (less derived) form.^[5] Synonyms: Scymnognathus whaitsi (Broom, 1912)

Gorgonops longifrons (Haughton, 1915)

A large specimen known from an incomplete and flattened skull about 35 centimetres (14 in) long. Orbit larger and snout longer than G. whaitsi, from which it may have descended. Beaufort West, Tropidostoma/Cistecephalus Assemblage Zone. Synonyms: Gorgonognathus longifrons (Haughton, 1915)

Gorgonops? eupachygnathus (Watson, 1921)

A flattened, incomplete, medium-sized skull, probably a juvenile of either G. torvus or G. whaitsi Synonyms: Leptotrachelus eupachygnathus (Watson, 1921); Leptotracheliscops eupachygnathus (Watson, 1921)

Gorgonops? dixeyi (Haughton, 1926)

A large, incomplete and flattened skull, from Chiweta Beds, Nyassaland. Placement uncertain. Probably Low Cistecephalus Assemblage Zone equivalent (= middle of the Wuchiapingian Stage).^[6] Synonyms: Chiwetasaurus dixeyi (Haughton, 1926)

Gorgonops? kaiseri (Broili & Schroeder, 1934)

A large (about 35 centimetres (14 in) long), incomplete skull, with a high snout and narrower in the rear than other species, from the "High Tapinocephalus zone" (earlier than the other species, most probably Pristerognathus Assemblage Zone) Synonyms: Pachyrhinos kaiseri (Broili & Schroeder, 1934)

Classification

Below is a cladogram from Gebauer's 2007 phylogenetic analysis.^[7]

Gorgonopsia

Gorgonops

Njalila

	Sycosaurus

	Clelandina

References

^ Owen, R. (1986). Descriptive and illustrated catalogue of the fossil Reptilia of South Africa in the collection of the British museum. British Museum (Natural History). pp. 27–29.
^ Lydekker, R. (1890). Catalogue of the fossil Reptilia and Amphibia in the British Museum (Natural history) Part IV. British Museum (Natural History). p. 111.
^ Seeley, H. G. (1895). "Researches on the structure, organization, and classification of the fossil reptilia.—Part IX. section 1. On the Therosuchia". Annals and Magazine of Natural History. 13 (6): 375. doi:10.1080/00222939408677718.
^ Smith, R.H.M.; Keyser, A.W. (1995). "Biostratigraphy of the Tropidostoma Assemblage Zone". Geological Survey of South Africa. 1: 18–22.
^ ^a ^b Sigogneau-Russell, D. (1989). "Theriodontia I". In H.-D., Sues (ed.). Encyclopedia of Paleoherpetology. Gustav-Fischer-Verlag. ISBN 978-3-89937-033-1.
^ Jacobs, L.L.; Winkler, D.A.; Newman, K.D.; Gomani, E.M.; Deino, A. (2005). "Therapsids from the Permian Chiweta Beds and the age of the Karoo Supergroup in Malawi". Palaeontologia Electronia. 8 (1): 28A.
^ Gebauer, E.V.I. (2007). Phylogeny and evolution of the Gorgonopsia with a special reference to the skull and skeleton of GPIT/RE/7113 ('Aelurognathus?' parringtoni) (PDF) (Ph.D. thesis). Tübingen: Eberhard-Karls Universität Tübingen. pp. 1–316.

[Owen1876-1] Owen, R. (1986). Descriptive and illustrated catalogue of the fossil Reptilia of South Africa in the collection of the British museum. British Museum (Natural History). pp. 27–29.

[2] Lydekker, R. (1890). Catalogue of the fossil Reptilia and Amphibia in the British Museum (Natural history) Part IV. British Museum (Natural History). p. 111.

[3] Seeley, H. G. (1895). "Researches on the structure, organization, and classification of the fossil reptilia.—Part IX. section 1. On the Therosuchia". Annals and Magazine of Natural History. 13 (6): 375. doi:10.1080/00222939408677718.

[4] Smith, R.H.M.; Keyser, A.W. (1995). "Biostratigraphy of the Tropidostoma Assemblage Zone". Geological Survey of South Africa. 1: 18–22.

[:0-5] Sigogneau-Russell, D. (1989). "Theriodontia I". In H.-D., Sues (ed.). Encyclopedia of Paleoherpetology. Gustav-Fischer-Verlag. ISBN 978-3-89937-033-1.

[6] Jacobs, L.L.; Winkler, D.A.; Newman, K.D.; Gomani, E.M.; Deino, A. (2005). "Therapsids from the Permian Chiweta Beds and the age of the Karoo Supergroup in Malawi". Palaeontologia Electronia. 8 (1): 28A.

[GEVI07-7] Gebauer, E.V.I. (2007). Phylogeny and evolution of the Gorgonopsia with a special reference to the skull and skeleton of GPIT/RE/7113 ('Aelurognathus?' parringtoni) (PDF) (Ph.D. thesis). Tübingen: Eberhard-Karls Universität Tübingen. pp. 1–316.

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