Swine Nutrition for Pork Producers
Swine Nutrition for Pork Producers
Nutrition Guide
Preface
  The objective of the Swine Nutrition Guide              emphasizing a balanced approach to this sometimes
remains the same: to provide the reader with a            complex subject but also briefly explaining the
simple, easy-to-read source of practical information      specific mechanics as well.
on the nutrition of the pig. Fundamentally, the pig
has changed little since 1989 when the first edition        Chapters 6 to 9 discuss the day-to-day feeding of
was released; of course, genetic progress has been        various classes of swine. Because nutrition cannot
impressive, but the list of nutrients required by the     be considered in isolation, other aspects of manage-
pig and how they are used have remained much the          ment are also included. The authors have taken a
same.                                                     more factorial approach to the design of feeding
                                                          programs than in the first edition. This reflects
   What has changed over the past 6 years is the          what we believe to be a growing trend in pig nutri-
tremendous growth in knowledge on how to define           tion - to define nutrient requirements on the basis of
nutrient requirements and how to provide these            specific traits and desired levels of productivity,
nutrients more precisely and effectively with a           with economics being considered in all instances.
keener eye to economics. Perhaps even more
striking has been the rapid progress of pork produc-        Chapter 10 considers the processing and manufac-
ers in their ability to take advantage of this informa-   turing of pig diets, a step that is often taken for
tion and apply it "on the farm."                          granted - at considerable peril. Mixing and delivery
                                                          of feed to the pig represents the final step in the
  The target audience of the Swine Nutrition Guide        nutrition continuum. Failure to manufacture diets
also remains the same: pork producers, workers in         properly can erase any gains achieved through
agribusiness and extension, and students. The             improved diet formulation or ingredient selection!
common interest is information on how the pig uses
nutrients, how to determine the nutrient require-           Chapter 11 provides information on toxins or
ments of a given class of swine and how feedstuffs        antinutritional factors that may be present in feeds
can be most effectively employed in the diet. We          and feedstuffs, either as a consequence of contami-
sincerely hope that the Guide continues to fill this      nation or due to the natural composition of the
need - serving as a single source reference on these      feedstuff itself. A new chapter (12) on water has
topics.                                                   been added to the second edition of the Swine
                                                          Nutrition Guide, discussing both nutritional and
  The book is not intended to make all readers pig        quality issues.
nutritionists, but rather to provide an understanding
and appreciation for the importance of a holistic           We trust that the interests of the individual pig are
approach to the subject; the pig does not exist in a      not lost in the pages of this book. Readers will note
vacuum and to be successful, feeding programs             that considerations of pig behaviour creep into
must be designed with many factors in mind,               many chapters as we learn more about this aspect of
including genetics, the environment, herd health          the animal that we serve - and who has served us so
and, of course. economics.                                well. Our respect and affection for this creature, at
                                                          once both intelligent and stubborn, fun and frustrat-
  The format of the second edition remains the            ing, neat and messy, motivate us to again dedicate
same as the first. Chapters l, 2 and 3 provide            the Swine Nutrition Guide to the pig.
background on the more fundamental aspects of
nutrition. Chapter 4 describes more than 40 com-
mon ingredients, including recommendations on              J.F. Patience
how they might be used in pig diets. Chapter 5 then        P.A. Thacker
explains how to successfully formulate diets,              C.F.M. de Lange
Table of Contents
1. The Science of Nutrition                                                                                                                                                                       I
    What is Nutrition                                                                                                                                                                             1
    The Role of the Computer                                                                                                               .                                                      3
    The Future of Nutrition                                                                                                                                                                       4
2. Gastrointestinal Physiology                                                                                                                                                                    5
    The Role of the Digestive Tract ..                                                                                                                                      .                     5
    Types of Digestive Tracts                                                                                                                                                                     6
    Eating and Swallowing                                                                                                                                                                         7
    The Stomach . . . . .. . . .. . . .. . .. . . . .. . .. . . .. . . .. . . .. . . .. . . . . . . .. . . .. . . . . . . .. . . .. . . . .. . .. . . . . . . .. . . . .. . .. . .. . . . .. .    7
    The Small Intestine                                    .                                                                .                                          .            .             9
    The Large Intestine .. . . . .. . . . .. . . . . . .. .. . . .. .. . . .. . . .. . . .. . . .. . . . .. . . . . . . .. . .. . . . .. . . .. . . .                                            12
3. Nutrients                                                                                                                                                                                     15
    What is a Nutrient . . . . . . . . . . . . . . . . .. . .. . . .. . . .. . . . . . . .. . . .. . . .. . . .. . .          . .. . . . . . . . . . . . .. . . .. . .. . . .                    15
    Nutrient Availability                                                                                                                                                                        15
    Nutrient Requirements ..                                                                                                                   . . .                   .                         16
    Energy                                                                                                                                                                                       16
    Amino Acids . . . . . . . .. . . .. . .. . . .. .. .. ..                        . .. . . .. . .. .. .. .. .. . . . .. . . .. . . . .. . . .. . . .. . .. . . ..                              22
    Minerals                                                                                                                                                                                     31
    Vitamins .. . .                                 ..                                                                                                                .                          40
    Essential Fatty Acids .                         . . . . . . . . . . . .. . . . . . . . . . . .. . . .. . . .. .. .. . . . . . . .. . . .. . . . .. . . .. . . . .                            48
    Water                                                                                                                                                                                        49
4. Ingredients                                                                                                                                                                                   51
    Ingredient versus Nutrient ..                                                                                                                                                                51
    Animal Sources                                                                                                                                                                               53
       Blood Meal .. . . .. . . . . . . . .. . . .. . . . .. . . .. . . . . . . . . . . . .. . . .. . .. .. . .. .. .. . .. . . .. .. .. .. . .. .. .. ..                                        54
       Bone Meal                                                                                                                                                                                 55
       Feather Meal                                                                                                                                                                              55
       Fish Meal ..                                                                                                                                    ..                                        56
       Fish Silage                                                                                                                                                                               57
       Meat and Bone Meal                                                                                       .. ..                                                                            59
       Whole Milk                                                                                                                                                                                60
       Dried Skim Milk                                                                                                                                                                           61
       Whey                                                                                                                                                                                      61
    Plant Sources . . . .                                                                            .                         .                  . ..                                           63
       Alfalfa                                                                                                                                                                                   63
       Barley                                                                                                                                                                                    65
       Barley: High Moisture                                                                                                                                                                     67
       Barley: Hulless                                                                                                                                                                           69
       Beet Pulp                                                                                                                                                                                 71
       Buckwheat                                                                                                                                                                                 71
       Cano la Meal .                                                                                     . .                                   .. ..                                            73
        Canola Seed: Full Fat                                                                                                                                                          74
        Corn                                                                                                                                                                      .    76
        Corn By-Products                                                                                                                                                          .    78
        Dried Bakery Product                                                                                                                                                      .    80
        Fababeans                                                                                                                                                               .      80
        Field Peas                                                                                                                                                          ..         82
        Grain Dust                                                                                                                                                              .      84
        Grain Screenings                                                                                                                                                               84
        Lentils                                                                                                                                                                        86
       Molasses                                                                                                                                                                        87
       Oats                                                                                                                                                                            87
       Oats: Naked .. . .                                     .                        . .                                                                                             89
       Oat Groats . .                       .                                                                                                                  . .                     90
       Potatoes and Potato Products . . . .. . . .. .. . . .. . . .. .. .. . . .. . . . . . . .. . .. .. .. .. .. .. . . . . .. . . ..                                                 90
       Rye                                                                                                                                                                             92
       Soybeans: Full Fat                                                               .                                                                   .               ..         93
       Soybean Meal                                                                                                                                                                    96
       Sunflower Meal                         .                                                                          . .                                                           97
       Sunflower Seeds . .. .                                                  .                                                                                               .       98
       Triticale . . .. .. . .                      .                                                                .                                                        .       100
       Wheat                                                                                                                                                                          101
       Wheat Bran .. . . . . . .. . . .. . . . . . . .. . . .. . . .. . .. . .. .. . . .. . . . .. .. .. . . .. . . .. .. . . . . .. . . .. . .. . . . . . . . .. . .                 104
       Wheat Shorts                                     .                           . .                                                                                               105
       Wild Oat Groats                                    .                          .                       . . .                                                           .        105
    Fats and Oils .           .               .                             . .                                                                                            ..         107
    Mineral Ingredients ..                                                                                                                                                  ..        112
    Probiotics                                                                                                                                                                        113
    Synthetic Amino Acids                                                                                                                                                             115
    Organic Acids                                                                                                                                                                     115
    Feed Flavours . . . .. . . . . . . . . . . . . . . .. . . . . . .. . . .. . . .. . . . .. . . . . . . .. . . .. . . .. . . .. . . .. .. .. . . . . . . .. . . . . . . ..          116
• ••
                                              •
                                              . , • ... .
                                          •       •              •
FEATHER MEAL       FISH MEAL           FISH SILAGE (CAPELIN)
BARLEY             BARLEY:HULLESS
BEET PULP                                                                               CANOLASEED    CANOLA MEAL
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                                                                        I
                                                                                                               What is Nutrition?
Figure 1-la. Cost of production budget: all costs.                                                                What exactly is nutrition? It is the science that
                                    ..
                                                                                                               studies the nutrients needed by animals: how much
                                                                                                               of each is required in a healthy diet, how they can
                                                                                                               be supplied in an economical fashion andhow the
                                    •
                                                                                                               body utilizes them for maintenance, growth and
                                                                                                               production. Nutritionists are concerned with
                                                                                                               improving the chemical analysis of feeds and
                                                                                                               feedstuffs to better reflect their true value to the
       II                                         FEED                                                    11   animal. Information is drawn from other fields of
            111111111                                 72%                                      11111111        science, such as biochemistry and physiology, to
                        11111                                                         111111                   help us understand the basic processes in the body
                             111111111111111111111111111111111111111111111111111111111
                                                                                                               that dictate the ultimate fate of the feed that animals
                                                                                                               eat. The essence of nutrition is to define the
                                                                                                               nutrients required by the animal to perform at a
Figure 1-1 b. Cost of production budget: variable
costs only.
desired level, identify a suitable source of these
nutrients and match the two (requirement and
source) in a diet formulation.
   Diets, therefore, cannot be formulated in a                Diets can not be formulated in a vacuum. Like designing
vacuum; the total circumstances surrounding the               a barn, feed formulation must consider the total manage-
                                                              ment situation including genetics, housing, health, and of
animal including its environment and the type of
                                                              course. economics.
farming operation involved must be considered. In
the same way that an engineer designs a barn with a
                                                                  The nutrients supplied by a given ingredient are
view towards the producer's management style and
                                                              sometimes difficult to determine with precision, due
his overall production objectives, the nutritionist
                                                              partly to the limitations of the chemical analyses
must formulate diets that will complement other
                                                              currently available and in part to the many factors
aspects of the farm. No single diet will suit all
                                                              that influence how available these nutrients might be
farming operations.
                                                              to the animal.
                                                              t>
                                                                   t<>
                                                                   o 0
                                                          NUTRIENTS
          QUANTITY
           EATEN                  I       t»    <>
          PER DAY                 \ 0          I> 0
                       "-.. \ <> g <>o
                                      i   t»    <> 0
                                  MIXED DIET                  INGREDIENT A               INGREDIENT B
Pig nutrition embraces three components: nutrient requirements, nutrient sources, and daily feed intake.
                                                          2
    It is also a challenge to determine the intake of         computer has led to errors in diet formulation, but
diets, since feed intake will obviously dictate               invariably these can be traced to human mistakes in
nutrient intake. The physical capacity of the gas-            programming.
trointestinal tract plays a role, particularly in nurs-
ing sows, weanling pigs and growing pigs. The                 Photo 1 - 1.
importance of feed intake in practical nutrition is
becoming increasingly apparent, in part because of
the considerable variation that exists among farms -
in the order of 30% in the grow out period, for
example. Measuring true feed intake is being
viewed as increasingly important, although it tends
to be a time consuming process.
                                                          3
The Future of Nutrition                                      growth and production. It will allow us to
                                                             formulate diets for very specific conditions, such as
   What does the future hold for the science of
                                                             the growth of lean as opposed to fat in the carcass;
nutrition? Already, the advent of biotechnology is
                                                             and to integrate environment, disease and genetics
having an effect. Reduced costs of synthetic amino
                                                             into the final ration. The costly waste of excess
acids means that we will have more options for
                                                             nutrients will be eliminated and pork production
balancing diets, and that our dependency on con-
                                                             will become a much more efficient process.
ventional protein sources such as soybean meal and
canola meal will diminish.
                                                                The rise of the environment as a key issue in pork
                                                             production, and the role of the diet in determining
   Both improved crops and superior genetic quality
                                                             the nutrient content of slurry, dictates that nutrition
of pigs will alter the way in which we formulate
                                                             and the environment will be intimately linked in the
diets. The most exciting development, however,
                                                             future. Indeed, diet formulation with a view to
will be a vastly improved ability to understand how
                                                             minimizing nutrient waste not only benefits the
the pig grows and uses nutrients. This knowledge
                                                             environment, but it often leads to reduced produc-
will be of tremendous benefit because it will expand
                                                             tion costs. It has forced us all to question bow we
our ability to match the pig's diet with its needs for
                                                             balance diets and design feeding programs, particu-
                                                             larly with respect to amino acid (nitrogen) and
                                                             phosphorus supply.
CROP \
                                                         4
 2. GASTROINTESTINAL PHYSIOLOGY
   The science of nutrition is closely linked to other            The breakdown of food into simpler and smaller
scientific disciplines, one of the most important              compounds is called digestion. Digestion not only
being physiology. Gastrointestinal physiology is               converts proteins into amino acids, but it also
the study of the stomach, the small and large intes-           breaks carbohydrates down into sugars such as
tines and related tissues. It is a subject that helps us       glucose or fructose. Fats in the diet are converted
better understand nutrition.                                   into their constituent parts, largely free fatty acids,
                                                               monoglycerides and related compounds. Minerals
The Role of the Digestive Tract                                are also separated into individual elements. Salt is
   Before feed can be used by the pig, it must first           an example. Salt is chemically known as sodium
be absorbed from the digestive tract into the body.            chloride and is broken down into its individual
From a physiological point of view, the material               molecules of sodium and chloride. Although not
contained in the digestive tract is considered to be           digested, some vitamins must be modified in the
outside the body. There are very good reasons for              gut to facilitate their transfer across the gut wall.
this. Some parts of the undigested food are actually           All of these processes are required in order to
toxic to the body; the intestinal wall acts as a barrier       prepare the nutrients for absorption. The enzymes
to prevent, or at least attempt to prevent, the entry          involved in the digestion of feed are represented
of unwanted compounds. For example, some intact                below. The dual processes of digestion and ab-
proteins can cause an allergic reaction in the animal.         sorption are discussed in more detail later in this
The proteins in the diet are therefore broken down             chapter.
into their constituent parts called amino acids,
before being allowed to enter the body.
e,oSE
Feed ingredients must be broken down in the gastrointestinal tract into individual nutrients such as protein into amino
acids or starch into glucose.
                                                           5
Types of Digestive Tracts                                      As a result of the differences in the digestive
   There is an old saying that "You are what you            tracts of cattle and pigs, their diets are also differ-
eat!" In actual fact, all animals must cat according        ent. The ruminant digestive tract lends itself to the
to what "they are". Each member of the animal               use of feeds with a high fibre content while that of
kingdom has a digestive tract that determines the           the pig utilizes more easily digested. low-fibre feeds
type of diet best suited to the animal and dictates         such as grains.
how it should be fed. For example, cattle and other
ruminant animals have a complex stomach which                  All animals with a simple stomach are not alike
includes four distinct segments each of which has a         in terms of nutrition. Chickens and pigs both have
specific function. The rumen is the largest of these        simple stomachs but differ in many areas.
and functions as a large fermentation vat containing
both bacteria and protozoa. The action of these
microbes serves to degrade many feed components
and convert them into more simple compounds.
This fermentation process allows the cow to take
advantage of the "bugs" that can break down fibre
or roughage into products that, upon entering the
intestinal tract, can be digested and absorbed.
                                                        6
       PIGS DIFFER FROM CHICKENS
        * The pig has a stronger sense of
     taste, so palatability is of greater concern
     with swine than with poultry.
                                                             The Stomach
                                            33.5%               Once food enters the stomach, the next phase of
                                                             digestion begins. The stomach serves as a large vat
                                                             in which food is mixed with acid and digestive
                                                             enzymes plus a substance called intrinsic factor.
                                                         7
   Acid secreted in the stomach serves to alter the              In summary, the stomach of the pig secretes acid,
structure of food proteins making them more                   pepsinogen and intrinsic factor which all contribute
accessible to digestive enzymes and initiating the            to the digestion and absorbtion of food. The stom-
action of certain digestive enzymes. For example,             ach also acts as a regulator of food entry into the
the stomach secretes a compound called                        small intestine, helping to prevent either overloading
pepsinogen. When pepsinogen is exposed to the                 or deprivation.
acid of the stomach, it is converted into pepsin, a
very potent enzyme involved in the digestion of                  The stomach of the pig is susceptible to ulcers,
proteins.                                                     especially in the esophageal region. Ulcers are
                                                              common in growing and adult swine, although most
   The intrinsic factor is an interesting compound            producers do not recognize the problem until death
also secreted by the stomach. It is absolutely                occurs. Symptoms of bleeding ulcers include dark
critical in the prevention of a disease called perni-         feces and anemia, both due to the loss of blood into
cious anemia, which is due to a vitamin B12 defi-             the intestinal tract. Animals with a chalky, white
ciency. The intrinsic factor must bind to the                 colour may also have a bleeding ulcer. Ulcers can
vitamin B12 supplied by the diet if the vitamin is to         impair growth in swine and in severe cases, result in
be absorbed. Supplements of the vitamin will not              death. Detailed information on ulcers is surprisingly
prevent pernicious anemia if the intrinsic factor,            limited, but it is suggested that economic losses are
which cannot be added to swine diets, is not present          greater with subclinical ulcers which reduce animal
to allow for its absorption.                                  performance, than from death loss itself.
  The secretory activity of the stomach must not be             The cause of gastric (stomach) ulcers is not really
underestimated. For example, in one study, 40 kg              known. Many factors are associated with them but
pigs were found to secrete four to eight litres of            more research needs to be carried out in this area.
total fluid per day.
                                                                      CURRENT THEORIES ON THE
  A major role of the stomach is to control the rate                     CAUSES OF ULCERS
of entry of food into the small intestine and thus
contribute to a more gradual supply of nutrients                * Some suggest that "stress" is involved but
from the gut. After a single meal, the stomach can              this has not really been studied nor quantified.
mete out digesta to the lower gut over a period of              * Diet type may be involved with wheat and
18 hours or more.                                               corn showing more of a connection with ulcers
                                                                than barley or oats. This may be due to their
   It has been postulated that in the young pig, the            different fibre contents.
formation of the 'milk clot' in the stomach is                  * Pelleted diets may also contribute to ulcers
critical in slowing the entry of food into a relatively         and yet the number of animals fed pelleted diets
immature small intestine. The problem with diets                without apparent problems suggest that it is not
containing little or no milk is that clotting is im-            the sole cause. The fine grind required to manu
paired and transit time is increased. An excessive              facture firm pellets is probably more of a factor
rate of passage of food into the small intestine                than the pelleting process itself.
overloads the system. The problem is compounded                 * Other dietary factors that may be involved
by the reduced digestive abilities of the small                 include fineness of grind, the presence of milk
intestine of the young pig. This is extremely                   proteins, high fat content, especially of unsatu
important to consider at the time of weaning. Once              rated, long-chained triglycerides and deficiencies
poorly digested material overwhelms the gut,                    in thiamine and vitamin E.
undesirable bacterial fermentation begins and                   * Ulcers also appear to be a greater problem in
scouring results. lt is recommended that recently               minimum disease herds, possibly because other
weaned pigs fed non-milk diets should be limit-fed              health problems are reduced or because animal
for five to seven days to help prevent scouring.                performance is higher.
                                                          8
Photo 2 - I.                                                   Photo 2 - 2.
Cross section of the intestinal tract, showing the villi       A more powerful microscope shows the micro-villi
extending into the lumen of the gut(top). The mus-             (top) typical of the small intestine. The dark oblong
cles of the gut, which allow it to mix and move its            structures below the micro-villi are called
contents, appear at the bottom of th� photo. The               mitochondria; they act like little power plants to
micro-vllli, too smaJI to be seen in this photo appear         drive the many activities of the gut.
opposite.
                                                           9
also help to regulate its activity. It is quite clear            The structure of the small intestine is uniquely
then that the small intestine does not work in                designed to accomplish its objectives of digestion
isolation, but is influenced by many other tissues in         and absorption of nutrients. The wal I is heavily
the body. Once food enters the small intestine from           muscled which permits the intestine to expand and
the stomach, digestion speeds up and absorption               contract in a rhythmic manner to move material
begins. More enzymes are added to the digesta, as             along its length and to assist in mixing enzymes,
well as mucous, buffers and bile. Some of the                 bile and other secretions into its contents. The
enzymes are produced locally by the intestine;                process also ensures that material ready for absorp-
others arrive from the pancreas.                              tion is brought into contact with the absorptive cells
                                                              found along the intestine surface. The constant
   Specific enzymes have specific roles to play.              mixing motion is absolutely critical if the process of
Carbohydrates, such as starches and sugars, are               absorption is to be efficient.
broken down by the action of enzymes which are
very specific with respect to the carbohydrates they             The small intestine is designed to maximize
will attack. Amylase breaks down starches such as             absorption. The surface consists of finer-like
those found in grains. Sucrase degrades sucrose               projections called villi which increase the surface
(table sugar) and lactase degrades the 'milk sugar',          area of the gut and thus increase its absorptive
lactose.                                                      capacity. Along the villi are further projections,
                                                              called microvilli which also increase surface area
   Proteins are digested by a totally different set of        and thus, the absorptive capacity of the small
enzymes. Proteins must be broken down into                    intestine.
amino acids before crossing the intestinal wall.
Protein digestion begins in the stomach but be-              Another interesting feature of the small intestine
comes much more effective in the small intestine.         is the way in which the absorptive cells grow. They
Examples of protein-degrading enzymes include             are called enterocytes and are formed at the base of
trypsin, chymotrypsin, elastase and                       the villi and then migrate up along its surface as
carboxypeptidase. The protein-degrading                   they mature. The maturation process is important
(proteolytic) enzymes attack proteins at very spe-        because the more mature these cells are, the greater
cific locations in their structure.                       is their content of digestive enzymes. The migra-
                                                          tion of enterocytes along the surface of the villi
   Fats in the diet must also be digested before          from the base to the tip, ensures that the mature
being absorbed. An enzyme called lipase, derived          cells receive maximum exposure to the contents of
from the work lipid which is another name for fat,        the intestine; meanwhile, the immature cells located
breaks fats into smaller parts which are then com-        at the base of the villi are protected rom the physi-
bined with bile salts to form a compound that can         cal forces of food moving along the length of the
cross the intestinal barrier. Bile salts are produced     intestine. Once the enterocytes reach the tip of the
in the liver and are an absolute necessity for proper     villi, they are sloughed off into the lumen of the
fat digestion.                                            intestine, to be digested and absorbed. These
                                                          sloughed off cells, together with enzymes secreted
    lt can be seen that a very competent system has       into the intestine, constitute what are cal1ed endog-
been devised to allow the pig to digest the various       enous protein. This endogenous protein has been
components in the diet. Failure of any part of this       estimated to represent anywhere from 25 to 75% of
system will result in impaired utilization of food. It    the protein digested and absorbed by the gut. The
is a complex system, with details far beyond the          diet makes up the remainder.
scope of this book. lt is important, though, to
recognize that digestion is not a simple matter and             The cells have a very brief existence even under
that what and how we feed the pig, how we house               conditions of good health. The life expectancy of
it, how old it is and how healthy it is can all influ-        an enterocyte in a one week old piglet is only two to
ence the final outcome of the digestive processes.            four days. Sometimes the enterocytes do not reach
                                                         10
THE SMALL INTESTINE OF THE PIG                                                            BLOOD VESSELS
         �����2----/
                                                                                              Moving through
                                                                                              the lumen, feed Is
                                                                                              digested and absorbed
                                                                                              Into the body of the
                                                                                              pig. Muscles surround
                                                                                              the Intestinal tissue;
                                                                                              their contraction
                                                                                              moves the dlgesta
                                                                                              along the tract and
                                                                                              helps mix In digestive
                                                                                              enzymes.
                                                                Folds in the
                                                                intestine are lined
                                                                with projections called
                                                                villi. The villi
                                                                increase the surface
                                                                area, thus enhancing
                                                                both digestion and
                                                   ,            absorption.
                                                       -,
                                                            '>1--.
                                                       '�"-.
                                                                                             DISCARDED CELLS
                                                   ����' �<',<
                                                     ."�" , " ' '\..o
                                                               $�
                                                                 ":/    o   0}
                                                                                 -,
                                                                                             !1
       The individual villus consists of
       cells that form at the base and migrate
       to the tip, where they are sloughed off
       Into the lumen of the intestine. As the
       cells migrate they mature, Increasing
       their ability to break-down and absorb
       nutrients from the lumen of the gut. It
       takes a cell 2-5 days to move from the
       base to the tip. Each day, literally
       billions of cells are discarded along the
       length of the pig's gastrointestinal
       tract. The villi are also the site of
       absorption, where nutrients pass
       through the cells to enter the blood
       stream.
                                                       11
maturity, but are sloughed off into the intestinal
lumen prematurely. As a result, the 'oldest' cells in
the system arc still immature. If this happens, the
digestive ability of the small intestine is impaired
due to the absence of mature cells and their rich
                                                               "'..
                                                               CD
                                                               .c
                                                                      100
                                                               0      80
source of digestive enzymes.                                   Ill
                                                               .c
                                                               <      60
                                                               ell
    Several things have been shown to influence the
                                                               'i     40
lifespan of absorptive cells. Research suggests that           iii
                                                               CD
one of the factors responsible for the post-weaning            Cl
                                                               0      20
ple, even a few days off feed wiU lower the rate of
cell turnover and impair the intestine's ability to            Figure 2-3. Nutrient Digestion Along the Gastrointes-
                                                               tinal Tract of the Pig.
produce new cells.
                                                          12
by the large intestine. lf the vitamins are excreted             impair this activity. Simply stated, for best per-
in the feces and re-eaten, they will then contribute             formance, the pig should be fed a diet that is suited
to the vitamin nutrition of the pig. Some people                 to its digestive abilities. Since these abilities change
believe that pigs housed on fuJly-slatted floors                 with age, the better quality and more expensive
which have Jess exposure to manure, have a higher                ingredients should be used with the younger pig
requirement for dietary vitamins. There is very                  than with the adult sow or boar.
little scientific evidence to support this. The pru-
dent approach is to ensure that the diet is properly
supplemented with preformed vitamins to meet the
requirements of the pig. In this way, the pig is not
                                                                 Additional Reading and References
                                                                 Lloyd, L.E., B.E. McDonald and E.W. Crampton.
dependent on microbial fermentation to supply even
                                                                 1978. Fundamentals of Nutrition. W.H. Freeman
a portion of its vitamin requirements.
                                                                 and Co., San Francisco. 466 pp.
    Some of the information in Figure 2-2 must be
interpreted with great care. It suggests that nitrogen,          Low, A.G. 1989. Research into the digestive
                                                                 physiology of pigs. In. (E.J. van Weerden and J.
and thus protein, is absorbed by the large intestine.
It is true that nitrogen is absorbed, but unfortu-               Huisman, eds.) Nutrition and Digestive Physiology
                                                                 in Monogastric Farm Animals. pp. 1 - 15, Pudoc,
nately, not as amino acids. Thus, if an essential
                                                                 Wageningen.
amino acid is not absorb from a feed by the end of
the small intestine, it will not be absorbed intact.
The large intestine does not have the ability to                 Moran, E.T., Jr. 1982. Comparative Nutrition of
                                                                 Fowl and Swine: The Gastrointestinal System,
absorb essential amino acids but rather absorbs
                                                                 University of Guelph, Guelph.
nitrogen as a simpler compound called ammonia.
Nutritionists who are measuring the proportion of
lysine or other amino acids digested from a given                McDonald, P., R.A. Edwards and J.F.D.
                                                                 Greenhalgh. 1973. Animal Nutrition. Oliver &
feedstuff will collect digesta as it leaves the small
                                                                 Boyd, Edinburgh.
intestine rather than collect the feces. This is done
quite painlessly by surgically placing a collecting
tube in the intestinal tract of the pig. If nutritionists        Pond, W.G. and K.A. Korpet. 1978. The Biology
                                                                 of the Pig. Cornell University Press, Ithaca.
use feces to determine amino acid digestibility, their
                                                                 371 pp.
results will be incorrect. The bacteria in the large
intestine break down lysine and other amino acids
and are also able to manufacture amino acids from                Whittemore, C.T. 1987. Elements of Pig Science.
non-protein nitrogen.                                            Longman Handbooks in Agriculture, Harlow.
                                                                 181 pp.
                                                            13
14
3. NUTRIENTS
    Although diets are mixed using ingredients,                    The essential nutrients can be divided into
what really matters to a pig are the nutrients                 categories of minerals, amino acids, vitamins, fatty
contained in those ingredients. The relative                   acids, energy, and water (Table 3-1 ). If a diet is to
proportions of barley, wheat, soybean meal, and                be properly balanced, the nutritionist must know the
other feedstuffs in a diet are determined on the basis         pig's requirement for each nutrient and the amount
of the nutrients they provide to a pig. When a                 supplied by each ingredient.
nutritionist formulates a diet for pigs, no less than
41 essential nutrients are considered.                         Table 3-1. Essential Nutrients in the Diet of a Pig.
    In this chapter, the utilization of nutrients by           Amino Acids Minerals            Vitamins       Other
pigs and means to estimate the available nutrient
levels in pig feed ingredients are discussed. For              Arginine      Macro             Vitamin A Energy
information on nutrient allowances in diets for the            Histidine     Calcium           Vitamin D Linoleic
various classes of pigs, the reader is referred to             Isoleucine    Chloride          Vitamin E    Acid
chapters 6 - 9. Water, which can also be classified            Leucine       Magnesium         Vitamin 812 (Omega-6
as a nutrient, is addressed in chapter 12.                     Lysine        Phosphorus        Biotin       fatty acids)
                                                               Methionine Potassium            Choline     Water
                                                               Phenylalanine Sodium            Folic Acid
What is a Nutrient?
                                                               Threonine     Sulphur           Menadione
    Knowing the difference between a nutrient and
                                                               Tryptophan                      Niacin
an ingredient, or feedstuff, is critical when
                                                               Valine        Micro             Pantothenic Acid
formulating a diet. A nutrient is a chemical
                                                               Cystine*      Cobalt            Pyridoxine
substance that is supplied by the diet and which pigs
                                                               Tyrosine"     Copper            Riboflavin
need to stay alive. Nutrients perform specific
                                                                             Iodine            Thiamine
functions in the body and their absence or
                                                                             Iron
deficiency will result in reduced productivity, health
                                                                             Manganese
problems and, possibly, death.
                                                                             Selenium
                                                                             Zinc
     An ingredient or feedstuff is the material used to
supply nutrients. For example, barley and com are
                                                               * These amino acids are considered semi-essential
ingredients which contain amino acids, energy, and             nutrients because they can be derived from methionine
other nutrients. They can supply a pig with some of            and phenylalanine, respectively. Other nutrients, such as
the nutrients it needs. A balanced diet mixes                  vitamin C, linolenic acid, fluorine, chromium, nickel, lead
ingredients, such as com or barley, that supply the            silicon, tin, and vanadium, may be essentiaJ in a pig's diet,
                                                               however, deficiencies are extremely unlikely and/or
nutrients in proper proportions. The pig has no
                                                               requirements for these nutrients have not been clearly
specific requirement for individual ingredients - it           established.
does not need barley or com, specifically, in its diet,
for it can obtain the necessary nutrients from other           Nutrient Availability
ingredients, such as canola or soybean (or in some
                                                                  Availability refers to the pig's ability to digest,
countries, even green bananas!). A pig may prefer
                                                               absorb, and utilize nutrients from a given
to eat one ingredient over another, but ingredients
                                                               ingredient. Very few nutrients are 100% available.
have no unique role in a pig's diet. Selecting the
                                                               A nutrient may be present in an ingredient but not
type of feedstuffs to supply nutrients is much less
                                                               available to the pig for both mechanical (e.g.,
critical than making certain the necessary nutrients
                                                               processing methods) and physiological (e.g.,
are supplied from the feedstuffs chosen.
                                                               digestive functions, age, state of health) reasons.
                                                               Defining the availability for each nutrient in each
                                                               ingredient is a difficult task because availability
                                                          15
varies from ingredient to ingredient and is
influenced by many factors.                                    :c
                                                               g_
                                                                     .
                                                                     5
                                                                     3
   How docs the nutritionist work around this
problem? In many cases, sufficient information is
                                                              ••
                                                              a:     2
available to permit at least an educated guess. Diets          CII
                                                              a:     1
will often include more than one basal grain and
more than one protein supplement. Thus, the                          o..L.�---�------.....-..1,--.--r--.-....,....-r....,............,
                                                                         o         2            4A          s           s           10
nutritionist 'hedges his position' across a number of
                                                                                   Nutrient Level In the Diet
ingredients, such that underestimating one might be
compensated by overestimating another. This                  * As the requirement (point A) is approached, the rate of
approach is not a totally desirable one, but until           improvement per unit of nutrient decreases.
more precise data becomes available, it is probably
                                                             Figure 3-1. Pig Response to Level of Nutrient in the
the most practical one.
                                                             Diet.
Nutrient Requirements
    What is meant by the word 'requirement' from a              Thus, requirement defines the level needed to
nutritionist's perspective? Traditionally, a                 achieve maximum performance but not necessarily
requirement has been defined as the amount of a              maximum profit. Each situation must be evaluated
given nutrient a pig requires to maximize                    to determine the best (most profitable) feeding
performance. Producers need to understand the                strategy.
criteria used to determine requirements because
how the requirements are determined can influence           Although we would all like to have neat, simple
the way producers use nutritional information.           tables defining an absolute value, they are not
                                                         realistically possible. Some compromise must
    One of the frustrations in defining a requirement    occur. The compromise does not diminish the value
is that maximum performance may occur at                 or importance of nutrient requirement tables, it
different nutrient levels depending on the response      merely emphasizes the importance of understanding
criteria being considered. For example, maximum          their origin. Three excellent publications that
growth rate may require more or less of an amino         provide useful requirement guidelines are the
acid than maximum feed efficiency. The amount of         National Research Council's Nutrient Requirements
a given nutrient required to maximize the immune         ofSwine (1988), the Agricultural Research
response may differ from the amount required for         Council's The Nutrient Requirements ofPigs
maximum feed intake. If carcass merit is used to         (1981), and the Australian Agricultural Council's
evaluate nutrient requirements, conclusions may          Feeding Standards for Australian Livestock - Pigs
differ from those reached using growth rate.             (1987).
                                                        16
into forms the cell can use for various purposes.                energy is called net energy (NE). NE is the best
Energy, due to the amount required, is generally the             estimate of the amount of useful energy that is
most expensive 'nutrient' in the pig's diet.                     supplied by feed ingredients.
Consequently, nutritionists focus considerable
attention on energy in feed formulation and feeding              Metabolizable Energy - Heat Increment= Net
management in order to minimize feed cost without                Energy
compromising performance.
Energy Components
   The total quantity of energy contained in a                                           Gross Energy
feedstuff can be partitioned into different                                                   I
components based on how the pig will use the                                   I
                                                                         Fecal energy
                                                                                                            I
                                                                                                  Digestible energy
energy (Figure 3-2). If an ingredient, such as barley
or wheat, is burned completely, the amount of                                                               I
energy released is called gross energy (GE). The
gross energy content can be determined by the                           Urinary energy              Metabolizable energy
bomb calorimeter. GE is thus the total amount of
energy contained in the grain; if the pig could digest
and utilize 100% of the energy, that is how much
                                                                                                            I
                                                                       Heat increment                    Net energy
would be available. The pig, however, digests or
absorbs only a portion of the gross energy. The
                                                                         Maintenance
                                                                                                            I
                                                                                                        Production
energy absorbed by the intestinal tract and actually
available to the pig is called the digestible energy
(DE). The DE content of any feedstuff can be
                                                                 Figure 3-2. Distribution of Dietary Energy.
determined by measuring the quantity of food
(energy) consumed and subtracting the energy lost
in the feces (calculated by weighing the feces and
analysing their composition).                                       Some scientists have attempted to use this
                                                                 information on heat increment to practical
                                                                 advantage by designing diets around heat
Gross Energy - Fecal Energy = Digestible Energy
                                                                 increments. The pig is a warm-blooded animal and
   _Some of the digestible energy is lost in the pig's           must maintain a constant internal body temperature
unne or released as gas from its gastro-intestinal
                                                                 irrespective of the temperature of its surroundings.
tract. Much of the energy lost in the urine results              Normal metabolism generates heat that is used in
from the excretion of unneeded nitrogen                          body temperature regulation. If a pig is chilled, it
compounds, such as urea. The amount of digestible                must eat more feed to keep warm or else it will lose
                                                                 weight or gain it at a slower rate. The heat
energy remaining in the pig after these losses is
called metabolizable energy (ME).                                increment is used to provide some of this required
                                                                 heat, so heat increment in a diet is desirable. The
Digestible Energy - (urine energy loss+ gas energy               pig will need less heat to maintain a constant body
loss)= Metabolizable Energy                                      temperature in hot weather and will actually lower
                                                                 its feed intake to reduce the production of metabolic
   A portion of metabolizable energy is lost when                body heat if it is in danger of becoming heat
the pig uses the nutrients in the feed as heat, i.e. heat        stressed; in this case, heat increment in the diet is
losses during eating, digestion and actual nutrient              detrimental. Obviously, in these two instances, pigs
metabolism. This portion is called the heat                      will require diets with different heat increments.
increment. Once the heat increment is used up out                The quantity of heat increment varies depending on
of ME, the energy actually available to the pig for              the nature of the diet.
maintenance and growth is left. This amount of
                                                            17
   The solution proposed by some scientists is to             There are, however, some important differences in
feed a diet high in heat increment (rich in fibre)            DE content, both expressed in absolute values as
when the pig is likely to be chilled. When the pig is         well as in fractions of GE content. In fact, barley
likely to be heat stressed, a diet low in heat                contains more GE but Jess DE than com. This
increment (rich in fat) is recommended. Under                 clearly indicates that it is more accurate to
Canadian conditions, this feeding approach means              formulate pig diets based on a DE basis rather that
feeding a high-fibre diet to dry sows in chilled              on a GE basis.
conditions, such as in outside housing in winter or
in a cool gestation barn (especially barns equipped              The proportion of digested energy lost in the urine
with individual stalls where sows cannot huddle to            is fairly constant for all grains and all protein
keep warm). A high fibre diet is never fed to                 sources, but differs between these classes of
weanling pigs, lactating sows, or growing pigs                ingredients. This difference is due in part to the
because growth rate would suffer. In the heat of              urinary energy lost during excretion of nitrogenous
summer, lactating sows and growing pigs should                waste products. Because of the differences between
receive a diet rich in fat to help maintain energy            protein sources and grains, there are theoretical
intake. Of course, these suggestions must be                  advantages to using ME values in preference to DE
considered in the context of cost and economic                values to formulate feed. However. note that many
benefit.                                                      of the published ME values have merely been
                                                              derived mathematically from DE. Direct assays
  Although net energy is the best estimate of how             (analyses) of ME are difficult since measuring the
much useful energy is supplied by a diet, it is rarely        amount of energy excreted in the urine poses a
used as a system for formulating swine diets in               number of logistical problems. Whether DE or ME
North America because of the difficulty in routinely          is used, be certain that both requirement and
and accurately determining the heat increment of              nutrient values are expressed in the same system.
feeding when different ingredients are fed to pigs.             The values in Table 3-2 demonstrate that the
NE systems are used more commonly in European                 differences in NE content between feedstuffs tend
countries, where a larger variety of ingredients are          to be larger than the differences in DE content. For
used to formulate swine diets.                                example, in a DE system. corn contains 12% more
                                                              energy than barley. In a NE system, this difference
  Table 3-2 shows the GE, DE, ME, and NE                      increases to 15%. For ingredients with extreme
content in selected pig feed ingredients. Most of             compositions such as alfalfa and wheat shorts (high
the ingredients listed are very similar in GE content.        in fibre) or soybean oil (high in fat), these differ-
Table 3-2. Proportions of Gross Energy (GE), Digestible Energy (DE), Metabolizable Energy (ME) and Net
Energy (NE) in a Sample of Ingredients.
Derived from: Ewan, 1989; Energy metabolism of farm animals. EAAP publication No .. 43. Pudoc Publishers,
Wagcningcn, The Netherlands.
                                                         18
ences are even more apparent. This again illustrates       hemicellulose, and lignin, which are essentially
that, in theory, it is more accurate to formulate diets    indigestible by swine. ADF consists only of
using an NE rather than DE or ME system.                   cellulose and lignin. ADF and NDF were
                                                           developed to evaluate forages for ruminant species.
  Energy is measured in units of either calories (cal)     They have also been found valuable by swine
or joules (J). The term used in this book is the           nutritionists but tend to underestimate the total fibre
kilocaloric, equal to 1000 calories. The joule is          of cereaJ grains.
more common in Europe and is becoming more
popular in Canada, but is not yet universally
                                                           Table 3-3. Fibre Content of Ingredients Used in Swine
accepted. One calorie equals 4.184 joules.
                                                           Diets.
  Defining the term fibre has been a source of Adapted from: Graham 1988. Anim. Plant Sci. 1 :76-80.
controversy for decades. For the purposes of this
book, fibre is defined as the portion of carbohydrate
in a diet that cannot be digested by the enzymes              There are large differences in the values obtained
produced by a pig. Pigs, like ruminants, rely on by these measurements, although they all attempt to
bacteria that are present in the digestive tract to utilize define the same component of feed ingredients -
dietary fibre. The pig does not have a requirement indigestible carbohydrate. Most Canadian feed
for fibre per se, although it provides a texture in diets testing laboratories report crude fibre, although acid
that helps digestive processes. Fibre is used to evaluate detergent fibre and neutral detergent fibre
ingredient quaJity because it is associated with reduced measurements are gaining popularity.
DE content and impaired utilization of other important
nutrients. Therefore. from a nutrient perspective, fibre      The ether extract (fat) content of a feedstuff or
has a generally negative impact on animal feed quality. diet is important because fat is rich in energy. All
                                                            other factors being equal, the higher the fat content
   Measuring fibre in feed ingredients is an area of in a diet, the more digestible energy it will contain.
active research. Table 3-3 defines the fibre content of Determining fat content provides some insight into
some common ingredients. The complex nature of how much energy is present. Fat content is
fibre is illustrated by the number of fibre values estimated by determining the portion of the sample
defined. Dietary fibre is a Swedish tenn that includes that is soluble in petroleum ether. This procedure is
lignin (a highly indigestible carbohydrate found in a standard one, but misses some important
such feedstuffs as straw) plus all complex sugars not compounds. such as energy-rich phospholipids. In
found in starch.                                            barley, for example, these phospholipids represent
                                                            as much as 25% of the total fat in barley grain and
  The detergent system, developed by P.J. van Soest contribute to its total energy.
in the United States, differentiates between two
kinds of fibre: acid detergent fibre (ADF) and                Ether extract will also include such components
neutral detergent fibre (NDF). NDF includes                 as waxes and pigments, which are of little
material in the plant cell wall, notably cellulose,         nutritional value. For this reason, the digestibility
                                                          19
of the ether extract varies among grains as they vary    Estimating Energy Contents in Feeds and
in the relative proportion of phospholipids and          Feed Ingredients
waxes (Table 3-4 ).                                          Evaluating the content of energy and other
                                                         nutrients in feed ingredients is an important step in
   What is the solution? Some nutritionists use          formulating diets for swine. Without knowing the
di ITcrent solvents in place of petroleum ether to       amounts of nutrients supplied by individual
extract the fat of feed grains thus developing a more    ingredients, one cannot, with any degree of
complete analysis. Common alternative solvents           certainty, put together diets that will satisfy the pig's
include diethyl ether, and a 2: 1 mixture of             needs. Unfortunately, there is a considerable
chloroform and methanol. Different solvents arc          amount of variation in available energy contents
used to answer different questions. As in most           between different samples of one feedstuff. For
analyses, there is no complete test for all answers.     example, recent research at the Prairie Swine Centre
                                                         demonstrated that DE content varied by as much as
Table 3-4. Digestibility of Ether Extract of              l 0% between various samples of substandard
Common Feedstuffs.
                                                         wheat, with the poorest sample similar to that in
                                                         regular barley. This demonstrates the use of
                                     Percent
                                                         average book values is not sufficient to estimate
Feeds tu ff                        Digestibility
                                                         nutrients in specific samples of feed ingredients.
Alfalfa meal                             54
Barley                                   86                However, it is simply too expensive and time-
Com                                      80              consuming to evaluate the nutritional value of
Soybean meal                             80              feedstuffs in feeding trials. There is thus a need for
Meat meal                                93              rapid, inexpensive, and practical tests for use in
Wheat                                    52              day-to-day diet formulation. Laboratory (chemical)
Wheat bran                               58              evaluation can be of great assistance in meeting this
                                                         need. World-wide research has attempted to
Adapted from: Animal FeedstuffTable. 1991. Centraal      develop chemical and alternative tests that give the
Veevoederbureau, Runderweg 6, 8219 PK Lelystad, The      true nutrient composition of ingredients and mixed
Netherlands.                                             diets, and accurately reflect the true feeding value.
                                                        20
DE (Kcal/kg of dry matter)*= 4168 - (9. lx Ash)+                  Table 3-5. Effect of Bushel Weight on Nutrient
( 1.9 x Crude Protein)+ (3.9 x Ether Extract) - (3.6              Content of Cereal Grains.
x NDF)**
                                                                     Bushel         Crude         Crude         Ether
*The contents of the chemical constituents in the diet are           Weight         Protein       Fibre        Extract
expressed in grams per kg of dry matter.                             kg (lb)                      -%-
**NDF - Neutral Detergent Fibre
                                                                     23 (5 I)         9.2           8.5          2.4
                                                                     22 (49)          10.4          7.6          1.8
  In Chapter 4, various equations will be presented                  20 (45)          10.6          8.9          2.5
that can be used to predict DE content in specific                   19(42)           11.0          9.6          2.5
samples of main feed ingredients.
  A common misconception is that bushel weight                      Table 3-6 offers guidelines for minimum bushel
is a good indicator of a cereal grain's nutrient                  weights of cereal grains destined for use in swine
quality, and DE content in particular. Premiums are               diets.
sometimes paid for grains that exceed normal
bushel weights, e.g., 23 kg (50 lb) barley or 26 kg
(58 lb) com. Unfortunately, like many of the .                    Table 3-6. Minimum Recommended Bushel Weights
factors used in the grading of grains, bushel weight              for Grains Used in Swine Rations.
is not a good indicator of feed value. Test weight
can be affected by many factors, such as the shape                   Grain               Bushel Weights (lb)
and surface dimensions of individual kernels, which                                  Standard      Minimum
do not necessarily reflect nutrient content.
                                                                     Barley             48                43
                                                                     Com                56                50
  Research results show very clearly that energy
                                                                     Wheat              60                55
concentration of grains does not fall in proportion
with bushel weight. The energy content of typical                 Guidelines for minimum bushel weights of grains used in
50 lb of barley is no different than that of 48 lb or             swine diets. [f grains below these standards are used,
even 46 lb of barley. In a recent study at Prairie                adjustments in energy content should be made.
Swine Centre, there was no relationship between
DE content and bushel weight in wheat samples that
                                                                  Energy Requirements
varied in density between 53 and 63 lb/bushel. In
                                                                    A pig requires energy for almost all its body
the case of very low bushel weights, energy values
                                                                  processes. Amino acids cannot be converted to
do appear to fall; therefore, very low bushel weight
                                                                  muscle proteins and feed cannot be digested or
grains should be used only in diets where energy
                                                                  wastes eliminated without energy. Body
concentration is not critical (gestating sows or
                                                                  temperature cannot be maintained , gestating sows
growing pigs over 60 kg body weight) and even
                                                                  cannot produce a fetus, nor can nursing sows
then , the concentration of other nutrients needs to
                                                                  produce milk without energy. Clearly, energy is
be adjusted to reflect changing energy levels.
                                                                  fundamental to all life.
  The rise in crude fibre coinciding with falling
                                                                    A pig's requirement for energy will be determined
bushel weight is often put forward as an argument
                                                                  by a number of factors. The pig's size is important
favouring the use of bushel weight as an indicator
                                                                  because energy needed for maintenance is directly
of grain quality. However, the rise in crude fibre,
                                                                  related to body size. It 'costs' more in terms of
which is low in energy, appears to be offset by a
                                                                  energy to maintain a 250 kg sow than a 180 kg sow.
similar rise in fat, which is high in energy (Table 3-
                                                                  The pig's productive state is also an important
5). Interestingly, bushel weight has been used as an
                                                                  factor. A lactating sow requires more energy than a
indicator of flour yield, but even this relationship is
                                                                  gestating sow since she is producing large quantities
suspect.
                                                                  of milk. A pig that is gaining weight requires more
                                                             21
energy than one that is not growing. As described            on growth, assuming normal ingredients are used.
previously, the environment in which a pig is                Because of limitations in gut capacity relative to
housed is also critical. In cold temperatures or wet,        energy needs, diets with a high concentration of
drafty conditions, energy required to maintain body          energy should be fed to younger pigs and lactating
temperature rises. Jf pigs can huddle with each              sows. These are called high nutrient density diets.
other, their energy requirements in cold weather are         Lower energy grains, such as oats, are too low in
lower than if they arc penned individually.                  energy to support maximum growth rate.
  The amount of energy required in feed is                     Refer to the specific chapters on feeding various
determined by considering both quantity offeed               classes of pigs for further information on energy
consumed per day and amount of energy required               requirements. Any general requirement must
per day. This principle is important. For example,           consider many variables, such as environmental
a man working at a job that involves considerable            temperature, desired performance, body size, and
physical exertion has a high energy requirement per          some score of genetic capacity.
day. In theory, this requirement could be met by
eating an enormous quantity of lettuce or other              Amino Acids
bulky, low energy, foods. In practice, this approach           Amino acids are the building blocks of protein,
will not work because the man simply cannot                  meaning that all proteins are made up of individual
physically consume enough lettuce to meet his                amino acids linked together like beads in a
energy needs. If he is given a diet of meat and              necklace. In nature, there are about 22 amino acids
potatoes, i.e., high energy foods, he can easily             that can link together to form proteins. Pigs can
consume enough food to meet his daily energy                 produce some of them from other substances; but
needs. The message here is that gut capacity plays           they cannot synthesize 10 amino acids so they must
an important role in formulating diets.                      be provided by the diet. These 10 amino acids are
                                                             called the 'essential amino acids' (Table 3-1). If
  The same principle applies to pigs. If the                 they are not present in the diet, or are present in
concentration of energy in the diet is too low, the          insufficient quantities relative to need, the animal
pig may be unable to consume sufficient amounts to           will be unable to grow properly.
meet its energy needs. A gestating sow is fed less
than her appetite demands to prevent excess weight             In addition to the 10 essential amino acids, there
gain. Her stomach capacity is more than sufficient           are two so-called semi-essential amino acids,
to meet her energy needs with most practical diets.          cystine and tyrosine, that can be synthesized only
Conversely, the lactating sow has a very high energy         from essential amino acids, methionine and
requirement to support milk production and                   phenylalanine, respectively. The term 'total sulphur
maintain her body condition in preparation for the           amino acids' (T.S.A.A.), refers to the sum of
subsequent breeding period. If she is fed a low              methionine plus cystine. Pigs require T.S.A.A. in
energy diet, she will be unable to consume                   addition to the requirements for methionine. This
sufficient energy per day, will lose body weight, and        requirement indicates the importance of monitoring
produce less than her maximum potential of milk.             both methionine and cystine levels in swine feeds.
                                                             At least 50-55% of the total T.S.A.A. requirements
   Pigs up to about a 60 kg body weight ( 130 lb)            must be supplied by methionine. In the same
being fed traditional ingredients may be in danger           manner, phenylalanine must supply approximately
of growth retardation because their gut capacity is          55% of the animals' requirements for phenylalanine
insufficient to accommodate the energy required for          plus tyrosine. The sum of phenylalanine and
them to grow quickly. In pigs with extremely high            tyrosine is also referred to as "total aromatic amino
(lean) growth potential, such as boars, energy intake        acids". Phenylalanine and total aromatic amino
may limit growth up to higher body weights.                  acids are rarely deficient in practical diets.
However, in most finishing pigs of 60 kg or heavier          Consequently, under practical conditions, tyrosine is
body weight, gut capacity no longer places a limit           much less important than cystine.
                                                        22
   When the pig eats protein as part of its diet. the        Estimating Protein and Amino Acid
intestinal tract breaks down protein into individual         Contents in Feeds and Feed Ingredients
amino acids. These amino acids are then                    The crude protein content is an estimate of the
transported into the blood and carried to various        amount of protein present in a feed ingredient or
tissues where they are used for a variety of             mixed diet. Since amino acid analysis is very
purposes. The most obvious function is to form           expensive and time-consuming, crude protein has
muscle protein. However, a host of other proteins        been adopted as a more practical indicator of feed
in cells are just as important since they support a      value. Direct analysis for amino acids will become
wide range of essential body functions. Enzymes,         a much more common procedure in the future.
such as those that help to digest food in the gut, or
those that help ensure that overall metabolism in              Crude protein is usually measured using the
tissues flows smoothly, are proteins. There are also         Kjeldahl method. According to this very old, but
carrier proteins that transport nutrients from one           simple and accurate method, feed is digested in acid
side of the cell wall to the other. Haemoglobin in           and the quantity of total nitrogen in the sample is
the blood is a protein; it transports oxygen from the        determined. The total nitrogen value is then multi-
lungs to the tissues to support cell metabolism. The         plied by the 'Kjeldahl' factor of 6.25 to generate the
blood also contains many other proteins, such as             crude protein value.
those which transport certain vitamins or minerals.
Milk contains proteins to nourish newborn piglets.             The crude protein content of feedstuffs must be
Immunoglobulins circulating in the blood protect             interpreted with great care for three main reasons:
animals against specific forms of disease.                   The true Kjeldahl factor varies from ingredient to
                                                             ingredient; not all of the nitrogen in a feed sample is
   Although protein synthesis is important,                  associated with true protein; and total protein is a
individual amino acids also perform other                    very rough estimate of the amino acids present.
functions. For example, tryptophan is converted
into serotonin, a chemical called a neuro-                     The first point The true Kjeldahl factor varies
transmitter, that is involved in the transmission of         from ingredient to ingredient. The Kjeldahl factor
signals by the nervous system. It is this function           of 6.25 assumes that the protein in the sample
that has prompted some people to recommend                   contains 16.0% nitrogen. This assumption is not
tryptophan supplements as a calming influence in             necessarily true. As mentioned previously, protein
the diet. Tryptophan can also be converted to the            consists of individual amino acids. The proportion
vitamin niacin. thereby reducing, but not                    of nitrogen in each amino acid varies from 7.7 to
eliminating, the need for niacin in the diet.                32.2 (Table 3- 7) so the actual amount of nitrogen
                                                             present in a protein will depend on the relative
                                                             proportions of each amino acid in the protein. As
                                                             the proportion of amino acids change, so will the
                                                             average proportion of nitrogen in the final protein.
                                                             Com protein contains I 6.0% nitrogen while the
                                                             protein in wheat or barley contains about 17.2%
                                                             nitrogen. Therefore, the correct factor for wheat
                                                             and barley is 5.83. Using 6.25 for these two cereals
                                                             will overestimate protein content by about 7%.
                                                        23
Table 3-7. Nitrogen Content of Amino Acids.                The second point: Not all of the nitrogen in a
                                                         feed sample is associated with true protein. Some
Amino Acid                     % Nitrogen                of the nitrogen in feed ingredients is in the form of
.Methionine                    9.4                       non-protein nitrogen. There are compounds that
Cystine                        11.6                      contain nitrogen but are not protein. Including them
Lysine                         19.2                      in the crude protein calculation will overestimate
Tryptophan                     13.7                      the quality of the ingredient. The information in
Phenylalanine                  8.5                       Table 3-8 summarizes these two points. It provides
Leu cine                       10.7                      the 'corrected' Kjeldah1 factor, two crude protein
lsoleucine                     10.7                      values (one obtained from the standard Kjeldahl
Threonine                      11.8                      factor, one from the corrected factor) and the
Valine                         12.0                      corrected crude protein value adjusted for non-
Histidine                      27.1                      protein nitrogen.
Arginine                       32.2
Glycine                        18.7                        The third point: Total protein is a very rough
Asparagine                     21.2                      estimate of the amino acids present. Total protein,
Aspartic Acid                  10.5                      even adjusted for non-protein nitrogen and using the
Serine                         13.3                      corrected Kjeldahl factor, is not a perfect indicator
Glutamine                      19.2                      of amino acid content. For example, in cereal
G lutamic Acid                 9.5                       grains lysine contributes to approximately 4% of
Pro line                       12.2                      corrected protein, while in soybean meal this value
Alanine                        15.7                      exceeds 7.5%. As lysine is generally the first
Tyrosine                        7.7                      limiting amino acid in pig diets, this means that
                                                         protein in soybean meal is close to two times more
                                                         valuable than cereal grains in meeting the pig's
                                                         amino acid requirements.
Table 3-8. Protein Estimation Adjusted for Corrected Nitrogen Factors and Non-protein Nitrogen Content.
Source: unknown.
                                                    24
  We do not suggest eliminating the crude protein            Com (N= 153 samples)
measure since it is a simple, rapid, and inexpensive
test which is required to provide an estimate of             % Lysine=% CP x 0.0224 + 0.057; r =0.64
                                                             % Methionine=% CP x 0.0192+0.015; r = 0.62
protein quality. However, failure to consider the
                                                             % T.S.A.A. = % CP x 0.0345 + 0.073; r = 0.56
underlying principles could lead to serious errors in
                                                             % Threonine= % CP x 0.0336 + 0.014; r = 0.84
diet formulation. For greatest precision, crude              % Tryptophan = % CP x 0.0026 + 0.041; r = 0.41
protein should be used in conjunction with
estimates of amino acid composition.                         Soybean Meal (N=277 samples)
  The pig requires amino acids that make up                  % Lysine = % CP x 0.0665 + 0.252; r - 0. 70
protein. It is possible to analyse ingredients for           % Methionine=% CP x 0.0 I I + 0.127; r = 0.44
their amino acid content but it is an expensive              % T.S.A.A. = % CP x 0.0255 + 0.157; r = 0.52
                                                             % Threonine=% CP x 0.0344 + 0.203; r = 0.65
procedure that costs as much as $150 per sample.
                                                             % Tryptophan = % CP x 0.0144 + 0.041; r = 0.62
How then can diets be formulated on the basis of
amino acids, when such anaJyses are so expensive?
                                                             Canola MeaJ (N=57 samples)
                                                        25
protein and less the profile of nutritionally superior        Table 3-1 O. Effect of Increasing Total Nitrogen (Crude
albumens and globulins. Thus. lysine decreases as a           Protein) on the Lysine Content of Wheat and Oats.
proportion of the total. Oats are an exception to this
                                                              Crude Protein      Proportion of Lysine in Total Protein
rule, since the prolamine protein proportion in oats is
                                                              in Grain           (Percent of Total Amino Acids)
low. The effect of di fTcring nitrogen contents of wheat      (%)
and oats on lysine content is shown in Table 3-10.                                 Wheat                Oats
               ��·   AJbumlnaandGlob<II�
                                                              Adapted from Eppendorfer, 1978. J. Sci. Food Agric.
                                                              29:995-100 I .
Protein Content
Adapted from Hosenay, R.C. 1986. Principles of Cereal         (i.e. diets are formulated based on total rather than
Science and Technology. American Association of Cereal        available amino acid levels) then animal
Chemists, Inc.
                                                              performance will likely be reduced if soybean meal
Figure 3-3. Relationship of Protein Content to                is replaced by canola meal. However, research at
Protein Type in Cereal Grains.                                various institutions, including the Prairie Swine
                                                              Centre, has demonstrated that when diets are
                                                              formulated based on available amino acid levels in
Amino Acid Availabilities in Feed                             canola meal and soybean meal. animal performance
Ingredients                                                   can be maintained at high levels even if large
   In addition to the total amino acid content in feed        quantities of canola meal are included in the diet.
ingredients, the availability of these amino acids
should also be considered in practical feed                      Differences in availabilities between feedstuffs
formulation. Over the last decade, a considerable             can be attributed to various factors. These factors
amount of research has been conducted that                    include fibre levels, levels of anti-nutritional factors
demonstrates large differences in amino acid                  that interfere with nutrient digestion and utilization,
availabilities between various feedstuffs. For                protein quality, and heat damage. For example, raw
example, the availability of lysine in canola meal is         soybeans contain anti-nutritional factors called
lower than that in soybean meal. If these                     trypsin inhibitors, which reduce the trypsins' effec-
differences are overlooked in a feed formulation              tiveness (trypsins are digestive enzymes produced
Table 3-9. Amino Acid Profile of Various Protein Fractions in Corn (g/IOOg of protein).
                                                         26
by pigs). When soybeans are processed (oil extrac-             protein level in the diet is raised. Because separat-
tion or roasting), these trypsin inhibitors will be            ing endogenous secretions from those of dietary
inactivated.                                                   origin is difficult, estimating true amino acid
                                                               digestibilities in feedstuffs for swine is not yet
    Amino acid availabilities are routinely estimated          possible.
 from apparent ileal digestibilities, i.e., the disap-
pearance (absorption) of amino acids from the                      There is, however, a concern about apparent
digestive tract prior to the end of the small intestine        amino acid digestibilities in pig feed ingredients
(terminal ileum). Heal digestibilities are a better            related to these endogenous secretions. More
reflection of amino acid availabilities than                   specifically, the apparent amino acid digestibilities
digestibilities measured over the entire digestive             in a mixture of ingredients many not be the same as
tract because essential amino acids that enter the             that calculated from the inclusion level of the
large intestine, or cecum. may be altered by resident          various ingredients in the diet and their apparent
bacteria. The bacteria may break down some of                  digestible amino acid contents. This may cause
these amino acids, convert them to other amino                 errors in feed formulation. This concern can be
acids, or even manufacture new amino acids from                attributed to the effects of dietary amino acid levels
ammonia, thus contributing to an inaccurate digest-            ( or inclusion levels of the protein source in the test
ibility measurement. Essential amino acids that are            diet) on observed apparent amino acid digestibilities
digested by bacteria present in the pig's large                (Figure 3-4a). For example, the apparent amino
intestine and cecum are not available to the animal.           acid digestibilities in cereal grains, and com in
Therefore, to measure the net amounts of amino                 particular. are generally lower than in protein
acids extracted from the diet, digesta is collected            sources such as soybean meal. This is because
before it enters the large intestine. Measuring                amino acid digestibilities in cereal grains are deter-
amino acids excreted in the feces wil I not reflect            mined at lower levels of amino acids in the test diet.
unabsorbed amino acids, but rather unabsorbed                  This corresponds to low apparent amino acid
amino acids after possible alteration by the bacteria.         digestibilities (Figure 3 - 4a).
Many nutritionists now formulate diets using ilea!
digestible amino acids rather than total amino acids              Apparent amino acid degestibilities in protein
in a feedstuff.                                                sources, such as soybean meal and canola meal, are
                                                               determined at higher amino acid levels in the test
   Table 3-11 summarizes the estimated apparent                diet, corresponding to higher apparent amino acid
ileal digestibility of amino acids from common feed            digestibilities (Figure 3-4a). If corrections are made
ingredients. The word 'apparent' refers to the fact            for the minimum endogenous gut amino acid losses,
that the values are not adjusted for what are called           then the corrected "true" amino acid digestibilities
endogenous (internally produced) secretions.                   are generally independent of the dietary amino acid
                                                               level (Figure 3 - 4a). Minimum gut amino acid
   Amino acids consumed in the diet are not the                losses can be derived from feeding protein free
only amino acids entering the small intestine.                 diets or from regression to zero protein intake when
Endogenous secretions, which include sloughed off              various diets are fed with varying protein levels
intestinal cells, digestive enzymes, and possibly              (Figure 3-4b ). However, it should be stressed that
hormones, are added by the stomach, the salivary               the actual endogenous gut amino acid losses are
glands, and the small intestine. This process was              higher than the minimum losses and they arc
described in chapter 2. If these endogenous secre-             affected by various dietary and animal factors. This
tions were constant, apparent digestibilities could be         implies that these corrected "true" digestibilities arc
easily corrected to true digestibilities. However,             merely a means to improve the precision of diet
that is not the case, and many factors appear to               formulation; they are not an accurate reflection of
influence the quantity of endogenous secretions in             the actual true digestibility of amino acids in
the gut. An important observation is that endog-               feedstuffs.
enous gut protein losses appear to increase as the
                                                          27
Table 3-11. Average Apparent AvailabiUty of Amino Acids (%) in Common Ingredients.
Alfalfa Meal 55 47 62 47 54 52 22
Barley 74 68 79 64 68 71 72
Blood Meal 66 84 78 81 80 85 74
Canola Meal 72 74 78 68 64 78 77
Corn 78 65 86 68 64 78 76
Fish Meal                     85            86           87             80           76           83        65
   (high quality)
Meat & Bone                   7l            70           77             66           55           72        54
   Meal
Oat Groats 83 79 83 75 80 83 76
Peas 78 83 77 69 67 71 60
SBM (44%) 82 84 85 76 80 79 74
SBM (47%) 83 85 85 78 80 81 78
Sunflower                     78            74           87            72            77           75        73
Meal
Triticale 80 73 83 64 70 78 82
Wheat(HRS) 83 72 84 71 79 79 81
Whey 88 81 88 82 82 87 90
Derived from: Centraal Veevoeder Bureau ( 1994 Table of Feeding Values of Animal Feed Ingredients); Heartland
Lysine ( 1994 Swine and Poultry Digestibility Tables); BioKyowa ( 199 I Digestible Amino Acids and Digestible Amino
Acids for Swine, BioKyowa Technical Review 2); Rhone Poulenc (1990, first version, Nutrition Guide); Degussa
(Ileal Digestible Amino Acids in feedstuffs for Pigs).
                                                        28
      95         .                .                                      -i      0.4
                                            - --- ------·
                 ............. -1 ...... � ...............
�
�     90                                                                 c
                                                                          c,     0.3
�     85                          ,,.. .,     '-                         �
                                 /o                                      :§
:0                           /                     "True"                        0.2                              1 - "true" dig.
;
Ill   80                 /                                                �
                                                   Apparent
                 .,, I
Cl)                                                                       G)
c,                                                                        >      0.1
      75                                                                  0
0                                                                         CJ
                                                                          Cl)              Minimum endogenous methionine losses
      TO                                                                 a:::      0
           0          0.1                   0.2    0.3        0.4                      0         0.1       0.2        0.3           0.4
                     Dietary Methionine (% OM)                                                 Dietary Methionine (% in OM)
Figure 3-4. Relationship between Amino Acid Level in Diet and Observed Apparent and Calculated1 "True"
Ilea! Amino Acid Digestibilities (Figure 3-4a), and the Relationship between Amino Acid Level in Diet and
Observed Recovery at Distal Ileum (Total Endogenous and Non-digested Dietary Amino Acids)(Figure 3-4b).
'According to a review of the literature, it can be estimated that the minimum endogenous gut amino acid losses
(AAend; g/kg dry matter intake) are 0.38 - lysine, 0.10 - methionine, 0.3 l - methionine plus cystine, 0.51 - threonine,
0.18 - tryptophan, and 0.24 - isoleucine. The "true" digestibility (TD,%) can be calculated from observed apparent
digestibilities (AD,%) and the amino acid level in the test diet (AAd, g/kg dry matter) as follows: TD= 100 x (1-
[AAd-{AAd x AD/100} -AAend)/AAd). Note that the slope in the right hand graph represents the true digestibility.
Derived from Fan, M. R. 1994. Methodological considerations for the determination of amino acid digestibility in
pigs. PhD. Thesis. University of Alberta.
   If diets are formulated based on 'true' rather than                      One of the unfortunate features of reduced
apparent ileal digestible amino acid contents in feed                    availability of amino acids due to heating is that
ingredients, then adjustments may have to be made                        lysine is the amino acid most affected. Lysine can
to the requirements as well. Requirements ex-                            interact with sugars in a chemical reaction called
pressed as apparent digestibilities (g/kg of diet) can                   'carmelization'. This reaction is stimulated by heat
be converted to those expressed as 'true'                                in the presence of moisture, the exact conditions
digestibilities, simply by adding the minimum gut                        present in the drying process. Carmelization, as its
amino acid losses (g/kg of diet; g/kg of dry matter x                    name suggests, is associated with a colour change in
dry matter content of the diet/I 00).                                    the grain or protein. Thus, darkened grains or
                                                                         protein supplements tend to be down-graded for
   Unfortunately (apparent, or true, ileal)                              livestock due to concerns about carrnelization and
digestibilities do not always provide a good esti-                       associated changes in lysine availability. At present,
mate of amino acid availability. The amino acids in                      there is no assay available that will allow us to
some ingredients are digestible but unavailable                          routinely evaluate this chemical (un)availability.
because they are absorbed in a form which renders
them useless to the animal. These amino acids are                        Amino Acid Requirements
excreted, mostly in the urine. This occurrence is a                           Clearly, dietary protein is very critical to the
concern especially for heat treated ingredients of                       pig. None of the body processes described above
animal origin - milk products, blood meal, meat                          could occur without the necessary proteins being
meal, fish meal- but also applies to some ingredi-                       present. Consequently, inadequate amino acid
ents of plant origin.                                                    intake from the diet resulting in impaired protein
                                                                    29
synthesis has a profound effect on a pig's health and                                                                The explanation above brings up the concept
productivity. That amino acids present in the diet                                                             of the 'first limiting' amino acid. In Figure 3-5,
are balanced according to the pig's need is very                                                               lysine would be the first limiting amino acid in the
important. If all other amino acids are present in                                                             deficient diet because the amount present is lower,
quantities well above requirement, but lysine is                                                               relative to requirement, than any other amino acid.
missing or inadequate, the pig will still be unable to                                                         Methionine would be considered second limiting.
grow to its full potential because it must have all                                                            There are no third limiting amino acids in this
amino acids present in adequate quantities. Each                                                               example. Lysine is the amino acid discussed most
individual protein is created by a specific sequence                                                           often because in practical diets for swine, it is the
of amino acids; the lack of any single amino acid                                                              one most likely to be first limiting. Thus,
can prevent formation of the total protein.                                                                    nutritionists pay particular attention to lysine.
                                                                                                               Remember, however, that while adding extra lysine
      The importance of a proper balance of amino                                                              to a deficient diet will be beneficial, you must also
acids is illustrated in Figure 3-5. Tn the adequate                                                            consider the supply of other limiting amino acids if
diet, all amino acids arc present at 100% of                                                                   the diet is to be fully balanced.
requirement. Thus, when the body requires amino
acids for protein synthesis, each will be present in                                                             The balance in which amino acids are required by
sufficient quantity. In the inadequate diet, both                                                              pigs can be determined in a number of ways. Until
lysine and methionine are present at levels well                                                               recently, the most common approach was to study
below requirement. If lysine is present at 50% of                                                              individual amino acids and determine the require-
the requirement, it will place this upper limit on the                                                         ment of each one in terms of maximum growth rate
rate of protein synthesis. Not only will body                                                                  or carcass quality. A second approach, gaining in
functions be impaired, but other amino acids                                                                   popularity, is to relate the requirement for all amino
supplied in adequate quantity will be wasted.                                                                  acids to one reference amino acid, often lysine.
                                                                                                               This approach emphasizes the concept of amino
                                               Deficient Diet
                                                                                                               acid balance and integrates the requirement for all
                    r: r r r                             ,e::;          r: r: r:                      c.
       c
              100
                                                                                                       UI      10 essential and two semi-essential amino acids.
                                                                                                       z
       "E                                                                                                      For example, the requirement for lysine is defined
        e                                                                                              �
      ·;;
       O"                                      ,e::;      w
                                                          z
                                                                               z              ..
                                                                                             L:;       /::     in absolute terms (e.g., 1.3% in a starter diet). The
      ...."
                                                                                              z        o6
                                 w                        z      c                                     w
      0::                                       UI
                                                                               <               "'              requirement for the other amino acids is then
                                                                                               .."'
              50                 z                        <             w                              z
                    w                           !:                      z      x              >
                                                                                                       z
                                                                        z
       0                 UI                               .J                                  u
       c<>          z    z       0:,     w      z         <                    0.
                                                                                                       .sj     established relative to lysine .
       ..           z                    z      0                              0
                                                                                              I"
                                                          .J
                       0                        -x               UI     0      ....    w
        e           i, ;::
                                 UI
                                 .J      0:,    ...UI
                                                          )-
                                                          z
                                                          w      .,
                                                                 z      w
                                                                        a:
                                                                               a.      !:              >
                                                                                                       z
                         "'x
                                                                                )-     .J
      0..
                    a:
                    <
                                 0
                                 �       w
                                         .J     l
                                                          x
                                                          0.     ., ...x a:...
                                                                 >                     <
                                                                                       >
                                                                                                       UI
                                                                                                       :t
                                                                                                       a.            Table 3-12 provides an estimate for the optimum
                0
                                                                                                                   amino acid balance, also referred to as the ideal
                                 ESSENTIAL AMINO ACIDS
                                                                                                                   dietary protein. Lysine is set at l 00 and all other
                                                                                                                   amino acids are defined relative to I 00. For exam-
                                               Deficient 01e1                                                      ple, the amount of tryptophan in a diet should be
              100   r.r.        /""'.:   r               ,e::;          r r r                         ,e::;        about 19% of lysine, while threonine should be
      cc
                                                                                                      I
                                                                                                      UI
                                                                                                                   65% of lysine. Thus. in a diet containing 0.85%
      -�g-
      6
      c
                                                          w                                  ,e::;                 lysine. there should be at least 0.16% tryptophan
                                               ,e::;
                                                                                              "' ..,w
                                                          z
                                                                               z                                   and 0.55% threonine. Remember that the balance
                                 UI             UI        z L:;
                                                          .,                   <              !
                                                                                              ..
      �       60                                z         <             w
                                 !:                                                                    i:
       0            UI   UI
                         z                      z         .,<           !:     :t            >
                                                                                             o         z           in which amino acids arc required by pigs will vary
                                                                                                       3
                                                                               0.
      c             !:   0
                                 0
                                 :,
                                         w
                                         z      0                       z
                                                                               ...0.
                                                                               0       w     l!!
      e"            z            .,
                                 w                        )-     UI     0              z                           somewhat with body weight, (lean) growth rate,
                                                ...UIi                                       �
                                                                                                       I
                    e    ;::             0                z !:          UI
                                                                        a:             ::;
      a".                                :,               w
                                                                                                                   level of milk production, and diet composition. For
                         "':c                                    .,"'                         ...2
                                                                               )-
                    a:           0       w                              :c     a:             i=
                                                                               .... <>
                                                          :t
                                 !!
                                                                 )-
                    <                    .J     l         0.            ....
                0                                                                                                  example, as pigs grow heavier relatively more
                                 llSSENTIAL AMlNO ACIDS                                                            protein is required for body maintenance functions.
                                                                                                                   Because proteins required for body maintenance
Figure 3-5. Diagram Illustrating What is Meant by                                                                  functions contain large quantities of threonine and
the Term Amino Acid Balance as it Relates to Evalu-
                                                                                                                   T.S.A.A. relative to lysine, requirements for
ating Swine Diets.
                                                                                                              30
threonine and T.S.A.A., expressed as a proportion            However, minerals have other important functions
of lysine, will increase as pigs grow heavier. In a          to fulfill if the pig is to perform well. Many
similar fashion, requirements for threonine and              enzymes will not function unless certain minerals
T.S.A.A., relative to lysine, will reduce as lean            are present. Muscles could not contract and nerves
growth rates in pigs continue to increase. The               could not send signals in the absence of minerals.
concept of ideal protein, the optimum balance in
which amino acids are required by pigs, has proven               Minerals can be assayed directly in feedstuffs
very useful in practical swine diet formulation. This        with great accuracy and precision. Samples are first
concept a11ows for a rapid estimation of require-            ashed using very high temperatures (550°C) or
ments for essential and semi-essential amino acids           strong acids to remove all but the mineral content.
once the requirements for one amino acid have been           This residual material is then dissolved in weak acid
established. Determining the optimum amino acid              and assayed by various methods.
balance for different groups of pigs under varying
conditions remains an important area in swine                    Minerals can be divided into two types:
nutrition research. Refer to the specific chapters on        macrominerals and microminerals (Table 3-13).
feeding various classes of swine for further infor-          They are differentiated solely by the relative
mation on amino acid requirements and optimum                amounts present in the diet. Macrominerals are
dietary amino acid balances.                                 present in greater concentration and are therefore
                                                             measured in terms of percent or grams per kilogram
                                                             (1 g/kg equals 0.1%). Microminerals are present in
Table 3-12. Suggested Optimum Amino Acid                     much smaller quantities and are measured in terms
Balance in Protein {Ideal Protein) for Swine.                of parts per million or milligrams per kilogram ( 1
                                                             ppm equals I mg/kg). Thus, the concentration of
Amino acid                       Suggested balance           rnicrominerals in the diet is about one-tenth to one-
Lysine                                  100                  ten thousandth that of the macrominerals. To put
Threonine                               60-70                ppm and percent in perspective: 1 ppm is I second
Methionine                              30                   out of 11 .5 days; 1 percent is approximately one
Methionine plus cystine                                      second out of 1.5 minutes.
(T.S.A.A.)                              56-64
Tryptophan                              18-20                    The quantity of each mineral present in a pig's
Isoleucine                             60                    body reflects, in a general way, the amount required
Leu cine                               100                   in the diet. Table 3-13 shows the approximate
Phenylalanine                          60                    concentration of minerals in the animal body. Note
Phenylalanine+ tyrosine                 100                  that calcium and phosphorus are present in by far
Valine                                 70                    the highest concentration, while minerals such as
Histidine                              30                    selenium and chromium can be found only in trace
Arginine                               45                    amounts. Calcium and phosphorus are involved in
                                                             the skeleton and are thus present in large quantities,
                                                             while selenium supports the activity of certain
                                                             enzymes, a role which requires only trace amounts.
Minerals                                                     Importance should not be equated with quantity; a
   Minerals are an important constituent of the pig's        deficiency of a micromineral can be just as serious
diet. As a group, they perform far more functions            as that of a macrornineral.
in the body than they are normally given credit for.
We tend to associate minerals, such as calcium and
phosphorus, with the hard structures of the body
because of their involvement with bones and teeth.
                                                        31
Table 3-13. Approximate Mineral Composition of the           Although limestone is not expensive, supple-
Animal Body.                                             menting large quantities of calcium 'costs' a consid-
                                                         erable amount in terms of the total diet because
Macrominerals             Microrninerals                 limestone dilutes all other nutrients. Although
Element         %         Element          ppm           calcium is added to the diet at least expense by
                                                         using limestone, limestone contains no energy or
Calcium        l.50       Iron             20-80         protein, so adding it in place of barley or com or
Phosphorus     l.00       Zinc             10-50         any other basal grain will reduce the diet's energy
Potassium      0.20       Copper           1-5           and amino acid content unless other adjustments are
Sodium         0. 16      Manganese        0.20-0.50     made. To provide a rough estimate of the true cost
Chlorine       0.11       Iodine           0.3-0.6       of additional calcium in a diet, multiply the cost of
Sulphur        0.15       Molybdenum       1-4           limestone required to meet the new levels by
                          Selenium         1.7           approximately 2 or 3. This equation accounts for
                          Chromium         0.08          the cost of overcoming nutrient dilution by lime-
                                                         stone as well as the cost of adding the calcium
Adapted from: McDonald, et al. 1973.
                                                         itself.
                                                        32
the body to generate energy in one location for later          calcium costs about forty cents per tonne while
use in a different part of the cell. Although none of          0.1 % additional phosphorus costs two dollars. As
this activity requires large amounts, phosphorus is            with calcium, part of this cost is associated with the
nonetheless important to keep the body functioning             cost of the phosphorus itself and part is overcoming
properly.                                                      the diluting effect of the added phosphorus source.
                                                               Commonly used sources of supplemental phosphate
  A major concern with respect to dietary phospho-             are shown in Table 3-15.
nis levels is the proportion the pig actually utilizes.
A part of the phosphorus supplied by cereaJ grains
and vegetable proteins is bound to phytate and is              Table 3-15. Feed Grade Phosphate Sources.
much less available than in other forms. Table 3-14
shows the amount of total phosphorus present in                Source            Phosphorus     Calcium Sodium
common ingredients and an estimate of phosphorus                                                 (%)
availability.
                                                               Bone meal            12.0         24.0
                                                               Defluorinated
Table 3-14. Total Phosphorus Content and Phospho-              phosphate            18.0         31.0         4.0
rus Availability of Some Common Ingredients.                   Diammonium
                                                               phosphate            20.0         trace
Ingredient      Total             Phosphorus                   Di calcium
                Phosphorus (%)    Bio-availability(%)          phosphate            18.0         21.0
                                                               Monocalcium
Corn,dry        0.25              15
                                                               phosphate            22.0         16.0
Corn,
high moisture   0.25              49
Barley                                                         Note: Actual nutrient composition may vary depending
                0.29              31
                                                               on the supplier. Nutrient composition should be con-
Wheat           0.34              50
                                                               firmed by chemical analysis.
Oats            0.31              30
SBM, 47%        0.60              25
Canolameal      1.07              21                               Given the low availability of phosphorus in most
Peas            I.OJ              47                           swine feed ingredients, the cost of phosphorus, and
Wheat                                                          the contribution of phosphorus excreted with swine
middlings       0.95              45                           manure to environmental pollution in areas with
Wheat bran      1.10              35                           extensive swine operations, there is much interest in
Meat and
                                                               enhancing the efficiency with which dietary phos-
bone meal       4.75              68
                                                               phorus is utilized by pigs. A means to accomplish
Derived from NRC, 1988 and table of feeding values of          this is to add enzymes, called phytases, to the pig's
Animal Feed Ingredients (Centraal Veevoeder Bureau)            diet. Various studies have shown that these en-
The Netherlands.                                               zymes enhance the availability of phytate phospho-
                                                               rus in pig feed ingredients. This reduces the need
   Whereas phytate is an undesired component of                for inorganic phosphorus in pig diets. Concerns
swine diets, it plays a very important role for                with the use of these phytases include cost and
growing plants that need it to store phosphorus for            product stability. Phytases are generally unstable at
later use. Late application offertilizer may increase          higher temperatures which is a concern when feeds
total phosphorus in the seed but much of the addi-             are pelleted. It should be noted that some ingredi-
tional mineral is in the form of phytate and less              ents such as wheat and triticale contain some
available to the pig when it eats the plant.                   endogenous phytases.
  Phosphorus is a much more expensive ingredient                  The ratio of calcium to phosphorus, and more
than calcium, so maximizing utilization should be a            accurately, available phosphorus, in the diet should
priority. Presently, the addition of 0.1 % additional          also be considered. Dietary calcium to phosphorus
                                                          33
levels of 3: I and 1.25: I for available and total            0.05% sodium (compared with a requirement of
phosphorus, respectively, are thought to optimize             0.10%). Vegetable-based protein ingredients are
calcium and phosphorus utilization. The impor-                much richer in sodium, containing 0.30 - 0.50%. A
tance of the ratio appears to be greater if calcium is        good, economical source of sodium is common salt
present at or slightly above requirement. In practi-          (39% sodium). Feed companies sell four types of
cal diets the ratio of calcium to available phospho-          salt defined according to colour. White salt con-
rus in the diet should be maintained at 2.7 to 3.4.           tains only NaCl, red salt contains iodine, blue salt
This corresponds with calcium to total phosphorus             contains iodine and cobalt, and brown salt contains
rations of approximately 1.1 to 1.5. If the diet              trace minerals. For most swine diets, salt will be
contains plenty of calcium and phosphorus, the ratio          added as part of the premix or supplement. If the
becomes much less important. A deficiency in                  premix contains other sources of iodine or trace
vitamin D also impairs the utilization of calcium in          minerals, only pure (i.e., white) salt should be
the diet.                                                     added to the diet.
                                                         34
from feed has reduced the scouring associated with        hydrochloride, a source of the amino acid lysine.
poor-quality water, a chloride deficiency (which          Both chloride sources could bias the salt value if
results in reduced feed intake, thus reduced scour-       chloride is used to estimate salt content. Thus, if a
ing), is likely to have developed. A pig's perform-       diet is found to contain excessive 'salt', be sure to
ance will suffer more as a consequence of the             consider the method of analysis before worrying
chloride deficiency than it would have due to the         about a possible formulation or mixing error.
water quality problem.                                    Consult a nutritionist if you suspect a problem.
                                                         35
action may provide some relief, a chloride defi-            (38.0% potassium) and potassium chloride (50.5%
ciency may result since water rarely contains much          potassium) are logical choices.
chloride. Therefore, even partial removal of dietary
salt requires some care; consult a qualified nutri-         Sulphur
tionist since a deficiency in chloride will depress           Most of the sulphur present in a pig's body is
feed intake and slow growth rate.                           found in the amino acids, methionine and cystine.
                                                            Practical diets are never deficient in sulphur per se
Magnesium                                                   and supplementing a diet with pure sulphur is not
  About two-thirds of the total magnesium in the            required. However, the sulphur amino acids need to
body is found in bone. Magnesium is important in            be added via the protein supplement to meet the
stimulating the activity of many enzymes that               pig's requirements. (see Amino Acids)
catalyse chemical reactions in the body. It is also
required for normal nerve and muscle function, and          Micro minerals
for fat and protein utilization.                              The microminerals required in a pig's diet are
                                                            discussed in the following sections. Some
  Like potassium, common feed ingredients supply            microminerals are present in sufficient quantity in
sufficient magnesium to meet the pig's normal               basal ingredients and are not normally
requirements. For example, a cereal grain-based             supplemented. Others are routinely supplemented
diet will contain about 0.1 % magnesium, more than          to prevent deficiency symptoms. Table 3-17
twice the pig's requirement. If magnesium must be           summarizes the information on the microminerals
supplemented, magnesium oxide (51-59%                       including those which are normally supplemented in
magnesium) can be used.                                     practical diets, maximum tolerable limits for the
                                                            pig, and common supplemental sources.
Potassium
   Potassium is an important mineral. It helps to           Cobalt
maintain both water and acid-base balance, supports           Although there is some evidence that cobalt may
the transport of nutrients across cell membranes,           be involved in several bodily functions, cobalt's
and is involved in maintaining heart rate.                  major role is as a constituent of vitamin B12• A pig's
                                                            requirement for cobalt is very low; if vitamin 812 is
   Potassium tends to be overlooked in nutrition            properly supplemented, there is no need to add
because natural ingredients supply more than                additional cobalt to swine diets.
enough of it to meet a pig's requirement. Common
protein supplements, such as soybean or canola              Copper
meals, are particularly good sources of potassium.            Copper is essential for formation of red blood
For this reason, combined with the fact that cereal         cells. A copper deficiency may lead to anemia, a
grains often supply all the potassium the pig               type very different from that caused by an iron or
requires, a potassium deficiency is not usually a           vitamin B12 deficiency. Copper plays an important
concern in practical diets.                                 role in certain enzyme systems, most importantly
                                                            those involved in energy metabolism. Finally,
   Excess potassium may depress the absorption of           copper is required for normal hair pigmentation.
many nutrients from the intestinal tract but this
situation appears to require dietary levels in the            Although a copper deficiency is very rare in
range of2.0% (practical diets contain less than 1%          swine, copper is always supplemented in a pig's
potassium) combined with a wide                             diet. Normally, only 6-12 ppm is required. Usually
potassium:sodium ratio.                                     this amount is supplied by copper oxide (75%
                                                            copper) or copper sulphate (25% copper), the latter
  Potassium rarely needs to be supplemented to              being the more available source of copper. Metallic
healthy swine, but if it docs, potassium bicarbonate        copper is poorly absorbed so is not used in diets.
                                                       36
Table 3-17. Maximum Tolerable Limits and Supplemental Sources of the Microminerals.
Cobalt No 10
analysis.
"Legal upper limit according to the Canadian feeds Act (see chapter I I).
  Copper sulphate, also called 'bluestone' because             Adding copper sulphate to a diet at growth
of its blue colour, was a popular disinfectant in the        promoting levels will produce black sticky feces in
days before more powerful and effective agents               pigs. Also, in regions of high livestock density, the
were discovered. For example, copper sulphate was            use of copper as a growth promotant has sparked
used in foot baths to treat lame animals.                    criticism due to the potential accumulation of
                                                             copper in the soil. Although swine can tolerate
  Copper is now added to swine diets in high levels          copper levels in excess of 250 ppm, sheep have
to act as an inexpensive growth promotant. It is a           been found to be highly susceptible to copper
more effective growth promotant in diets for starter         toxicity. Total dietary copper levels for sheep
and grower pigs than for finishing pigs. Adding up           should not exceed 25 ppm. Consequently, swine
to 125 ppm copper (from copper sulphate) to feed             diets containing copper at growth promotant levels
is legally permitted in Canada. This amount is well          must not be fed to sheep.
above the amount required for nutritional purposes
but costs less than one dollar per tonne.
                                                           37
Chromium                                                     known goitrogenic compounds in rapeseed meal.
   Until recently, chromium was not considered an            Canola meal, with very low levels of glucosinolates,
essential nutrient for swine. Recently, however,             can be used in swine diets (including those of
considerable interest has been generated in the              breeding animals) without fear of impairing thyroid
effect of chromium and chromium-containing feed              function.
additives on the performance of growing-finishing
pigs and sows. Chromium, via its involvement with              Primary iodine deficiency is prevented by includ-
the so-called glucose tolerance factor. may be               ing iodine in the mineral premix. Various forms of
required for the utilization of large amounts of             iodine can be used including potassium iodide
sugars. There are also some indications that chro-           (68.5% iodine), calcium iodate (63.5% iodine), or
mium may improve the animal's immune status and              EDDI (ethylene diamine dihydro-iodide; 79.5%
increase the effectiveness of insulin. Insulin,              iodine). Adding iodized salt (0.007% iodine) at the
produced by the pig, is an important hormone that            rate of 2 kg per tonne of feed provides sufficient
is required for energy and amino acid utilization.           iodine to meet the pig's requirement. Feeds of
                                                             marine origin, such as fish meals, are also rich
   Several studies have demonstrated that including          sources of iodine.
inorganic chromium in practical swine diets does
not affect animal performance. Other studies have            Iron
shown that adding certain chromium-containing                  Iron, a component of hemoglobin in the blood, is
compounds, such as chromium picolinate, to                   required to help the blood transport oxygen from
finishing pig diets improves animal performance,             the lungs to the body tissues. An iron deficiency
especially the lean yield in the carcass and                 can be very serious because the oxygen-carrying
reproductive performance in sows. Further research           capacity of the blood is impaired. lron is also a
is required to determine the optimum level of                constituent of many important enzymes throughout
chromium and the form in which it should be                  the body. Symptoms of deficiency include poor
included in pig diets and the conditions under which         vigour, lethargy. and a pale, white skin colour.
it is most effective.
                                                               Iron is always supplemented in pig diets, although
Iodine                                                       pigs would probably do quite well without it. A
  Iodine's main role in the body is as part of the           pig's requirement for iron is not very high and its
hormone, thyroxine. Thyroxine, produced by the               body has the ability to retain iron very well, espe-
thyroid gland, is involved in regulating a pig's             cially if dietary sources are scarce. Furthermore, as
overall metabolic rate. An iodine deficiency,                the need for iron rises, so does the ability of the
resulting in impaired thyroid function, will have a          pig's gut to extract iron from the diet. If market
serious effect on swine growth and productivity.             pigs or sows appear iron deficient, the cause is more
                                                             likely a bleeding ulcer, or other illness, than a
  Iodine deficiency leads to a condition known as            dietary deficiency. However, iron is very inexpen-
goiter, which refers to an enlargement of the thyroid        sive and is added to the diet as cheap insurance.
gland. Goiter can be observed as a swelling in the
neck where the thyroid gland is located. Goiter                A definite exception to the pig's limited need for
causes reproductive failure or the birth of weak or          iron is the neonatal pig. NeonataJ pigs grow very
dead, hairless piglets.                                      rapidly and require a great deal of iron to support
                                                             this growth. Sow's milk is a poor source of iron so
  Goiter is not caused only by a dietary deficiency          supplements are imperative. Injectable iron in a
of iodine. Certain ingredients in the diet impair            single dose of approximately 150 mg should be
iodine utilization by the thyroid gland. The highly          administered no later than 3 days of age. A second
successful rapeseed breeding program, which                  injection may be given at approximately 21 days
resulted in the development of canola, was in part           (see Chapter 7).
motivated by a need to remove glucosinolates, well-
                                                        38
   [fan iron injection causes sudden mortality, the       Selenium
little pigs are possibly suffering from a vitamin E         The history of selenium as a nutrient is an inter-
and/or selenium deficiency. If this is the case, the      esting study. Initially selenium was considered only
solution is to treat the sow herd with either inject-     as a toxic substance, responsible for alkali disease
able vitamin E/selenium or increase the quantities of     and blind staggers in cattle. More recently nutri-
each element present in the diet.                         tionists have learned that animals must have sele-
                                                          nium in their diet. Selenium helps the body protect
  An iron deficiency leads to anemia, which in turn       against the breakdown of cell membranes. It works
causes stunted growth and reduced disease resist-         with vitamin E in this regard, so the two elements
ance. Some producers have seen almost 'magical'           are often discussed together.
recovery from scours in pigs 2 to 4 weeks of age
merely by providing iron that was deficient earlier         Because selenium was first identified as a toxic
in the pig's life. Avoid extreme excesses, however        substance and more recently has been found to
- some research suggests that too much iron may           cause cancer, government agencies are very con-
actually predispose young pigs to diarrhea.               servative in allowing selenium supplementation of
                                                          livestock feeds. ln Canada, 0.3 ppm can be added
   Iron should be injected in the neck rather than the    to diets for all classes of swine. Supplemented with
ham muscle. The neck location avoids the possibil-        adequate vitamin E, this amount would appear to be
ity of staining expensive cuts (e.g., the ham) and        adequate under most circumstances. Nonetheless,
making the product undesirable to the consumer.           reports of selenium deficiency persist, especially in
                                                          areas where the soil is low in selenium, thus a low
   Ferrous sulphate (20 - 32% iron) is the first          natural supply in the grain. Adding more than 0.3
choice of iron supplement in a diet. A less desir-        ppm of selenium to a diet requires a veterinary
able second choice is ferrous carbonate (36 - 45%         prescription.
iron); iron oxide is nothing more than rust and is of
little nutritional value. Iron oxide may appear on          Selenium deficiency in pigs result in mulberry
some feed labels because it is used as a colouring        heart disease and liver necrosis. But, before adding
agent to ensure a uniform red colour. This practice       selenium to a diet in response to a problem with
is most common in mineral supplements or                  mulberry heart disease, review the situation
premixes. Pigs eating such diets produce red              carefully. If the problem is occurring in recently
manure due to iron oxide's poor digestibility.            weaned pigs, supplementation of the starter diet
                                                          may be of limited value because the piglets will eat
Manganese                                                 very little before succumbing to the disease. In this
  Manganese acts as an important enzyme activator         case. increasing the vitamin E content of the nursing
so is critical for normal utilization of carbohydrate,    sow diet or injecting pregnant sows with injectable
fat, and protein in a pig's diet. Lameness is a           vitamin E-selenium is the preferred course of
symptom of deficiency because manganese is                action. Some additional attention to the diagnosis
involved in bone formation. Manganese deficiency          of mulberry heart disease is also advised; infections
can also impair normal reproductive performance.          of Strep. Suis Type II have been suggested as a
causing such problems as irregular estrus cycles,         plausible alternative diagnosis. Discuss your
late sexual maturity, and weak piglets at birth.          situation with a qualified nutritionist and/or veteri-
                                                          nary pathologist before beginning any treatment.
  Supplements are required because basal dietary
ingredients cannot be relied on to supply sufficient            Concerns about selenium toxicity must not be
manganese. Manganese is routinely added to the                underestimated because the difference between
mineral premix, usually as manganous oxide (55 -              required and toxic levels is so small. While the
65% manganese) or manganese sulphate (27 - 28%                requirement is in the range of 0.1 to 0.3 ppm, only
manganese), so a manganese deficiency in swine is             7.0 ppm (25 times the requirement), is toxic.
very rare.
                                                         39
  Selenium supplements in the diet are provided by         controlling£. coli scours. However, these levels
sodium selenite (45% selenium) or sodium selenate          are higher than is allowed in swine feeds in Canada.
(40% selenium). Because selenium is required in            In addition, if these high zinc levels arc fed for too
such small amounts, 0.7 gm of sodium selenite per          long. e.g., more than two to three weeks, they may
tonne will supply 0.3 ppm. Many feed companies             result in zinc toxicosis. Alternative means to
use diluted selenium sources to ensure proper              control the negative effects of£. coli scours are
distribution in the mix and to avoid the risk of           preferred.
toxicity. In some instances, selenium can be admin-
istered by injection, often as a combination of            Vitamins
vitamin E and selenium, to deal quickly with                 Vitamins are one of the six major nutrient classes
suspected problems.                                        in a diet. The other classes are energy, protein,
                                                           minerals, essential fatty acids, and water. A pig
Zinc                                                       requires vitamins to support or stimulate the many
  Zinc is an important component in a pig's diet.          chemical reactions that occur in the body as part of
Zinc is a constituent of many enzymes and acts as a        normal metabolism. Although they are present in
co-factor of others. It is also a part of the hormone,     small quantities, vitamins perform very important
insulin.                                                   functions. As is the case with all nutrients, the
                                                           quantity present is not an indicator of importance.
  Zinc deficiency in the pig results in poor overall
growth, reduced appetite, and poor hair growth. A            Vitamins can be analysed but the cost is high. Also,
specific syndrome in swine, known as parakeratosis,        once they are added into a mixed feed, the levels may
is the result of a zinc deficiency. It shows up as skin    be too low to be accurately analysed. It is rare to test
lesions which may open and later develop scabs.            for vitamins unless a problem arises. Many feed
Parakeratosis is associated with poor growth rate          manufacturers test for one or two vitamins, e.g., ribo-
and impaired feed efficiency.                              flavin and vitamin A, in spot samples to ensure proper
                                                           formulation and mixing of premixes.
  The utilization of zinc can be impaired by exces-
sive calcium in the diet. Indeed, dietary calcium has        Deficiencies of a vitamin or group of vitamins
a profound effect on zinc requirements. At recom-          must be avoided to maximize swine performance.
mended calcium intakes, 50 ppm zinc appears to be          Unfortunately, the role of vitamins in nutrition has
quite adequate; excessive calcium can more than            often been misunderstood; vitamins have in many
double the amount of zinc required to prevent              cases been viewed in almost the same way as drugs.
deficiency symptoms. Phytic acid. common in                For example, vitamin X will prevent spraddle leg or
cereal grains and protein supplements of plant             vitamin Y will improve estrous behaviour. Al-
origin, also reduces the pig's utilization of zinc.        though individual vitamins have specific functions
                                                           and a deficiency will result in certain symptoms,
  Zinc should always be supplemented in a swine            this does not mean that adding additional quantities
diet. Consequently, it is included in the mineral          of a vitamin or vitamins will improve health or
premix as zinc oxide (70-80% zinc) or zinc sulphate        performance. As always, the essence of nutrition
(36% zinc). As zinc supplied in the form of a              must be to balance all nutrients according to need
sulphate is more available to the pig than zinc            and not to assume that one nutrient possesses
supplied in the form of an oxide. zinc sulphate is         magical qualities.
preferred over zinc oxide.
                                                                 At least some vitamins arc naturally present in the
   Recent studies have shown that including ex-                diet and need not be supplemented. Others are
tremely high levels of zinc oxide (up to 3000 ppm              present, but are either poorly available (niacin) or
of zinc) in d icts for newly weaned piglets increased          arc in amounts well below the requirement (vitamin
feed intake and growth rate. It is speculated that at          E) so that supplementation is necessary. Some
these pharmacological levels, zinc is effective in             vitamms, such as vitamin C (ascorbic acid), are
                                                          40
synthesized by the pig. Others, such as biotin, are          of the situations that increase vitamin breakdown
produced by the microbes present in the lower gut.           and avoid them as much as possible. In general, the
Under some conditions, natural synthesis may be              fat soluble vitamins are less stable than the water
impaired, resulting in greater need for supplementa-         soluble vitamins. There are exceptions; for exam-
tion. For example, sulpha drugs in the feed or water         ple, vitamin C is very unstable in the diet.
may reduce or even eliminate biotin synthesis in the
gut. All vitamins can be produced chemically so                 Vitamin supplement manufacturers have recog-
that supplementation, when necessary, is not a               nized the problem of stability. Vitamins A, D, and
problem other than cost. Overall, vitamin fortifica-         E are supplied encased in gelatin that protect them
tion generally represents less than 4% of the total          from breakdown, but which still allow for absorp-
feed cost.                                                   tion in the gut. These gelatin beads also contain
                                                             anti-oxidants which provide further protection for
  Identifying vitamin deficiencies is not a simple           the vitamins inside.
task because the symptoms are rarely specific. For
example, a niacin deficiency impairs appetite and              The presence of trace minerals, such as copper
growth rate, and causes severe diarrhea. Many                and iron, in the diet tend to increase the breakdown
other factors can produce the same symptoms.                 of some vitamins. Poor quality fats may also cause
Consequently, symptoms must be considered in the             a problem. The concern regarding trace minerals is
context of other information, such as feed analysis,         greater in premixes than complete feeds since in a
herd history, and a thorough review of health status,        premix, trace minerals are present in much higher
to ensure an accurate vitamin deficiency diagnosis.          relative concentrations and are thus in more intimate
                                                             contact with vitamins.
  Vitamins are categorized according to solubility
characteristics. Vitamins A, D, E, and Kare called             Choline chloride is a particular stress factor for
the fat soluble vitamins. The B vitamins and                 vitamins. Breakdown can be as much as three to
vitamin Care called the water soluble vitamins.              four times higher in premixes containing choline
The fat soluble vitamins are stored in the pig's body        chloride than those without it.
so that brief periods of dietary deficiency rarely
cause a problem because the pig can draw on                    Table 3-18 summarizes information on the stabil-
reserves. The water soluble vitamins, because they           ity of individual vitamins. To ensure maximum
are not stored in the body, must be present in the           vitamin stability, premixes should be stored in a
diet on a more or less regular basis.                        cool, dry, dark location and inventories should be
                                                             turned over reasonably quickly, particularly in the
 In Canada, all diets based on cereal grains must be         summer when the temperature and humidity rise.
supplemented with vitamins A, D, E, K, and B12, as
well as riboflavin, niacin, and pantothenic acid.              A premix containing only vitamins can be stored
Sow diets should contain added choline, and prob-            up to 6 months with minimal deterioration, except
ably folic acid and biotin. Other vitamins, if               for folic acid, which deteriorates more rapidly.
supplemented, are probably being included only as            Vitamin-trace mineral premixes, sometimes called
a safety factor.                                             micro-premixes, which do not contain calcium,
                                                             phosphorus, or salt, can be stored up to 4 months.
Vitamin Stability                                            However, losses of some vitamins under good
   Vitamin supplements deteriorate with age; certain         storage conditions will exceed 10%. Folic acid and
environmental factors accelerate the rate of deterio-        vitamin C, if present, are the least stable vitamins in
ration. Agents that tend to stimulate vitamin degra-         such premixes.
dation include moisture, oxygen (air), light, and
heat. pH is another concern but the producer can do            Macro-premixes, which contain supplements of
little to regulate it, for pH regulation is the feed         all vitamins and minerals (micro and
supplier's responsibility. A producer must be aware          macrominerals) should not be stored for more than
                                                        41
four months, since losses in excess of I 0% can          often added to provide further protection. Perhaps
occur for many vitamins including folic acid,            the greatest danger exists in vitamin-mineral
pyridoxine, vitamin 03,vitamin A, thiamine,              premixes that contain both vitamin A and trace
menadione (vitamin K), and if present, vitamin C.        minerals. For this reason, it is wise not to keep such
                                                         premixes longer than 4 months. Because modem
  The maximum time for storing complete feeds is         vitamin A sources are stable and because most
probably 4 months; losses of I 0% or more will           premixes contain far more vitamin A than a pig
occur after that period. Vitamins C and K, and           actually requires, a 4 month storage period should
thiamine will be particularly affected.                  not be excessive provided the premix is kept dry
                                                         and is not heated or exposed to light.
  Acceptable storage periods can be increased if
extra vitamins are added to the premix. By provid-             Vitamin A concentration is expressed in Interna-
ing large overages, shelf-life is extended because           tional Units (IU), an arbitrary scale that permits
losses due to storage are offset by higher original          comparison among different sources. One IU of
vitamin concentrations. Actual overages can be               vitamin A is equivalent to 0.3 ug of retinol (vitamin
calculated based on expected losses due to process-          A alcohol), 0.344 ug of retinyl acetate (vitamin A
ing and length of the storage time (Table 3 - 18).           acetate) or 0.55 ug retinyl palmitate (vitamin A
                                                             palmitate).
Vitamin A
   Vitamin A, known chemically as retinol, plays a             Fish oils are good natural sources of vitamin A.
major role in vision, bone development, reproduc-            Alfalfa meal contains carotenoids which are chemi-
tion, and in the formation of mucous membranes               cals that the animal can convert to vitamin A. As a
(i.e., lining of the lungs and intestine). Vitamin A         rule, one can assume that for swine, 1 mg of Beta-
deficiency can result in night blindness or un-co-           carotene is equivalent to 260 TU of vitamin A.
ordination. In sows, a deficiency may result in the          There are some indications that Beta-carotene itself
birth of dead. weak, or malformed piglets. Signs of          plays a role in swine reproduction, independent of
vitamin A deficiency include fetuses with small or           vitamin A. However, more research is required in
no eyes, a cleft palate, or hydrocephalus. These             this area. For example, some studies indicate that
symptoms, however, can be caused by other factors,           Beta-carotene appears effective only when adminis-
including some of genetic origin, so a vitamin A             tered via injections and not when included in the
deficiency cannot be diagnosed merely on the basis           feed. Other studies suggest that vitamin A, when
of malformed piglets. Vitamin A deficiency in the            administered in an injectable form, may enhance
sow will not cause abortion. In any event, vitamin           reproductive performance in sows just like inject-
A deficiency is rare now that diets are properly             able Beta-carotene.
supplemented.
                                                               Beta-carotene is widely distributed in certain
  Vitamin A is readily destroyed when exposed to             foods. Most green materials, such as alfalfa, are
heat, moisture. and light. The process can be                relatively rich in Beta-carotene since it is generally
accelerated if the vitamin is in close contact with          associated with chlorophyll (the compound which
trace minerals and/or unstable fats. Vitamin A               imparts the green colours in plants). There are
activity can best be maintained by storing the feed          exceptions, such as carrots and tomatoes, which
in a dry, cool, dark area.                                   contain carotene but not chlorophyll. A compound
                                                             called xanthophyll, although part of the carotenoid
  Vitamin A in feed is supplied as esters (com-              family, has no vitamin A value. It is often used in
pounds), such as vitamin A palmitate or vitamin A            poultry rations based on cereal grains to provide the
acetate, which are more stable than other forms.             yellow pigment in egg yolks, but xanthophyll has
They are also manufactured in tiny gelatin capsules          no value in swine diets.
that reduce deterioration rate and ensure both
stability and uniform distribution. Anti-oxidants are
                                                        42
  Hays may lose much of their vitamin A activity if             Vitamin D is formed naturally by exposure of the
left in the sun too long to cure, since heat and              pig's skin to sunlight. The ultraviolet rays in
sunlight destroy the carotenes. Cereal grains                 sunlight activate an enzyme responsible for convert-
contain essentially no vitamin A.                             ing a natural compound (7-dchydrocalciferol) into
                                                              vitamin D. For this reason, pigs housed indoors
Vitamin D                                                     require preformed vitamin D supplements in their
   A pig requires vitamin D for the proper utilization        diet.
of calcium and phosphorus, including stimulating
the gut to absorb them. Vitamin D also plays a very             Stability concerns, similar to those described for
important role in normal bone metabolism. A                   vitamin A, also exist for vitamin D, although it is a
vitamin D deficiency is most likely to appear as              somewhat more stable compound. Modern
lameness or other signs of disturbed calcium and              processing methods, generous diet formulation
phosphorus utilization. Pigs that are deficient may           standards, and proper storage practices combine to
have swollen joints, broken bones, or stiffness.              overcome potential concerns. ln terms of natural
However, various infectious agents can also cause             sources, most fish oils are excellent. Cereal grains
stiffness and swelling so determination of a vitamin          contain no vitamin D.
D deficiency cannot be made without detailed
diagnostic tests.
Table 3-18. Stability of Vitamins in Premixes and During Pelleting and Extrusion.
                                                      Stability
                       Very High          High       Moderate                     Low         Ver:i::Low
Vitamin                 Choline        Riboflavin Thiamine Mono                 Thiamine      Menadione
                        Chloride         Niacin     Folic Acid                     HCI         Ascorbic
                          B12          Pantothenic  Pyrodoxine                                    acid
                                          acid, E        DJ
                                         Biotin          A
                                                         (losses/month)
Premixes without
choline and trace          0             <0.5%                  0.5%               1%             2%
minerals
Premixes with
choline and             <0.5%              1%                    8%                15%            30%
trace minerals
                                                         43
  There are two chemical forms of vitamin D                 Vitamin E deficiency symptoms include muscle
referred to as vitamin 02, also called ergocalciferol,    weakness and liver damage. Reproductive impair-
and vitamin 03, known as cholecalciferol. Vitamin         ment is often used as a sign of vitamin E deficiency,
03 is the only form available in commercial supple-       but because the sow can store vitamin E for a long
ments since poultry have difficulty utilizing Vitamin     time and in considerable quantities, effects on litter
02 and because the cost of manufacturing vitamin          size are extremely rare.
D2 is greater than that for vitamin Dr Vitamin D
activity is expressed in International Units, where l           Researchers now know that vitamin E acts with
TU is equivalent to 0.025 ug of pure vitamin 03•              selenium to protect against some diseases. Thus,
                                                              nutritionists take great care to ensure that both
  Excess vitamin D must be avoided because it can             vitamin E and selenium are properly supplemented
lead to many health problems, such as accumulation            in a diet. If one is deficient, the requirement of the
of calcium in the kidney and blood vessels which,             other is raised. Conversely, a high dietary level of
in severe cases, will cause death. In some parts of           one reduces, but does not eliminate, the need for the
the world, high potency vitamin D preparations are            other. A vitamin E deficiency can be prevented by
used as rat poison.                                           supplying a properly balanced diet containing both
                                                              vitamin E and selenium supplements.
                                                         44
able to interchange these measurements because              Vitamin B12
different sources may be expressed in different               Vitamin B12, also known as cyanocobalamine, is
ways. One IU of vitamin Eis equivalent to I USP             required in extremely small quantities in the diet. It
unit which in turn equals l mg of dl-a-tocopheryl           used to be called the 'animal protein factor' because
acetate, 0.74 mg of d-a-tocopheryl acetate, 0.91 mg         before Vitamin B12 was discovered, producers knew
of dl-a-tocopherol, and 0.67 mg of d-a-tocopherol.          that animals receiving diets containing ingredients
The acetate form is preferred because it is more            solely of plant origin would develop pernicious
stable.                                                     anaemia. Now we know that plants are incapable of
                                                            producing vitamin B12 so diets based on grains are
  Vitamin Eis present in many feedstuffs but can be         completely devoid of it. Today, synthetic vitamin
destroyed by curing, artificial drying ( of grains),        B12 is readily available so pigs can grow quite well
and storage. The tocopherol content of many feed            on diets devoid of animal products. The vitamin is
ingredients are listed in Table 3-19. However, in           synthesized by bacteria so that pigs with access to
practical swine diets, sufficient supplemental              feces can obtain vitamin B12 from this source.
vitamin E is added to meet requirements so the              Vitamin B12 cannot be absorbed from the gut
natural sources provide an additional safety margin.        without the aid of the 'intrinsic factor', which is
                                                            secreted by the stomach of healthy animals. Ab-
Table 3-19. Alpha-tocopherol Content (mg/kg) of             sence of intrinsic factor, even with adequate vitamin
Feed lngredients.                                           B12, will result in pernicious anaemia.
                                                       45
synthesis, nitrogen excretion, and the maintenance               Choline deficiency has been imp! icated as a cause
of hair, skin, nerves, and sex glands. A deficiency            of spraddle leg in swine although there is very little
results in impaired growth rate, dermatitis, loss of           scientific support. Many experiments have evalu-
appetite, and spasticity of the hind legs.                     ated the effect of choline on spraddle leg and none
                                                               have concluded that choline was beneficial. The
  Biotin is found in cereal grains and other                   use of choline to prevent spraddle leg is question-
ingredients commonly fed to swine. The                         able and should be carefully considered from a
availability of natural biotin in most feedstuffs tends        cost:benefit perspective. Some producers spend as
to be poor and variable. The biotin in com is more             much as five or six dollars per tonne of sow diet to
available than that in wheat, but there is much less           provide generous supplementation of choline.
of it. Thus, the earlier belief that supplemental              Using good flooring materials that improve footing
biotin is not required in swine diets has been                 in the farrowing crates has proven to be a more
reconsidered and common practice now includes                  effective solution to the problem at less cost. Other
biotin in diets for the breeding herd and growing              approaches, such as tracing genetic predisposition in
boars and gilts that may be selected for breeding.             the breeding herd and removing parents of piglets
                                                               that seem predisposed to spraddle leg, is also
  Avidin, a naturally occurring compound in egg                effective in some herds.
whites that binds with biotin and makes it
nutritionally unavailable, can be added to swine                 The pig can synthesize choline from the amino
diets to induce deficiency symptoms. On the basis              acid, methionine. Thus, the dietary requirement for
of this research, we now understand that sows                  supplemental choline will depend on the pig's
deficient in biotin may develop hooflesions that               methionine status. Choline is supplied by the salt,
can lead to premature lameness. Various studies                choline chloride. It is very hygroscopic (attracts
show that adding biotin to sow diets increases                 water) so must be carefully stored. Pure choline
reproductive efficiency. Wheat or barley based sow             chloride contains 86.78% choline. Commercial
diets are generally supplemented with 150 to 250               supplements. available in dry forrn, contain either
mcg of biotin per kg of diet. Much higher levels are           50% or 60% choline chloride. Liquid choline
used in some cases, but the benefits are far from              chloride (70%) is also available.
clear. Fifty to one hundred (50-100) mcg of biotin
may be added to barley or wheat based diets for                Folic Acid
growing-finishing pigs. Since biotin is an                       Folic acid is the name given to the vitamin. while
expensive vitamin, adding an excessive safety                  the term, folacin, refers to compounds derived from
margin does not make economic sense. When                      the vitamin. Folic acid is involved in a number of
required, biotin is available in synthetic form.               functions, including cell division and growth, and
                                                               amino acid and nitrogen metabolism. Folic acid
Choline                                                        deficiency results in anemia (different from that
  Choline, unlike other vitamins. is required in the           caused by iron deficiency), reduced weight gains,
diet in a high concentration and acts more as a                and fading hair colour.
structural component of the body rather than in
support of chemical reactions. The body uses                     Until recently, it was believed that normal dietary
choline to produce chemical messengers called                  ingredients plus gut synthesis provided sufficient
neurotransmitters which allow nerves to send                   folic acid for pigs. Therefore, most swine diet
messages throughout the body. Choline also forms               premixes did not contain supplemental folic acid.
part of cell membranes, provides a surface active              However, recent information suggests that folic acid
agent required by the lungs, is involved in fat and            improves reproductive performance in sows.
cholesterol rnctabol ism, and acts as a precursor for
the formation of the amino acid, glycine. Signs of               Prolonged administration of sulpha drugs inhibits
choline deficiency include reduced weight gain,                growth of the bacteria that synthesize folic acid in
rough hair coat. decreased red blood cell count, and           the gut. The need for supplementation may depend
unbalanced and staggering gates.                               on the use of drugs in a particular herd.
                                                          46
Menadione                                                   Pantothenic Acid
   Menadione, a more common name for vitamin K,               Pantothenic acid was formerly known as vitamin
is involved in calcium and vitamin D metabolism.            B5• As a component of one specific co-enzyme, co-
The blood requires vitamin K to form clots; a               enzyme A, pantothenic acid is important for fat and
deficiency results in prolonged clotting time and in        carbohydrate metabolism. Deficiency symptoms
severe cases, haemorrhaging. A compound called              include slow growth, diarrhea. dry skin, reduced
dicoumarol, found in spoiled sweet clover, reduces          immune response, and a peculiar behaviour known
the blood's prothrombin content and raises the              as 'goose stepping' in the hind legs.
requirement for menadione supplementation.
Certain mycotoxins in the diet may have the same              The name pantotbenic acid is derived from the
effect. Sulpha drugs can reduce bacterial synthesis         Greek word 'pantothen' which means 'from every-
and place greater pressure on dietary sources of            where'. As its name suggests, pantothenic acid is
menadione.                                                  found in many feed ingredients. Barley, wheat, and
                                                            soybean meal are good sources, but biological
  Vitamin K is supplemented in diets by a number            availability in corn and sorghum is low. Although
of compounds, such as menadione                             cereal grains are good sources of the vitamin, swine
dimethylpyrimidinol bisulphite (MPB-22.7%                   diets are generally supplemented with the synthetic
menadione), and menadione sodium bisulfite                  form called d-calcium pantothenate (88.3% d-
complex (MSBC-33.0 or 16.5% menadione).                     pantothenic acid). The form called di-calcium
Alfalfa meal is a rich. natural source of menadione.        pantothenate provides 44.0% d-pantothenic acid,
                                                            since the I-isomer is biologically inactive.
Niacin
   Another name for niacin is nicotinamide, which is Pyridoxine
the biologically active form of the vitamin. Niacin      Pyridoxine was formerly called vitamin 86.
is a relatively stable compound not adversely          Pyridoxine, like other vitamins, plays an important
affected by heat. It is involved in the metabolism of role in enzyme function. Over 50 pyridoxine-
many dietary components including fats, carbohy-       dependent enzymes have been identified.
drates, and proteins. Because of niacin's central      Pyridoxine is particularly important in amino acid
role in metabolism, nutritionists are careful to avoid metabolism and the formation of neurotransmitters,
a deficiency. Deficiency symptoms include reduced the chemical messengers of the nervous system.
weight gain, dry skin, poor appetite, and diarrhea.    Deficiency symptoms include reduced appetite and
                                                       growth rate. In severe cases, nervous disorders.
   Cereal grains contain considerable quantities of    including convulsions, occur.
niacin but its availability in corn, wheat, oats, and
sorghum is poor. Conversely, the niacin in               Cereal grains are good dietary sources of
soybeans is highly available. Niacin is readily        pyridoxine. Because of the wide distribution of
available in synthetic form at a reasonable cost for   pyridoxine in conunon feed ingredients, pyridoxine
supplementation in swine diets.                        deficiency in swine is very rare and most diets, with
                                                       the possible exception of starter diets, are not
   Tryptophan in the body can act as a precursor for   normally supplemented.
niacin; diets containing excess tryptophan require
less niacin. A diet marginal in tryptophan will place Riboflavin
greater pressure on the need for niacin. However,        Riboflavin is a critical co-factor in energy
tryptophan is the second or third limiting amino       metabolism and also plays a role in fetus
acid in diets based on corn, so dietary tryptophan is  development. Formerly called vitamin B2,
unlikely to provide appreciable amounts of niacin.     riboflavin is supplemented in all swine diets
Theoretically, 50.0 mg of tryptophan will provide      because cereals are a poor natural source. Whey
 1.0 mg of niacin.                                     powder, commonly used in starter diets, is the only
                                                       47
ingredient used in swine diets that is a good source             The essential fatty acids (EFA) perform a number
of riboflavin. Supplemental riboflavin is available            of functions. They become an important part of cell
commercially as a 96% pure powder.                             membranes. thus help to ensure that cells function
                                                               normally. They are also involved in the synthesis of
  Riboflavin deficiency results in anestrus in gilts.          a group of very important hormones called
cataracts, a stiff gait, and reduced growth rate. It is        prostaglandins. These hormones are required for
also teratogenic, meaning a deficiency results in the          many body functions, including reproduction.
birth of deformed piglets.
                                                          The requirements for essential fatty acids in pig diets
Thiamine                                                are not well established. The recent interest in the
   Thiamine was once called vitamin 81• A pig's         role of essential fatty acids in human health has revived
requirement for thiamine increases in proportion to     the interest in the metabolism and utilization of
its energy intake because thiamine is required to       essential fatty acids by pigs as well. There is, for
maintain normal energy metabolism. Symptoms of          example, considerable debate on the efficiency with
deficiency include depressed appetite, vomiting,        which pigs can convert linolenic acid to other
depressed body temperature, and a slower than           polyunsaturated fatty acids that belong to the family
normal heart rate. Deficiency (for research             of omega-6 fatty acids and that are required for normal
purposes) can be induced by feeding raw fish,           animal metabolism and reproduction. Some studies
which contains the enzyme, thiaminase, that             suggest that the ratio of omega-6 to omega-3 fatty
destroys thiamine.                                      acids in the diet is as important for the conversion of
                                                        these omega-3 fatty acids as the levels of these fatty
  Grains are generally good sources of thiamine.        acids in the diet itself.        An essential fatty acid
Thiamine is present in the germ and hull portion of     deficiency is unlikely in normal diets for growing
cereals so wheat germ and bran contain very high        swine because the requirement appears to be very low
levels of thiamine. Flour, because of its limited       (approximately O. l - 0.3% of the diet) and because
quantity of wheat germ and bran, is a poor source of vegetable fats tend to be good sources. Even the small
thiamine. Diets are not generally supplemented          amount of fat present in wheat or barley apparently
with thiamine because most common ingredients           supplies more than sufficient EFA to meet
are rich sources of the vitamin. Some nutritionists     requirements.
believe supplementation is required, especially in a
young pig's diet, so small amounts (1 mg/kg) may          The actual requirements for EFA by sows and boars,
be added to starter diets. Synthetic thiamine is        however, are difficult to establish. The synthesis of
usually provided as thiamine hydrochloride (89%         some important reproductive hormones depends on
thiamine) or thiamine mononitrate (92% thiamine).       EFA: the requirement for the breeding herd is
                                                        suspected to be substantially higher than it is for
Essential Fatty Acids                                   growing   animals. Because EFA is stored in body fat
  As indicated previously, dietary fats are made up     reserves,  any signs of deficiencies in sows may not
of sub-units called fatty acids. At least one, linoleic occur   until the third or fourth parity. Based on a
acid, is essential in a pig's diet. Linoleic acid, a    factorial estimation, sows may require 0. 75% linoleic
member of the omega-6 family of fatty acids, is a       acid in the diet. If this estimate is indeed correct, a
dietary essential because it performs a specific role   barley-soybean meal based diet may be deficient in
and cannot be synthesized from other compounds.         linoleic acid. No estimates are available for the
Linolenic acid, a member of the omega-3 family of       requirements    for linolenic and arachidonic acid in
fatty acids, may also be required in the diet. A        swine.
third fatty acid, arachadonic acid, also a member of
the omcga-6 family of fatty acids, can be consid-                An EFA deficiency will impair growth rate and feed
ered a semi-essential fatty acid. Pigs require                 utilization, cause dry, scaly skin, and possibly result
arachadonic acid but can synthesize it from linoleic           in hair loss. Reproductive performance also suffers.
acid if supplied in sufficient quantities in the diet.         One experiment demonstrated that EFA deficiency
                                                          48
resulted in impaired water holding capacity of muscle,           N.F.l.A. 1981. Specialty ingredients handbook. Na-
and that heart muscle structure and colour changed.              tional Feed Ingredients Association. West Des Ma-
                                                                 rines, IA.
  Vegetable fats are rich sources of EFAs. The rea-
son cereal-based diets present a deficiency risk is be-          NRC 1980. Mineral tolerance of domestic animals.
cause barley and wheat contain much less fat than com.           National Academy of Sciences. Washington, D.C.
The recent practice of adding vegetable oil, or whole
seed canola or soybeans to sow diets should remove               NRC 1988. Nutrient requirements of swine. 9th
all concern about a possible deficiency. The fact that           Edition. National Academy of Sciences. Washing-
the requirement has not been defined, however, leaves            ton, DC.
some room for question.
                                                                 Noblet, J. and Y. Henry. 1991. Energy evaluation
Water                                                            systems in pig diets. Manipulating Pig Production
   Water is often called the 'forgotten nutrient' because        Ill. Australasian Pig Science Association, Attwood,
it is so often ignored. Yet, water is as important as            Victoria, Australia. pp. 87-103
any other dietary component and must be supplied in
sufficient quantities to ensure maximum productivity.        Noblet, J. and J.M. Perez. 1993. Prediction of di-
Water is critical to life and good health. For detailed      gestibility of nutrients and energy values of pig diets
information on water, refer to chapter 12.                   from chemical analyses. J. Anim. Sci. 71: 3339-
                                                             3398.
                                                            49
50
 4. INGREDIENTS
  The science of nutrition, as it applies to diet              feeding pigs is limited to using one or two grains
formulation, involves essentially five steps, as               (wheat, barley, corn) and one protein source (soy-
outlined in chapter 5: define feeding objectives,              bean meal), it would be a worthwhile exercise to
establishment requirements needed to achieve the               study diets utilized in other parts of the world,
objectives, select ingredients, formulate diets and            where these ingredients represent less than half of
evaluate diets once made. This chapter focuses on              the total diet, the remainder consisting of other crop
ingredients and how they might be used in diets.               (eg. lupins, peas, canola, tapioca) or animal (blood
                                                               meal, meat and bone meal, feather meal) products.
  In recent years, swine nutritionists have made
huge advances in defining the pig's nutrient require-            Nutrient based diet formulation, or formulation
ments and further developments in this area are                on the basis of nutrients irrespective of source, is
almost a certainty. Progress in defining the nutrient          not new, but a surprisingly significant emphasis on
specifications of ingredients has been somewhat                ingredients remains in our culture. While com and
slower and now provides a serious challenge to                 soybean meal may represent a standard against
commercial nutritionists. The precision with which             which other ingredients may be compared, and
we can define nutrients supplied to the animal                 therefore give a level of comfort to pork producers
appears to be considerably less than that with which           due to experience and tradition, it is essential to
we can define nutrients required by the pig. The               recognize that equivalent performance can be
two major challenges relate to nutrient variability            achieved with many other ingredients.
and nutrient availability. Consequently, to consist-
ently achieve the level of animal performance                    Therefore, ingredients should be selected on the
expected on an ongoing basis, and to do so at the              basis of composition, cost, availability and palat-
lowest possible cost, nutritionists must address these         ability. There is nothing magical about one ingredi-
two issues. Failure to do so will result in either             ent versus another, provided resulting diets are
excessive feed costs or variable animal performance            properly formulated.
or both!
                                                                 Composition is the first consideration. ls the
Ingredient Versus Nutrient                                     nutrient composition of an ingredient known with
  Pig diets should be formulated primarily from the            some reasonable degree of accuracy? If so, the
perspective of nutrients and less so on the basis of           ingredient can be used to its maximum level with
ingredients. Over many centuries of domestication,             confidence. If not, then nutritionists tend to be
the pig has demonstrated a marvellous ability to               more conservative and use less than they otherwise
perform successfully using a diverse array of ingre-           might.just to avoid unpleasant surprises!
dients; the pig is an omnivore, meaning it has the
ability to utilize feedstuffs of animal or vegetable
origin. The history of the pig is filled with exam-
ples of this culinary diversity.
                                                          51
Table 4-1. Consistency of Various Protein Sources Used in Swine Diets.
Animal Sources
Fish Meal                  5                  62.7               60.3 - 66.6                    2.72
Meat Meal*                29                  54.2               49.8 - 58.5                    2.04
"This analysis contains data from samples described as both meat meal and animal or feeding tankage.
Example of variation in protein content observed in commercial samples of the ingredients listed. Above example
refers to samples collected and analysed during April and May, J 995.
Supplied by Federated Co-operatives Ltd .. , Saskatoon, Saskatchewan.
  Based on nutrient composition, can the ingredient          associated with processing of offal is resulting in a
be included in the diet and still meet the needs of          more consistent product than was possible in the
the pig? For example, corn can be used in place of           past.
wheat, or vice versa, in a starter diet, because they
are both high energy grains. Oats would not be                 Does the ingredient contain any anti-nutritive or
used to replace either wheat or corn, because they           toxic compounds? For example, the old rapeseed
are too low in energy.                                       meal contained glucosinolates that reduced feed
                                                             intake and impaired animal health. The new canola
   Is the composition consistent? As a rule, proteins        meal has essentially eliminated such problems.
derived from vegetable sources, such as soybean
meal or canola meal, are relatively consistent from            Based on composition, is the product economical
batch to batch and from supplier to supplier. Con-           compared to other available ingredients? Cost
versely, protein supplements manufactured as part            should be considered in terms of profit, not price
of the meat processing industry tend to be Jess              per tonne. If an ingredient can reduce the overall
consistent. One can obviously use a consistent               cost of production, it should be used. If not, a more
product with a greater degree of confidence. Table           economical ingredient should be selected.
4-1 illustrates this point. Quality control data
supplied by the Feed Department of Federated Co-               Finally, is the ingredient appealing to the pig?
operatives Ltd. shows that products of plant origin          This is a difficult question to answer, because pigs
tend to be relatively consistent, as indicated by a          have a limited vocabulary! Unfortunately, we tend
smaller standard deviation (a measure of variabil-           to think we know what is attractive to the pig. It is
ity). Products of animal origin tend to be less              very important that one does not use human values
consistent and have a larger standard deviation.             to select ingredients for pig feed. Pigs eat many
This does not mean that they should not be used,             things that humans will not, and indeed. there are
but rather that more caution needs to attend their           things that humans will eat that pigs find
inclusion in the diet. Advances in the technology            unappealing.
                                                        52
  The basis for diet formulation is knowledge.             This chapter discusses the use of many ingredients
Lack of knowledge means uncertainty and this             in swine diets. Where possible, recommendations
translates into errors. The flow diagram in Figure       for inclusion in the diet are provided. In providing
4-1 helps determine ingredient usage.                    these guidelines, certain assumptions have been
                                                         made. For example, replacement of current ingredi-
                                                         ents with an alternative should not change the
                     INGREDIENT
                                                         overall nutrient composition of the diet. If so, a
 Is its nutrient composition, including availability,    depression in performance will probably occur.
                       known?
                                                           The following section contains a discussion on the
                                                         utilization of common and some not so common
                                                         ingredients in swine diets. Detailed feedstuff
                                                         composition tables appear in the Appendix. For
               Yes                 No                    further information on a particular ingredient, useful
                                     �                   articles are listed at the end of the chapter.
     Can the pig's nutrient              Determine
      requirements be met                  nutrient      Animal Sources
        if this ingredient               composition       The quality of feed ingredients derived from the
                                                         meat processing industry has improved consider-
            is used?
                                                         ably in recent years. Improvements in quality
                                                         control have resulted in products that are superior to
                                                         that sold even ten years ago. As with any product,
                                                         quality varies among suppliers and attention to good
               Yes                 No
                                                         purchasing practices, including regular feed testing,
                I
       Is it economical?
                                      �
                                           Reject
                                                         is advised.
                                                        53
Blood Meal                                                    spray drying process, the lower temperatures and
  Blood meal, as its name implies, is a high protein          shorter retention times, as compared to flash drying
ingredient manufactured from animal blood col-                results in a final product that is superior in terms of
lected from abattoirs. The nutrient composition of            amino acid availability. Spray-dried blood meal is
blood meal will vary a great deal among suppliers,            considerably more expensive than flash dried
due primarily to two factors: the method of process-          product, and is used most commonly in starter diets,
ing and the nature of the source material used in the         especially those employed for the young pig. Phase
manufacturing of the product. Book values will not            I diets may contain as much as 2% to 3% while
suffice, unless extremely conservative values are             Phase III diets may contain up to 5% to 7% spray-
employed. For example, one study of five different            dried blood meal.
batches of blood meal revealed that total lysine
varied from 7.7 to 10.6% of the dry matter! Differ-             The use of blood meal requires consideration be
ences in digestibility of lysine would further in-            paid to sulphur amino acid and branched chain
crease the variability of the product.                        amino acid levels. There is research demonstrating
                                                              that starter diets containing blood meal may be
  A variety of processes can be used to convert raw           deficient in methionine, while higher levels of
blood (approximately 20% dry matter) to the dry               blood meal (approaching 10%) in growout diets
powder useable in animal feeds. Vat drying is                 may result in problems with Jeucine levels being
essentially a batch process and results in the product        excessive or isoleucine levels being inadequate.
of lowest nutritional quality; in North America, vat
drying represents a very small portion of the total             Unlike meat and bone meal, blood meal is not a
blood meal available to the livestock industry. Vat-          particularly good source of calcium or phosphorus.
dried blood meal is not recommended for use in                The energy level in blood meal appears to be highly
swine diets, as the availability of nutrients is low;         variable, so individual suppliers should be asked to
for example, the cligestibility oflysine is probably          provide DE or ME values for their product.
less than 20%.
                                                         54
Additional Reading and References                           greatly increase the processing time needed; how-
Miller, E.R. 1990. Blood meal: Flash dried. in              ever, excessive treatment periods, even at lower
Non-traditional feed sources in swine production,           temperatures appear lo resull in an inferior final
P.A. Thacker and R.A. Kirkwood, eds. London:                product.
Butterworth. pp. 53 - 60.
                                                           55
Fish Meal
  Fish meals are high in protein (50 to 75%) and                        Fish Meal: Herring
amino acids. The protein is of good quality ( espe-                 DE                         3725
cially high in methionine) and is generally highly                  Crude Protein              71.0
digestible. Due to fat content (approximately 10%)                  Lysine                     5.82
the DE content is comparable to or higher than that                 Digestible Lysine          4.95
in soybean meal. The levels of most minerals,                       Digestible Threonine       2.41
particularly calcium and phosphorus are similar to                  Digestible Tryptophan      0.58
or higher than those in other protein sources. Phos-                Calcium                    2.75
phorus availability in fish meals is high.                          Phosphorus                 1.75
                                                        56
  When large quantities of fish meal are used in         these processing plants can obtain a high quality of
starter pig diets, close attention should be given to    protein supplement at a relatively low cost and
the level of lactose (normally supplied by milk          thereby increase the efficiency and profitability of
products) and minerals (calcium and phosphorus) in       their swine operations.
the diet. A concern with using fish meal in finish-
ing pig diets is its effect on meat quality. Problems          The potential feeding value of fish silage is
may arise with a fishy taint in carcasses from pigs          determined mainly by the quality of the material
fed fish meals. This can largely be attributed to the        being ensiled. Fish silage produced using a high
oil content of fish meal. In the finisher diet, the          percentage of whole fish will have a higher nutri-
amount of oil derived from fish products should not          tional value than will silage produced using offal.
exceed 1%.                                                   In addition, the type of fish used will affect the
                                                             quality of the silage produced. For example, silage
Additional Reading and References                            based on white fish is different from that based on
Gore, A.M., R.W. Seerly and M.J. Azain. 1989.                herring and it is important to differentiate between
Menhaden fish meal and dried whey levels in starter          these when discussing their nutritional value.
diets. Univ. Georgia Swine Res. Rep. P. 11.
                                                               White fish silage has a dry matter content of about
Stoner, G.R., J.L. Nelssen and R.H. Hines. 1988.             20%. On a dry matter basis, it contains approxi-
Replacing dried skim milk with select menhaden               mately 70% crude protein, 3% ether extract and
fish meal in a high nutrient dense diet. Kansas State        16% ash. In contrast, herring fish silage has a dry
Univ. Swine Res. Rep. P. 57.                                 matter content of about 35% and on a dry matter
                                                             basis, it contains only 43% crude protein and 8%
Stoner, G.R., J.N. Nelssen and R.D. Goodband.                ash. However, it has an ether extract content of
1989. Effect of fish meal quality on the growth              over42%.
performance of weanling pigs. Kansas State Univ.
Res. Rep. P.70.                                                The energy and mineral content and amino acid
                                                             profile of fish silage closely resembles that of fish
Wiseman, J., S. Jaggert, D.J.A. Cole and W.                  meal made from the same type of raw material (see
Haresign. 1991. The digestion and utilization of             section on fish meal). As a consequence, fish silage
amino acids of heat treated fish meal by growing-            would appear an excellent source of protein and
finishing pigs. Anim. Prod. 53:215-225.                      minerals for use as a supplement to cereal grains.
                                                             However, some of the concerns that apply to fish
Fish Silage                                                  meal ( oxidizing oils, thiamin supplementation.
  Fish silage is produced by adding organic acids to         contamination with mercury, effect on carcass
whole or parts of fish. The addition of acid acti-           quality) apply to fish silage as well (sec fish meal).
vates enzymes present in the raw fish and thus
stimulates the break down of the tissue protein. The           Fish silage can be used quite successfully in swine
resulting product has a nutrient content similar to          rations. The results of a research trial conducted at
fish meal except that fish meal has a dry matter             the University of Georgia using weanling pigs fed
content of about 90% while fish silage has a dry             either 0. 3, 6 or 9% fish silage for six weeks are
matter content in the range of 15-30%.                       presented in Table 4-2. It can be seen that the
                                                             performance of weanling pigs fed diets containing
  The process of making fish silage offers the               3 or 6% fish silage was not significantly different
potential of utilizing wastes from the fishing indus-        from the control group while those fed diets con-
try in areas where the quantity of waste material is         taining 9% fish silage gained weight at a slower
insufficient to justify the production of fish meal.         rate. The major factor responsible for this reduction
Producers with operations located in the vicinity of         in growth rate appeared to be a reduction in intake.
                                                        57
Table 4-2. Performance of Weanling Pigs Fed Diets                 There would appear to be some problems when it
Containing Graded Levels of Fish Silage.                        comes to feeding fish silage to breeding stock. The
                                                                results of one experiment in which fish silage was
                                  Fish Silage(%)
                                                                fed to sows during gestation are shown in Table 4-4.
                       0        3        6     9                Preweaning mortality has been shown to be signifi-
Daily gain (kg)        0.42     0.40     0.43    0.39           cantly higher when diets containing 6% fish silage
Daily feed (kg)        0.87     0.89     0.91    0.80           arc fed to sows during gestation. The reason for
Feed conversion        2.07     2.22     2.12    2.07           this increase in mortality has not been dctennined.
                                                           58
Meat and Bone Meal                                             content of meat meal is usually about 8%, since any
  Meat and bone meal, or often referred to as meat             additional fat in the raw material is removed in the
meal, is a by-product from the animal packing                  processing tallow.
industry. The crude protein content of meat and
bone meal (50%) is slightly higher than that of                  The major problem with meat and bone meal is
soybean meal while its amino acid profile is surpris-          variability of nutrient composition. For example,
ingly similar to that of soybean meal although                 one survey of only 17 samples revealed lysine
containing less tryptophan. However, the amino                 content ranged from 2.38% to 3.86%. Tryptophan
acids in meat and bone meal are generally less                 varied even more, from 0.20% to 0.69% and
available to the pig (see below). The energy content           threonine ranged from 1.81 % to 2. 72%. Amino
of meat meal varies due to differing mineral con-              acids were not the only nutrient to vary. Calcium
tent, but overall is less than that of barley or canola        content was 0.9% to 15.8%. This lack of uniformity
meal. The calcium and phosphorus content is a                  makes it a difficult product to use to maximum
bonus since phosphorus is an especially expensive              advantage in swine diets.
nutrient. Care must be taken to consider the sodium
and chloride (salt) content of meat meals. The salt               The degree of variability will depend on a number
content has been reduced in recent years. but is still         of factors. If the meal is produced from a single
sufficiently high to require adjustment of added salt,         source of relatively constant composition, then the
especially in the diets of young pigs.                         meal will reflect this in being consistent. However,
                                                               it is made from a variety of ingredients, such as
                                                               whole carcasses, offal and blood, and contains beef,
          Meat and Bone Meal                                   poultry and swine, the final product will likewise be
      DE                          2825                         less consistent. If this can be considered in the diet
      Crude Protein               50.0                         formulation, no harm will be done. For example,
      Lysine                      2.70                         hard offal, which includes bones, heads and hooves
      Dig. Lysine                  1.89                        is a less desirable component, because it results in a
      Dig. Threonine              1.09                         meat meal with a poorer amino acid profile due to a
      Dig. Tryptophan             0.17                         high collagen content. Soft offal, consisting of gut
      Calcium                     9.50                         material, tends to produce a superior product with a
      Phosphorus                  4.70                         more desirable amino acid profile.
                                                          59
example, increasing the cooking temperature from             Brooks, P. 1991. Meat and Bone meal: the under-
I 25°C to I 50°C has been found to reduce lysine             utilized raw material. Feedstuffs, volume 63,
availability from 84% to 38%.                                number 27, 1991. pp. 13-15, 22.
  As mentioned above, the amino acid availability            Cromwell, G.L., T.S. Stahly and H.J. Monegue.
of meat meal tends to be lower than comparable                1991. Amino acid supplementation of meat meal in
values for cereal grains and vegetable protein               lysine-fortified, corn-based diets for growing-
sources, For example, various studies have con-              finishing pigs. J. Anim. Sci. 69:4898-4906.
cluded that lysine is 54-67% available for meat and
bone meal, as compared to 81-87% for soybean                 Haugen, E.W. and J.E. Pettigrew. 1985. Apparent
meal and 71 % for fish meal. However, some                   digestibility of amino acids in meat meal as affected
individual meat and bone meal samples have lysine            by manufacturing variables. Proc. 46th Minnesota
availabilities equal to that of soybean meal.                Nutr. Conf., Minnesota Agric. Expt. Stn., pp. 154-
                                                             169.
  A potential concern if including meat and bone
meal in pig feeds is its contamination with harmful,         Knabe, D.A., D.C. LaRue and E.J. Gregg. 1987.
disease causing organisms such as salmonella.                Apparent digestibility of nitrogen and amino acids
However, various recent surveys demonstrate that             in protein feedstuffs by growing pigs. J. Anirn. Sci.
the incidence of salmonella contamination in                 67:441-458.
properly processed meat and bone meals (proper
temperature, no contamination of processed with               Leibholz, J. 1979. Meat meal in the diet of the
unprocessed material) is very low. In fact, some             early-weaned pig. III. Meat meal quality and the
surveys demonstrate that feedstuffs of vegetable             processing of meat meals. Anim. Feed Sci. Tech.
origin, such as cereal grains and soybean meal can           4:53-61.
be contaminated with salmonella as well. A further
point worth noting is that there are a whole range of        Whole Milk
types of salmonellas. Only a very limited number               Whole cow's milk is usually not included in swine
of salmonella strains are considered harmful to pigs         rations because its value in human diets makes it too
or humans.                                                   expensive. However, it is perhaps nature's most
                                                             perfect feed and can be used very successfully in
  Success in using meat and bone meal will depend            swine rations. Milk provides more essential nutri-
on one's ability to define nutrient composition.             ents than any other feed ingredient. Whole milk is
High quality meal, purchased from a good supplier            very easily digested and is extremely palatable. The
with a sound quality control program, can be used            amino acid balance of milk protein is excellent and
at high levels in market hog diets ( 10-15%) as well         because of its high protein quality, lower levels of
as those for sows (5-10%). Otherwise. meat and               crude protein can be fed when using whole milk
bone meal should be limited to 5-7% of grower and            than the levels commonly advised in feeding stand-
gestation diets, 3-5% of lactation diets and 2% or           ards.
less in starter diets.
                                                               Milk provides energy in its milk fat and milk
Additional Reading and References                            sugar (lactose). It is an outstanding source of
Batterham, E.S., R.E. Darnell, L.S. Herbert and E.J.         calcium and a good source of phosphorus. It is high
Major. 1986. Effect of pressure and temperature              in vitamin A, rich in riboflavin and is a good source
on availability of lysine in meat and bone meal as           of niacin, thiamine, vitamin B-12 and other B-
determined by slope-ratio assays with growing pigs,          complex vitamins. However, it is not perfect. The
rats, chicks, and by chemical assay. Brit. J. Nutr.          iron content is extremely low and it is a poor source
55:441-453.                                                  of vitamin D. Therefore, it is important that a
                                                             vitamin-mineral premix be utilized when whole
                                                             milk is being fed.
                                                        60
  A nutritional breakdown of whole cow's milk is
                                                                          Dried Skim Milk
shown below. Basically five pounds of whole milk
                                                                    Digestible Energy            3850
will supply the same amount of nutrients as one
                                                                    Crude Protein                33.4
pound of complete feed.
                                                                    Lysine                       2.43
                                                                    Digestible Threonine          I .31
                Whole Milk                                          Calcium                      1 .20
       Digestible Energy            660                             Phosphorus                    l.00
       Crude Protein               3.50
       Lysine                      0.28
       Digestible Threonine        0.13                          Because of price, skim milk is not a commonly
       Calcium                     0.12                       used ingredient in swine diets. However, skim milk
       Phosphorus                  0.09                       may be diverted to animal feed if excess supplies
                                                              develop or if a given shipment fails to meet certain
                                                              quality standards. Since these standards may relate
  An example of a feeding regime using whole milk             to factors of Little nutritional importance, a real
is shown in Table 4-5. Best use of the protein in             opportunity exists. However, if the milk powder
whole milk can be made if no other protein supple-            has been over-heated during drying, then its nutri-
ment is used in the ration. However, since less dry           tional value will be compromised.
feed is to be fed, it is recommended that higher
levels of vitamin-mineral premix be included in the             Generally, even skim milk powder sold at distress
diet.                                                         prices is economical only in creep and starter diets,
                                                              where it can make up to 20 - 30% of the total
                                                              formula. Pigs of all ages can be fed skim milk.
Table 4-5. Replacement Scheme for Using Whole                 Their feces may become loose as they adjust to the
Milk in Swine Diets.                                          lactose in the milk but this is usually a temporary
                                                              problem and will disappear in a few days. The
Weight of               Meal                  Milk            problem of milk intolerance in pigs tends to be
Pig (kg)                (kg)                (Litres)          overestimated.
  18                      0.84                 1.80
  23                      0.86                 3.20           Additional Reading and References
  34                      1.28                 4.10           Kornegay, E.T., H.R. Thomas and C.Y. Kramer.
  45                      1.50                 5.00           1974. Evaluation of protein levels and milk prod-
  68                      2.12                 5.90           ucts for pig starter diets. J. Anim. Sci. 39: 527-535.
  91                      2.36                 7.00
                                                              Owsley W.F., D.E. Orr and L.F. Tribble. 1986.
'Meal should contain 95% cereal grain and 5% vitamin-         Effects of nitrogen and energy source on nutrient
mineral premix. Additional protein supplements are not        digestibility in the young pig. J. Anim. Sci. 63: 492-
required.
                                                              496.
                                                              Whey
Dried Skim Milk
                                                                When whole milk is treated with the enzyme
   Dried Skim Milk is of course skim milk which
                                                              rennet, the protein casein is precipitated and takes
has had the water evaporated from it. The only
                                                              with it most of the fat and about halfof the calcium
major difference between dried whole milk and
                                                              and phosphorus. The remaining liquid is called
dried skim milk is that in skim milk, most of the fat
                                                              whey. Sweet whey arises from the production of
and fat soluble vitamins have been removed so that
                                                              Swiss- or cheddar-type cheeses and acid whey from
all other components of the diet are increased
                                                              cottage cheese production. About 9 kg of sweet
proportionally.
                                                              whey or 6 kg of acid whey will be produced per
                                                         61
kilogram of cheese. Sweet whey contains slightly                In older pigs, concern is often expressed about
more crude protein ( 12 - 14 vs IO - 12% ), more              their ability to utilize the lactose in whey. It is
lactose (68 - 72 vs 60 - 65%) and less ash (7.5 - 9.0         known that as the pig ages, and no longer consumes
vs 9.5 - 1 l.5%) than acid whey. As its name im-              lactose, the special digestive enzyme called lactase,
plies, it also contains less lactic acid (1.5 - 3.0 vs        that breaks down lactose in the gut, will decrease.
5.5 - 7.5%). Whey contains almost all of the                  This is no surprise. since lactase is used solely to
lactose from milk and significant portions of the             degrade lactose, and if it is not present in the diet,
mineral fraction as well. Since casein has been               there is little reason for the gut to continue to
removed, the major protein in whey is                         produce it. Consequently, farmers have become
lactoglobulin, which fortuitously is of good quality.         reluctant to utilize whey in the diet of growing or
Lactose is desired in diets of newly weaned pigs,             finishing animals, even if a local milk processing
due to its digestibility. However, some commercial            plant can supply fresh whey at an economical price.
whey powders have had some of the lactose re-
moved so the content may vary from 35 to 58% of                  The truth of the matter is that pigs can tolerate
the dry matter. As the amount of lactose increases,           lactose quite well. Research in Wisconsin demon-
the protein content decreases from 26 to 16%.                 strated that the pig can tolerate up to 30 % lactose in
                                                              its diet. Since whey is about 70- 77 % lactose, this
  In its crude form, whey is a liquid; more than              translates into about 40 % whey in the diet on a dry
90% of fresh whey is in fact water. Although liquid           matter basis. Indeed. the scientists also learned that
whey can, and is, fed to swine, most has been dried           removal of lactose from the diet of the older pig
before inclusion in the diet. On a dry matter basis,          will not diminish its ability to digest lactose in the
fresh whey is 70-77% lactose (milk sugar), 17%                future. The pig's adaptability, in terms of diet, was
crude protein, 9% ash (minerals), 1.0% calcium and            reaffirmed!
0.8% phosphorus. Although the true protein in
whey is of good quality, it must be recognized that             The major problem with feeding liquid whey to
up to 25% of the nitrogen in whey is present as non-          swine is the high moisture content, and the pig's
protein nitrogen. Therefore, nitrogen assays tend to          inability to consume sufficient dry matter to grow
overestimate the true protein content.                        rapidly. Thus. care must be taken to ensure that
                                                              sufficient dry matter from other sources is provided
  Dried whey is generally used in the diet of early
weaned pigs, as a source of highly digestible lactose
and protein. However, excessive utilization of                Photo 4-1.
whey can lead to diarrhea; the exact amount will
depend on other factors in the diet. Generally,
starter diets will contain no more than 15 - 20 %
whey, if for no other reason than cost. Concerns
about scouring often lead nutritionists to use skim
milk powder rather than exceed 20 % whey in the
diet. Again, cost becomes a limiting factor in such
diets, since skim milk powder is also very expen-
sive.
                                                         62
to pigs to support normal growth. If liquid whey is      Plant Sources
being fed to pigs, the dry feed can be adjusted to         Feeds derived from plants tend to be relatively
contain less protein and more mineral:vitamin            uniform in composition within species. This helps
premix. Since whey is rich m protein. less supple-       avoid the variability problems associated with feeds
mental protein would be required in the dry feed.        derived from animal sources. I lowever, ingredients
The premix must be increased since the pigs would        derived from plant sources have other problems
cat fewer kilograms of dry feed per day. but their       which ma) limit their inclusion in swine diets. For
daily requirement would remain the same. The             example. unprocessed oats arc too low in energy to
exact adjustment would depend on the amount of           be utilized as a major constituent in most swine
 whey being eaten. Since the final ration, including     diets. Barley, although higher in energy than oats. is
 proportions of whey. grain and premix will vary         of limited value to nursing sows and baby piglets if
 depending on pig size and amount of whey con-           maximum performance is to be recognized. Some
 sumed, a qualified nutritionist should be involved in   plant products contain anti-nutritional factors that
 setting up feeding programs on individual farms.        impair digestion. reduce appetite, or may compro-
 Liquid feeding systems, in which the dry feed and       mise the health of the pig.
 water are mixed prior to feeding, offer considerable
 potential for liquid whey feeding. Also, wet-dry         Alfalfa
 feeders have been used for the same purpose.               There are problems associated with alfalfa
                                                          (Medicago sativa) which limit its usefulness as a
Additional Reading and References                         feedstuff for pigs. The protein and energy are
Ekstrom, K.E., N.J. Benevenga and R.H. Grurnmer.          poorly digested and it contains toxic factors such as
1975. Effects of various dietary levels of dried          saponins and tannins which reduce growth rates of
whey on the performance of growing pigs. 1. Nutr.         animals fed diets containing alfalfa. However.
105: 846-850.                                             despite the negative factors present, there is still
                                                          interest in the use of alfalfa as a component of
Forsum, E. 1975. Whey proteins for food and feed          swine diets and research continues to attempt to
supplement. In Protein Nutritional Quality of Foods       overcome these problems. If this work is success-
and feeds (M. Friedman, ed.). pp. 433-470. Marcel         ful, the use of alfalfa in swine diets may increase.
Dekker. Inc .. New York.
                                                         63
  The primary factor limiting the use of alfalfa in             Table 4-6. Growth, Feed Intake and Carcass Charac-
swine diets is its low digestible energy content. The           teristics of Pigs (54 to 100 kg) Fed Alfalfa.
crude fibre content of alfalfa is extremely high
compared with grains and oilseeds. Since the pig                                                � DietaQ:'. Alfalf�
has a simple stomach of relatively small capacity, it           Criteria                      0.0   20.0 40.0 60.0
is less able to utilize crude fibre than are other types
of farm livestock. Therefore, the digestible energy             Daily Gain (kg/day)          0.86 0.73 0.63 0.41
content of alfalfa is approximately half of that                Daily Intake (kg/day)         3.0 3.0 3.2 2.7
found in common cereal grains.                                  Feed/Gain                     3.6 4.1   5.0 6.7
                                                                Dressing (%)                 77.9 76.2 75.4 75.2
  Alfalfa ranges from 12 - 22% crude protein (N x               Backfat Thickness (cm)        3.9 3.5 3.2 2.9
6.25). Unfortunately, the protein in alfalfa is not
very digestible. The high crude fibre content of                Powley et al., 1981, J. Anim. Sci. 53: 308-316.
alfalfa prevents the digestive enzymes from gaining
access to the soluble cellular proteins. As a result,
the protein in alfalfa is only about 50% digestible.          Poor palatability is one factor accounting for the
Alfalfa contains a good balance of amino acids and          reduction in performance when high levels of
a reasonable level of lysine. However, because of           alfalfa are included in the diet of the growing pig.
the high fibre level, the availability of the lysine in     Saponins are a bitter tasting compound present in
alfalfa is likely to be low.                                alfalfa. Recently, cultivars of alfalfa varying in
                                                            saponin content have been developed. It would
  Alfalfa is characteristically high in calcium.            appear that cultivars containing lower levels of
However, it has only a moderate phosphorus con-             saponins are more palatable and support higher
tent. When grown on phosphorus-deficient soils, it          levels of performance than do the traditional alfalfa
may be very low in phosphorus. Therefore, rations           varieties. Future research may allow for higher
containing high levels of alfalfa require supplemen-        levels of alfalfa to be incorporated into swine diets.
tal phosphorus to meet the pig's requirement and to         However, at the present time, it is not possible to
narrow the wide calcium:phosphorus ratio present            justify the use of high levels of alfalfa meal.
in this forage. Alfalfa is a good source of most
vitamins and is an excellent source of vitamins A, E              The subject of whether or not it is beneficial to
and K. However, the advent of relatively cheap                  include alfalfa in diets fed to sows is controversial.
sources of these nutrients added via the premix has             While some studies have supported its use, the
resulted in a reduction in the need for alfalfa in the          benefits can often be attributed to the vitamin E and
diet as a source of vitamins.                                   selenium supplied by the alfalfa meal. Since
                                                                synthetic sources of both vitamin E and selenium
   Alfalfa should not be used in diets fed to wean-             are available, there does not appear to be any
ling pigs. Its high crude fibre content and low                 necessity for the inclusion of alfalfa in gestation
digestible energy level are likely to limit growth and          diets. If it must be used, it would seem wise to limit
reduce the efficiency of feed utilization when fed to           the levels of alfalfa fed during gestation to a maxi-
pigs of this weight range. Higher energy feedstuffs             mum of25 - 30% of the diet.
should be used as the foundation for a high quality
starter diet.                                                     Alfalfa meal should not be fed to sows during
                                                                lactation. Feeding alfalfa meal during lactation will
  It is recommended that no more than 5% alfalfa                reduce the energy intake of sows resulting in re-
be included in the diet of grower pigs. The data in             duced milk production. However, some producers
Table 4-6 illustrate the adverse effects of alfalfa             include bulky ingredients such as alfalfa meal in the
meal when included in the diet of market hogs. The              diets of sows during prefarrowing and early lacta-
reduction in gain would appear to be the result of              tion to prevent constipation. ff this is the case,
insufficient dietary energy to meet requirements for            higher levels of wheat or fat are needed to maintain
maximum growth.                                                 a high energy content.
                                                           64
  In summary, alfalfa meal should not be included              Barley
in diets fed to starter and grower pigs and lactating            Swine producers and feed manufacturers who arc
sows because of its high crude fibre content and low           oriented towards com are sometimes unaware of the
digestible energy level. In rations fed to the gestat-         wide use of barley (Hordeum vulgare) as a swine
ing sow, a maximum of 25 - 30% of the diet is                  feed and are often sceptical about its use. However,
suggested.                                                     barley is an excellent feed for swine and millions of
                                                               pigs are raised annually on barley-based diets.
Additional Reading and References
Baker, D.H., B.G. Harmon and A.H. Jensen. 1974.                  Unfortunately, barley is not just barley, There is a
Value of alfalfa meal and wheat bran in diets for              tremendous amount of variability in the types of
swine during prefarrowing and lactation. J. Anim.              barley available for use in swine production. Barley
Sci. 39:325-329.                                               can be either two-rowed or six-rowed, hulled or
                                                               hulless, awned or awnless. In addition, variation
Powley, J.S., P.R. Cheeke, D.C. England, I.P.                  exists among the lysine and starch content of
Davidson and WH. Kennick. 1981. Performance                    barleys. Even barley of the same genetic back-
of growing finishing swine fed high levels of alfalfa          ground can vary greatly since growing and harvest-
meal: effect of alfalfa level, dietary additives and           ing conditions can have profound effects on the
antibiotics. J. Anim. Sci. 53:308-316.                         nutrient composition of barley.
Thacker, P.A., 1990. Alfalfa meal. Pages 1-12 In                 For example, the results represented in Table 4- 7
P.A. Thacker and R.N. Kirkwood eds. Non-tradi-                 indicate that differences in growth rates and feed
tional Feed Sources for Use in Swine Production.               efficiency between growing-finishing pigs fed
Butterworths Publishers, Stoneham, MA.                         samples of different barley varieties can be as much
                                                               as 609%. The observed differences in animal
Wallace, H.D., D.D. Thieu and G.E. Combs. 1975.                performance can largely be attributed to differences
Alfalfa meal as a special bulky ingredient in the sow
diet. Feedstuffs (Feb 3). pp. 24.
Table 4-7. Composition of Barley Varieties and Performance of Growing-finishing pigs (approximately 20 to
95 kg body weight) Fed Various Barley Varieties*
                                                                  Barley Variety
                               Boyer"       Camelot"           Clarkb     Harrington"       Hesk"        Steptoe"
  Chemical composition**
   Dry matter, %                 91.6          91.4             91.3           91.2          91.8          91.9
   Crude Protein, %               9.4          13.2             10.2           11.0           9.1           9.9
   Lysine,%                       .36           .44              .38            .42           .35           .36
  Animal performance
   Feed intake, kg/d             2.32          2.25             2.35           2.16          2.30          2.29
   Growth rate, kg/d              .78           .78              .80            .78           .79           .75
   Feed: Gain                    2.99          2.98             2.90           2.79          2.92          3.05
* The barley samples were supplemented with soybean meal and lysine to maintain similar lysine levels in all grower
(.75% lysine) and finisher (.60% lysine) diets and with vitamins and minerals. The inclusion levels of the barley
samples were 83.2 and 88.9% in the grower and finisher diet, respectively.
** As fed basis, analyses of the barley samples.
• Six-row winter barley varieties.
b Two-row spring varieties.
< Six-row spring variety.
Adapted from Michal et al., 1993. Washington State University Information Day Proc. Vol.8:69-74.
                                                          65
in unavailable energy content between the barley               Because of its lower energy content, barley finds
samples that were evaluated. Small differences in            only limited use in diets fed to starter pigs. How-
palatability (voluntary feed intake) were also               ever, when the cost of other cereals is very high it
observed. ln this study, Harrington barley (a two-           may be possible to include some barley in the diet
row spring barley variety) supported the best feed           of the weanling pig. Only good quality barley
efficiency while Steptoe (a six-row spring barley            should be used. Barley can be fed very successfully
variety) results in the poorest growth rate and feed         to growing pigs and in fact most pigs in Western
efficiency. As only one sample of each barley                Canada are raised on barley. To maximize growth
variety was evaluated, the differences in observed           rate, high energy cereals such as corn or wheat are
levels of animal performance may have been due to            often used in combination with barley. Growth
factors other than variety. For an estimation of the         rates in finishing pigs fed barley-based diets are
feeding value of barley, producers are thus encour-          generally similar to these fed corn or wheat-based
aged to have their barley tested before feeding it to        diets. This is because finishing pigs are generally
their pigs.                                                  able to compensate for reductions in diet DE
                                                             content with increases in daily feed intake in such a
  Barley is intermediate to wheat and oats as an             manner that the daily energy intake is dependent on
energy source for pigs. Its relatively high crude            diet DE content.
fibre content (5.1 %) is one of the major reasons for
the comparably low energy value. The digestible                 Barley can constitute the sole cereal grain in diets
energy (DE) content in individual barley samples             fed to sows during gestation. However, because of
may be predicts from the dry matter and crude fibre          its lower energy content, it may be beneficial to
content:                                                     include between 25-75% wheat in the diet of sows
                                                             during lactation.
DE (Kcal/kg of dry matter)= 4228 - 140 x Crude
fibre content(% in dry matter)."                                The performance of pigs fed barley-based diets
                                                             has been shown to be markedly improved as a result
  The protein content of barley is intermediate to
                                                             of pelleting. The reason for the improvement in
that of wheat and com and similar to the level
                                                             performance is a subject of much debate. Some
contained in oats. Relative to requirements, barley
                                                             researchers feel that the improvement is due to a
protein is generally low in lysine, isoleucine,
                                                             reduction in wastage, while others suggest that the
threonine, tryptophan and the sulfur containing
                                                             improvement is due to an increase in feed consump-
amino acids. The lysine content of barley is ap-             tion or to an increase in nutrient digestibility.
proximately equal to that of wheat. The amino acid
content of barley can be predicted using regression
                                                                Barley that is frozen or sprouted sometimes
equations based on crude protein. These equations
                                                             becomes available to the pork industry, and ques-
can be found in chapter 3.
                                                             tions arise regarding its feeding value. Recent
                                                             studies carried out at the University of Alberta
                                                             indicate that sprouting or frost damage will not
                  Barley                                     necessarily impair pig performance (Table 4-8). In
      Digestible Energy          3100                        this report, bushel weight fell as low as 42 lb, but
      Crude Protein               10.6                       all pigs performed equally, as compared to a control
      Lysine                     0.39                        diet based on normal barley.
      Digestible Lysine          0.27
      Digestible Threonine       0.23
      Digestible Tryptophan      0.10
      Calcium                    0.07
      Phosphorus                 0.35
                                                        66
Table 4-8. Utilization of Frozen or Sprouted Barley           Boyles, S.L., K.B. Koch. 1992. Feeding barley to
(Bonanza) by Growing Pigs'·                                   swine. ln: Feeding barley to cattle, swine and
                                                              poultry. North Dakota State University Extension
                        Control     Sprout     Sprout         Services. North Dakota Sate University of Agricul-
                        Frozen      Frozen     Frozen         tural and Applied Science, Fargo, North Dakota.
Grain Analysis
                                                              Michal, J.J., M.S. Han and J.A. Froscth. 1993.
Damage(%)                   0.1       18.7        6.9         Nutrient composition and feeding value of Boyer,
                            75        >75       >>75          Camelot, Clark, Harrington, Hesk and Steptoe
C. Protein (%)             12.1       13.3       11.0         Barley for growing-finishing pigs. Washington
                           11.4       11.7       12.2         State University Information Day Proceedings, Vol.
C. Fibre(%)                 6.5        7.3        7.3         8 pp. 69-74.
                            7.4        7.9        9.0
Bushel Wt (lb)              51         48         45
                            47         42         43          Barley: High Moisture
Pig Performance
                                                                     Barley is traditionally harvested after it has been
                                                                  field dried to a moisture content of 14-15%. This
Ave. Daily Gain (kg)      0.71        0.72       0.70
                                                                  drying is necessary in order to prevent deterioration
                          0.71        0.70       0.69
                                                                  of the grain during storage. Spoiled grain is useless
Ave. Daily Feed (kg)      2.27        2.24       2.23
                                                                  as an animal feed since molds growing on the grain
                          2.18        2.22       2.29
                                                                  may produce mycotoxins which cause poor growth
Feed Conversion           1.59        1.61       1.56
                                                                  and reproductive problems.
                          1.56        1.56       1.58
                                                                    Grain drying systems have been developed which
Adapted from Plett, and Aheme, 1987. Proc. Western
Nutrition Conf. pp. 128-131.                                      allow crops to be harvested at higher than normal
I Barley represented 80% of the diet in all cases. Initial        moisture levels and then dried for storage. This
pig weight averaged 20.8 kg; final weight was not                 artificial drying requires a lot of energy. As energy
specified.                                                        costs increase, it may become less attractive to
                                                                  harvest grain in this manner. Therefore, producers
                                                                  are looking for alternative harvesting methods. One
Additional References and References                              method that has generated considerable interest is
Batterham, E.S. 1990. Prediction of the dietary                   that of harvesting high moisture grain.
energy value of diets and raw materials for pigs. In:
(Wiseman, J. and DJ.A. Cole, Ed.) Feedstuff                         High moisture grains are preserved in a manner
evaluation. Butterworths, London, England, pp.                    similar to silage. In the absence of oxygen, acid
267-282.                                                          producing bacteria proliferate and produce acids by
                                                                  fermenting barley sugars and carbohydrates. The
Blair, R., B. Rakshit. J.M. Bell, V.J. Racz and K.A.              acids produced stop further bacterial action. The
Rosaasen. 1990. Dietary energy level for growing-                 ideal moisture content to ensile wet grain for swine
finishing pigs fed ad libitum. 1. Growth response.                appears to be in the 22-28% range. A moisture
Arch. Anim. Nutr. 9:793-804.                                      content greater than 28% leads to greater fermenta-
                                                                  tion, resulting in more acid production which tends
Blair, R., B. Rakshit, J.M. Bell. VJ. Racz and K.A.               to be less palatable to the pig. On the other hand,
Rosaasen. 1990. Dietary energy level for growing-                 the fermentation of grain containing less than 22%
finishing pigs ad libitum. 2. Carcass effects and                 moisture is relatively slow and the amount of acid
economical model of the responses. Arch. Anim.                    produced may not be sufficient to kill those organ-
Nutri. 9:805-813.                                                 isms that cause spoilage.
                                                             67
   A four-year research project conducted at Mon-                 Rates of acid application vary with the moisture
tana State University showed that high-moisture                 content of the grain and the intended length of
barley could be harvested an average of 12 days                 storage. The higher the moisture content of the
sooner than dry barley. By harvesting early, the risk           grain, the greater the amount of acid needed for
of losses due to high winds, rain, snow or frost is             proper preservation. Grain treated with an organic
minimized. Harvesting grain at a higher moisture                acid should have a moisture content of22% or less
content also results in higher yields compared to               when harvested. A higher moisture level would
traditional harvesting methods. Field losses are                make the chemical treatment expensive to use.
reduced because high-moisture barley does not
shatter like dry barley does. In the Montana State                The major disadvantage of acid-treated grain is
University tests, high moisture barley yielded                  the corrosive effect the acids have on most types of
 16. 7% more grain compared with similar fields                 metal in handling and storage equipment. The
which were harvested as dry grain. In addition,                 corrosion is usually most severe on surfaces in
high moisture grain helped to control weeds, espe-              contact with the grain or vapour for the first weeks
cially wild oats since reduced shattering resulted in           following grain treatment. It may also react with
fewer weed seeds left in the field to germinate.                concrete, especially if the concrete is newly cured
                                                                and unweathered.
   Although high moisture barley can be stored as
silage, several problems arise when it comes time to              In terms of chemical analysis, there is little
feed the product. Since the grain contains moisture,            difference between high moisture barley and regular
spoilage is always possible. Therefore, high mois-              barley on a dry matter basis (Table 4-9). High
ture grain must be fed soon after being taken from              moisture barley may be slightly higher in crude
storage. Untreated high moisture barley will heat               fibre than regular barley since it loses less hull
within hours of removal from the silo because of                during threshing. It may also be slightly higher in
microbial activity. Because of this, high moisture              protein content owing to the lighter and smaller
barley diets must be mixed on a daily basis and in              kernels saved by harvesting high moisture barley
amounts readily consumed over a short period of                 which are slightly higher in protein but lower in
time. Once a silo is opened, it is necessary to                 carbohydrate.
remove approximately 7.5 cm per day from the
exposed surface of the silo to control spoilage. If
this does not match the daily feed requirements of              Table 4-9. Chemical Analysis of High Moisture and
the pigs, some feed may be wasted.                              Dry Barley.
                                                           68
  High moisture barley may be used effectively as a               Several reports have indicated that high moisture
feed grain. In formulating diets with high moisture            barley may be more palatable than dry barley. This
grain, adjustments must be made for excess water.              improvement in palatability may be due to the fact
To convert wet grain to dry grain equivalent, divide           that high moisture grain is easier to roll and the
the dry matter content of the dry grain by the dry             rolling results in a flaky product which contains
matter content of the wet grain to establish an                very little in the way of fines or dust. This lack of
adjustment factor. This factor will tell how much              dust and fines in the ground product may be a factor
more high moisture barley must be fed to supply                in improving palatability. However, in most in-
amounts of nutrients as dry barley on a dry matter             stances, when differences in dry matter are consid-
basis (Table 4-10).                                            ered, consumption is about equal for high moisture
                                                               and dry barley. Since high-moisture grain has the
                                                               same feeding value as field dried grain when
Table 4-10. Amount of High Moisture Grain Neces-               compared on an equal dry matter basis, the decision
sary to Supply the Same Amount of Dry Matter as
Regular Barley (90% OM).                                       to use high-moisture grain should be made on the
                                                               basis of cost, storage and handling and not on
  % Moisture            High Moisture Barley (kg)              clifferences in nutritional value.
                                                          69
the glurnes does not occur, allowing the hull to be            fibre, may in fact be higher in hulless barley as
removed during the threshing process in a manner               compared to hulled barley. The beta glucan content
similar to that which occurs with wheat and many               of barley is one of the factors that contributes to the
other cereals. These so called hulless varieties of            reduced feeding value of barley as compared to
barley would appear to have considerable potential             wheat and com, especially in starter pigs.
for use in swine rations.
                                                                  For example, the results summarized in Table 4-
   ln general, hulless barley contains a higher protein         12 suggest that performance of starter pigs fed
level and a lower crude fibre content than hulled              hulless barley is similar to that in pigs fed regular
barley. All other nutrients are generally present at a         barley: pigs fed the hulless barley eat slightly less
slightly higher level in hulless versus hulled barley.         and used feed slightly more efficiently that the pigs
This is to be expected when the diluting effect of             fed the regular barley.
the hull is removed.
                                                               Table 4-12. Performance of Starter Pigs Fed Condor
                                                               Hulless or Samson Hulled Barley.
               Barley: HuUess
       DE                          3250                                                    Hulled            Hulless
       Crude Protein                                                                       Barley            Barley
                                    13.7
       Lysine                      0.54                        Feed intake, kg/d            0.90              0.85
       Calcium                     0.24                        Growth rate, kg/d            0.59              0.58
       Phosphorus                  0.37                        Feed: Gain                   1.57              1.47
                                                          70
significantly better than that of pigs given hulled            Beet pulp has proven to be a popular ingredient in
barley. This improvement in feed efficiency. in              sow diets. It is often added to prevent constipation
combination with the lower levels of soybean meal            problems. Five to seven percent beet pulp in sow
required in order to meet the pigs requirements for          diets tends to solve all but the most challenging
essential amino acids, may provide sufficient                cases of constipation.
incentive to encourage the incorporation of hulless
barley in swine diets, particularly when feed costs            It should be noted that the addition of beet pulp to
are high. There is no data available on the feeding          swine diets tends to reduce the digestibility of
value of hulless barley for breeding swine.                  energy and nitrogen. This is unlikely to be a practi-
However, given the benefits of increasing nutrient           cal problem, as only small quantities arc generally
density in nursing sow rations, serious consideration        used commercially.
should be given to the use of hulless barley in the
lactation diet.                                              Additional Reading and References
                                                             Graham, H., K. Hesselman and P. Aman. 1986.
Additional Reading and References                            The influence of wheat bran and sugar-beet pulp on
Aherne, F.X. 1990. Barley: Hulless. 1n: Non-                 the digestibility of dietary components in a cereal-
traditional Feed Sources for Use in Swine Produc-            based diet. J. Nutr. 116:242 - 251.
tion. Butterworths Publishers, Stoneham, MA,
USA. (Thacker, P.A. and R.N. Kirkwood, Ed.).                 Stebbens, H.R. 1991. The digestion and utilisation
                                                             of food fibre by growing pigs. Ph.D. Thesis,
Thacker. P.A., J.M. Bell, H.L. Classen. G.L.                 University of Edinburgh, UK.
Campbell, and B.G. Rossnagel. 1988. The nutri-
tive value of hulless barley for swine. Anim. Feed           Buckwheat
Sci. Tech. 19:191-196.                                         At the present time, buckwheat (Fagopyrum
                                                             sagittatum) is most commonly grown as a grain for
Beet Pulp                                                    human consumption with small amounts used in
   Beet pulp (Beta Vulgaris) is becoming a less              pancake mixes, breakfast cereals and in certain
common ingredient in swine diets in Canada due to            breads and ethnic dishes. Buckwheat bas not been
changes in the sugar beet crop. Reduced availabil-           widely utilized as a livestock feed although it now
ity and consequent increases in the price have               appears that it has considerable potential for use as
forced nutritionists to seek alternative ingredients.        an ingredient in swine diets.
                                                        71
requirements of the growing pig, buckwheat protein            containing a higher level of buckwheat tended to
contains adequate levels of almost all of the essen-          have leaner carcasses in comparison with pigs fed
tial amino acids. Only isoleucine and methionine              barley. Therefore, when prices dictate, buckwheat
are not present at a level greatly in excess of re-           can replace barley in rations fed to growing pigs.
quirement. Unfortunately, buckwheat supplies a
relatively low level of digestible energy (3000 kcal/
kg). The prime factors accounting for its low                 Table 4-14. Performance of Growing Pigs (20-60 kg)
energy content are a high crude fibre content                 Fed Diets Containing Various Levels of Buckwheat in
( 12.0%) and a low level of fat (2. l %). This low            Combination With Barley.
digestible energy content is the prime factor limiting
the usefulness of buckwheat in swine diets.                                       Percent of Cereal as Buckwheat
                                                                                 0       25     50     75     100
   Another factor limiting the nutritional value of
buckwheat is the presence of a photosensitizing               Daily Gain (kg)   0.70     0.70 0.72 0.66       0.70
agent known as fagopyrin. Pigs fed high levels of             Daily Feed (kg)   1.95     2.12 2.27 1.94       2.01
buckwheat develop peculiar eruptions and intense              Feed/Gain         2.79     3.02 3.16 2.93       2.91
itching of the skin when exposed to sunlight. This            Backfat (mm)      30.90   31.10 29.00 27.30     27.50
condition is known as fagopyrism or buckwheat
poisoning. Only white or light-coloured areas of              Anderson and Bowland, 1984. Can. J. Anim. Sci. 64:
the skin are affected and only if they are exposed to         985-995.
direct sun I ight. If animals are kept indoors, away
from sunlight, they remain normaJ. Therefore,                   There is very little research conducted on the
under modern systems of confinement, fagopyrism               feeding value of buckwheat for the breeding herd.
is unlikely to be a problem for pigs fed buckwheat.           Based on its nutrient content, it is likely that buck-
                                                              wheat could be used in gestation diets. However,
  Buckwheat also contains several other anti-                 buckwheat should not be used if the gestating sows
nutritional factors that may limit pig performance.           are housed outdoors. In addition, because of its low
There is a trypsin inhibitor in buckwheat which may           energy content, buckwheat should not be fed to
decrease the digestibility of buckwheat protein. In           sows during lactation.
addition, condensed tannins are aJso present in
buckwheat but at a level considerably lower than              Additional Reading and References
those found in sorghum or fababeans. Despite its              Anderson, D. M. and J.P. Bowland, 1984. Evalua-
high quality protein, buckwheat should not be used            tion of buckwheat (Fagopyrum esculentum) in diets
in diets fed to starter pigs. Its high crude fibre            for growing pigs. Can. J. Anim. Sci. 64:985-995.
content and low digestible energy level are likely to
limit growth and reduce the efficiency of feed                Farrell, D.J. 1978. A nutritional evaluation of
utilization when fed to pigs of this weight range.            buckwheat tFagopyrum Esculentum). Anim. Feed
Other alternatives arc available and producers                Sci. Technol. 3:95-108.
would be wise to choose a higher energy feedstuff
as the foundation for their starter diets.                    Thacker, P.A., D.M. Anderson and J.P. Bowland
                                                              1984. Buckwheat as a potential feed ingredient for
  The results of a feeding trial in which buckwheat           use in pig diets. Pig News and Information. 5:77-
was used to replace 0, 25, 50, 75 or 100% of the              81.
cereal portion of barley-based grower diets is shown
in Table 4-14. Substitution of buckwheat for barley           Thacker, P.A .• 1990. Buckwheat. Pages 61-68 In
had no significant effect on feed intake, daily gain          P.A. Thacker and R.N. Kirkwood eds, Non-tradi-
or feed efficiency. In addition, pigs fed diets               tionaJ Feed Sources for Use in Swine Production.
                                                              Buttcrworths Publishers, Stoneham, MA.
                                                         72
Canola Meal                                                  since soybean meal contains 15% to 20% more
  Canola is a crop derived from rapeseed, but                digestible energy.
developed by Canadian researchers to contain low
levels of erucic acid ( < 2% of the oil) and                   Research on lowering the fibre content of the
glueosinolates (<30 um/gin the meal). Previously,            meal could help to overcome the relatively low
these two constituents of rapeseed meal had im-              energy concentration. Some breeding programs arc
paired its use in swine diets. The improved quality          selecting for reduced hull, as another way to
of canola has elevated the crop to huge levels in            increase DE.
Canada, surpassing al I other export crops except
wheat; canola in its various forms - seed, oil and             Research at the Prairie Swine Centre has revealed
meal - is now the second largest agricultural export         that if diets are formulated on an equal nutrient
from Canada.                                                 basis, canola meal will support performance
                                                             equivalent to that of soybean meal (Table 4-15).
   There are two types of canola currently grown:
Legend and AC Excel are commonly grown varie-
                                                             Table 4-15. Performance of Female and Castrated
ties of Argentine canola (Brassica napus), while             Male Pigs Fed Diets Containing Equal Levels of
Reward and Parkland are varieties of Polish canola           Digestible Nutrients {based on either soybean meal,
(Brassica rapa). Argentine cultivars of canola are           or a combination of canola and soybean meals).
generally later maturing but higher yielding than
Polish varieties.                                                                  Canola Meal       Soybean Meal
                                                             % Canola Meal             12.0                 0.0
  Canola meal is produced from the seed, following
                                                             % Soybean Meal             8.3                16.1
a series of processes designed to maximize the oil
                                                             Initial wt., kg           24.0                23.8
yield; unlike the soybean industry, canola is still
                                                             Final wt., kg            103.8               104.4
driven by oil production as opposed to the meal.
During processing, the seed is first warmed to
                                                             Ave. daily gain, kg       0.84               0.86
prevent shattering, cleaned and then passed through
                                                             Ave. daily feed, kg       2.38               2.49
roller mills to produce a flake. The flakes are then
                                                             Feed conversion"          2.84               2.94
conditioned using heat and moisture to prepare
them for oil extraction, which occurs through a
                                                             Carcass index            107.1               107.1
series of pre-pressing, mechanical extrusion and
                                                             Lean yield, %            49.6                49.4
finally hexane extraction. The solvent is then
                                                             P2 backfat, mm            17.3                17.3
removed from the meal using high temperature and
steam which also serve to ensure that myrosinase,            • Canola meal different from soybean meal, P < 0.05
an enzyme involved in glucosinolate metabolism, is           Source: de Lange, J.F. Patience and 0. Gillis. l 993. ls
denatured and thus rendered inactive. The meal               added biotin required in barley-based diets for growing -
emerges free of solvent, with a moisture content of          finishing pigs? Prairie Swine Centre Annual Report. pp.
8 - 10% and an oil content of less than l.5%.                35 - 38.
Gums, a phospholipid by-product of oil extraction,
may be added back to the meal at this point.
                                                                            Canola Meal
  The greatest limitation to greater canola usage is                DE                           3100
not palatability, although this unfortunately remains               Crude Protein                37.7
a topic of discussion. Countless experiments have                   Lysine                       2.16
demonstrated the ability of the pig to consume                      Digestible Lysine             1.60
canola-based diets with little or no resistance -                   Digestible Threonine          1.12
provided the diets are properly balanced and pre-                   Digestible Tryptophan         0.29
sented to the pig. A relatively low energy level,                   Calcium                       0.63
similar to that of barley, is a much greater problem,               Phosphorus                    1.0 I
                                                        73
  Canola meal is complementary to other feedstuffs          Canola Seed: Full Fat
used in swine diets and in particular, represents a           Considerable interest has been shown recently in
good "fit" with peas and other pulse crops. The             the possibility of incorporating whole canola seed
relatively higher sulphur amino acid content of             into swine diets. Dry growing conditions, early
canola helps to offset the low TSAA content of              frosts and other weather conditions may result in the
peas, while the high energy in peas counters the            production of off-grade canola seed which is
lower DE value of canola meal. Because conditions           unsuitable for crushing or export. This seed can be
that favour the growing of canola are also desirable        salvaged as a feed ingredient.
for peas, the two crops represent an excellent
complementary pair from the perspective of both               Unprocessed canola seed contains approximately
swine nutrition and crop selection.                         40% oil and 20% crude protein. Therefore, it can
                                                            be regarded as a high energy, relatively high protein
   Several research trials conducted at various             supplement. The level of other nutrients are similar
institutions across Canada have shown that canola           to those found in canola meal modified only by the
meal can be used as part or all of the supplementary        dilution effect of the oil.
protein in diets fed to breeding stock. Research
trials conducted at the University of Alberta showed
no reduction in litter size, birth weight or weaning                  Canola Seed: FuU Fat
weight when canola meal was fed to sows for two                   DE                          4750
successive parities. Therefore it would appear that               Crude Protein               20.7
canola meal can be used as the sole source of                     Lysine                      1.20
supplementary protein in diets fed to breeding                    Calcium                     0.39
stock. If high energy lactation diets are desired,                Phosphorus                  0.64
extra fat will be needed to compensate for canola's
low energy content. However, care should be taken
to ensure that sows are gradually adapted to diets            The addition of fat to swine diets has been shown
containing canola meal and not replace all of the           to increase growth rates and improve feed conver-
soybean meal in the diet abruptly by canola meal.           sion efficiency for growing pigs. In addition. the
                                                            reproductive performance of sows has been shown
   In summary, canola meal is a high quality product        to improve as a result of fat supplementation.
and when properly utilized and priced competi-              Unfortunately, there are many mechanical problems
tively, can reduce feed costs. Using typical feed           associated with adding fats and oils to swine diets,
grain and protein supplement prices, canola meal is         especially on farms using mix mills. However,
competitive with soybean meal if it can be pur-             many of these problems can be overcome through
chased at about 65- 75% of the cost of soybean meal         the use of whole canola seed as a fat source. Since
(47% protein) on a unit weight basis. When avail-           canola seeds are very small, a fine screen is re-
able at this price, even conservative nutritionists         quired. The experience of some farmers indicates
should feel comfortable allowing canola meal to             that a 1/8" screen in good condition will result in a
provide 25% of the supplementary protein in starter         reasonably good grind; some whole seeds may pass
diets ( 18+ kg), 50% in grower and lactation diets          through, but they represent a small portion of the
and 100% of the supplementary protein in finishing          total.
and gestation diets.
                                                              There is little information available on the
Further Reading and References                              nutritional value of whole canola seed for starter
Hickling, D. 1993. Canola meal: Feed industry               pigs. However, it would appear that the
guide. Canola Council of Canada, Winnipeg. MB.              performance of starter pigs may be enhanced as a
26 pp.                                                      result of including whole canola seed in starter
                                                            diets. The results of one feeding trial conducted at
                                                            the University of Alberta are presented in Table
                                                       74
4-16. This data indicates that the inclusion of 15%         inclusion, the palatability of the diet appeared to
canola seed resulted in improved growth and                 decrease and as a consequence, performance
increased efficiency. However, at higher levels of          declined.
Table 4-16. Performance of Starter Pigs Fed Diets Containing Whole Canola Seed.
Adapted from Shaw and Aherne, 1987, Univ. Alberta 66th Ann. Feeders Day Report. pp. 7-9.
  The results of a feeding trial in which whole       canola meal as the protein supplement could lead to
canola seed was included at 0, 3, 6. 9, 12 or 15% of glucosinolate hydrolysis. This may result in a
the diet of growing pigs are shown in Table 4-17.     reduction in performance. Therefore, it is recom-
In general, growth rates were improved at all levels mended that whole canola seed not exceed l 0% of
of inclusion but the optimum inclusion level ap-      the total diet if canola meal is present.
peared to be about 6% of the diet. However, it may
be possible to include whole canola seed at levels of    There does not appear to be any published infor-
up to 15% of the diet without any adverse effects     mation on the effects of feeding whole canola seed
on pig performance.                                   to breeding stock. Therefore, until more informa-
                                                      tion is available regarding the effects of feeding
  It is important to note that the above experiment   whole canola seed on reproductive performance,
utilized soybean meal as its source of additional     diets fed to pregnant or lactating sows should not
dietary protein. Addition of a myrosinase source      contain whole canola seed.
such as whole canola seed to a diet containing
Table 4-17. Performance and Carcass Composition of Pigs Fed Diets Containing Whole Canola Seed.
Adapted from Castell and Falk, 1980. Can. J. Anim. Sci. 60:795-797.
                                                       75
Corn
  Corn (Zea mays) sometimes referred to as maize                                 Corn
and related products have been popular ingredients                   DE                           3550
in swine diets for many years. Corn first became a                   Crude Protein                 8.5
popular swine ingredient in Ontario when new                         Lysine                       0.26
cultivars made it agronomically more feasible about                  Dig. Lysine                  0.17
30 years ago. The high yields achieved in areas                      Dig. Threonine               0.21
capable of growing com quickly translated into                       Dig. Tryptophan              0.04
more pounds of pork per acre than barley and the                     Calcium                      0.02
switch to corn was thus inevitable. However, its                     Phosphorus                   0.25
early introduction was met with a certain degree of
scepticism, due to concerns about carcass grades.
Proper diet formulation, and in particular maintain-             The protein content in corn is low and of poor
ing proper amino acid to energy ratios, soon over-             quality. Lysine and tryptophan are the first and
came such resistance. Pig diets based on com,                  second limiting amino acids, respectively, for
soybean meal and premix have become widely                     swine. The reason for the poor quality of the
accepted and have become standards against which               protein is that zein, the main storage form of
many alternative diets are compared. This does not             protein in the kernel, is a poor source of many
imply that corn (and soybean meal) based diets                 essential amino acids.
should always be the diet of choice. In many parts
of the world, equivalent of better performance is                The amino acid content of com can be predicted
achieved in pigs fed diets that contain feedstuffs             using regression equations based on crude protein.
other than corn as compared to pigs fed corn based             These equations can be found in chapter 3.
diets. For example, starch in corn is not as well
digested by young, newly weaned piglets as the                   Varieties of com that are higher in protein and
starch in oats or barley. However, com is fed as the           more specifically in lysine have been bred. The
main feed ingredient in diets for millions of pigs             lysine content in these varieties is approximately
and will continue to be a major feed ingredient in             30% higher than in conventional corn. The original
the future.                                                    high lysine varieties, and in particular Opaque 2,
                                                               were not widely accepted because of poor agro-
  Because of its high energy content, pigs perform             nomic characteristics. such as yield and disease
well on diets based on com. Carcass quality is not             resistance. However, more recently newer varieties
impaired, as evidenced by the fact that average                have been developed that have better yielding
carcass indexes in Ontario or Quebec, where com is             characteristics. Commercial hybrids of these new
the predominant grain, are no poorer than those in             varieties will become available over the next few
regions where corn is not used. The carcass fat                years. In addition new varieties of corn with ex-
from pigs fed on corn will be different than the fat           tremely high fat contents (up to 10%) may become
of pigs fed wheat or barley; it will be slightly softer        available in the future. The DE content of these
and perhaps yellower in colour. The softer consist-            varieties may be as much as 10% higher than those
ency reflects the content and quality of fat in com.           in conventional com.
Corn contains approximately 3.5% fat which is
substantially higher than that in barley and wheat                Perhaps the greatest concern surrounding the use
that contain approximately 1.7% fat. The fat that is           of corn in swine diets is its susceptibility to molds,
present in com tends to be highly unsaturated and              especially when wet weather arrives during critical
soft ( oily) as well; the colour is due to the presence        growing stages. The actual mold does not appear to
of a pigment called cryptoxanthin. In some parts of            be the problem; the fungi may consume some
the world, this colour is considered undesirable and           nutrients in the kernel, but the effect on feed quality
white varieties of corn are preferred.                         is questionable. Corn so moldy that the kernels
                                                               could hardly be separated, has been fed to pigs with
                                                          76
no adverse affects on performance. However, when                 On a dry matter basis, there is little difference in
the mold produces a mycotoxin, such as vomitoxin               the nutritive value of high moisture or dry corn.
or searalenone, the effects on swine can be very               Phosphorus is more available from high moisture
serious. Reproductive performance of sows appears              corn. As vitamin E tends to degrade faster in high
to be most susceptible, so that some pork producers            moisture corn than in dry com, higher fortification
have switched to barley-based diets for the breeding           with vitamin E may be required in high moisture
herd. This is not necessarily a practical solution,            corn based diets. Otherwise, the decision of which
since barley can also be infected, and lactation diets         storage method to use will depend on the relative
based solely on barley as the basal grain may                  costs and convenience of the two systems. For
contain insufficient energy to support maximum                 more information on high moisture cereal grains see
milk production.                                               "high moisture barley" earlier in this chapter.
  If a suspected outbreak of mycotoxicosis occurs,               As with other cereal grains, variation in the
there is reason to believe that an increase in the             feeding value of different samples of corn, due to
nutrient (energy, protein and vitamin) content of the          differences in variety, growing, harvesting, and
diet will be beneficial. Minimize other stressors in           drying conditions can be expected. It is, however,
the environment also since the pig is less capable of          surprising how little research has been conducted to
dealing with them. A broad spectrum antibiotic at              determine variation in feeding value of different
therapeutic levels is recommended for the same                 samples of (sub-standard) corn. Based on research
reason. The best course of action is to remove the             with chickens, which are more sensitive to changes
suspect feed until a proper analysis has been carried          in feeding value than pigs, it can be concluded that
out to determine if mycotoxicosis is the cause of the          com bushel weight is a poor indicator of feeding
problem. A variety of feed additives are currently             value of individual com samples. This is supported
being evaluated as potential treatments for                    by the observations made on starter pigs (Table 4-
mycotoxicosis. A more thorough discussion on                   18). These results suggest that some samples of
mycotoxins appears in chapter 11.                              immature dried com with an extremely low bushel
                                                               weight can support levels of performance similar to
  Com normally must be artificially dried to pre-              that in pigs fed regular com. Unlike in barley and
vent deterioration during storage or stored as a wet           wheat. fibre levels in com are generally low and do
grain, either treated with a preservative or held in an        not vary much between samples. Fibre is thus also
oxygen limiting structure. Drying of com must be               a poor predictor of feeding value of corn samples.
carefully controlled to prevent loss of nutritive              The best predictor of feeding value may be the
value. Research has shown that drying com at                   starch content. In high fat com varieties, the fat
temperatures of up to 110°C to a final moisture                content should be considered as well. In regular
content of 12-15% will have no effect on pig                   com. the starch content is approximately 60%,
performance. Exceeding l 50°C drying temperature               while it may be as low as 50% in some samples of
will reduce acceptability to the pig.                          com. Based on the French and Dutch net energy
Table 4-18. Nutritional Value of Immature Corn (different bulk density) for Starter Pigs Fed Corn-Soybean
Meal Based Diets.
                Bulk density (lbs/bu)        Gain (kg/day)            Feed intake (kg/day)        Feed efficiency
Control 1 *              57.4                     .55                          1.32                     2.40
Control 2**              58.7                     .57                          1.42                     2.49
Variety A#               50.6                     .59                          l.38                     2.34
Variety B#               43.7                     .60                          1.41                     2.35
Variety C#               43.6                     .48                          1.24                     2.58
Variety D#               45.7                     .51                          1.23                     2.41
* 1991Ontario harvest; ** 1992 Indiana harvest; #immature dried com harvested between Dec 4, 1994 and Jan. 12,
1993. Adapted from Patterson et al., 1993, J. Anim. Sci. 71 (supplement I): 157 (abstract).
                                                          77
systems it could be derived that the DE content will           Corn gluten feed consists primarily of the corn
be reduced by 20 Kcal/kg per percent reduction in            hull mixed with some com germ. Due to its high
starch content. This relationship should be con-             fibre and low energy content, poor amino acid
firmed in well controlled studies. As mentioned              balance and digestibility it is more favoured in
earlier, the amino acid content of different corn            cattle rations than in swine rations. lt contains
samples may be predicted from the amino acid                 approximately 23% protein, 64% lysine and 3000
content.                                                     Kcal DE per kg. If included in swine diets the
                                                             inclusion level should be less than 5% in starter and
Additional Reading and References                            nursing sow diets and 20% in growing finishing pig
Patterson, R .. J.K. Tuitoek and L.G. Young. 1993.           diets. Com gluten feed has been successfully used
Nutritional value of immature corn of different bulk         at high inclusion levels in dry sow diets.
density for young pigs. J. Anim. Sci. 17 (supple-
ment 1 ): 157 (abstract).
                                                                         Corn Gluten Meal
Burgoon, K.G., J.A. Hansen, D.A. Knabe and A.J.                    DE                          4150
Bockholt. 1992. Nutritional value of quality                       Crude Protein               60.0
protein maize for starter and finisher swine. J.                   Lysine                       l.03
Anim. Sci. 70:811-817.                                             Calcium                      0.06
                                                                   Phosphorus                   0.38
Adams, K.L. and A.H. Jensen. 1987. High-fat
maize for pigs and sows. Anim. Feed Sci. Techn.
17:201-212.                                                     Corn gluten meal is actually two products, con-
                                                             taining either 41 or 60% crude protein. The former
Corn By-Products                                             is generally a mixture of the 60% material and corn
   While the major portion of the com crop is                gluten feed. Thus, the 41 % meal is lower in energy
destined for use in livestock feeds, some is destined        as well as protein compared to the 60% meal.
for further processing. Three industries use corn as         Compared to corn, 60% protein com gluten meal is
raw material: the wet millers who produce starch             richer in energy and crude protein. However, like
and oil for human use, the dry millers who produce           the parent material, the balance of amino acids is
grits, meal and flour, and the fermentation industry.        poor with lysine and tryptophan being most limit-
In all cases, between 25 and 35% of the com used             mg.
this way will be returned in processed form to the
livestock industry as feed ingredients.                         Corn gluten meal can be used in the diet of
                                                             market hogs or sows at levels of20-30% of the diet
    Com gluten feed and meal are by-products of the          without affecting palatability or performance,
wet milling industry. During wet milling, the raw            assuming proper adjustments for nutrient composi-
corn is soaked or steeped to soften the kernel and           tion are made. Indeed, higher levels are possible
facilitate separation of gluten, protein and germ.           with careful ration formulation. Com gluten feed
Steep water may be collected and sold as animal              will be used to a much lesser extent it at all, due to
feed. It is rich in protein and B-vitamins. The              its lower energy content.
steeped corn is then de-germinated (removes the
germ which may be sold as com germ meal); all                  Com distillers dried grains are derived from the
that is remaining at this point is the starch, gluten        fermentation industry, the most prominent of which
and hulls. The hulls are removed and the residual            is the alcohol industry. Actually two by-products
centrifuged to separate starch from gluten. The              are made available to the livestock industry: com
corn starch is destined for the human food trade             distillers dried grains with solubles and com distill-
while the gluten is used as animal feed. For every           ers dried grains. Over the last years the production
 I 00 kg of com milled, there will be 62-68 kg of            has changed from primarily distillers (dried) grain
starch, 3 kg of oil and the remainder will be corn           to distillers (dried) grain mixed with solubles. Both
gluten meal. corn gluten feed and corn germ meal.
                                                        78
of these products contain approximately 27% crude                The energy content of distillers dried grains with
protein. Although both can be used effectively by              solubles is approximately equal to that of corn.
the swine industry, the majority is destined for use           Like the parent material, the amino acid profile of
in cattle rations because the type of protein is               the protein is not well suited to the pig and thus
particularly attractive to high producing animals              needs to be used in concert with a good quality
especially dairy cattle. An additional concern with            protein supplement. Although calcium and phos-
using distillers grains, as in many by-products, is            phorus levels are higher than corn, the differences
the variability in the product. For example, in a              are of only marginal economic benefit.
recent survey of only eight samples of distillers
dried grains with solubles, the lysine content varied          Additional Reading and References
between .43 and 90% Odour and colour seem to be                Cromwell, G.L., K.L. Herkelman and T.S. Stahly.
important in determining the feeding value; light               1992. Physical, chemical and nutritional characteris-
coloured samples that are free of burnt or smoky               tics of distillers dried grains with solubles for chicks
odour, are more likely to have good nutritional                and pigs. J. Anim. Sci. 71 :679-686.
properties. Tnclusion levels of distillers dried grain
with solubles of up to 5% in starter diets, 20% in             Holden, P.J. 1991. Corn gluten feed. In (Thacker,
grower-finisher diets and 40% in dry sow diets                 P.A. and R.N. Kirkwood, Ed.): Non-traditional
appear to have little effect on animal performance             feed sources for use in swine production.
when diets are properly formulated and when good               Butterworths Publishers, 80 Montvale Avenue,
quality products are used.                                     Stoneham, MA 02180, U.S.A., pp. 131-138.
Table 4-19. Effect of Bushel Weight on Rate and Efficiency of Growth and on Digestibility of Energy in Corn.
Relative Change1
Corn Bushel Weight                         Rate of Gain                Feed Efficiency     Digestible Energy
Alberta
60   (100)                                      100                           100                  100
56   (93)                                       103                            99                  101
55   (92)                                        96                            99                  99
54   (90)                                       100                            99                  99
54   (90)                                        97                            99                  97
Manitoba
59   (100)                                      100                           100                  100
58   (98)                                        96                           IOI                  101
58   (98)                                        99                           99                    99
57   (97)                                       99                            100                    9
53   (90)                                       98                            99                   100
I
 Comparisons were carried out by arbitrarily setting the values for the highest bushel weight at I 00 and calculating
each parameter as a percentage of the highest bushel weight sample. Thus, in an Alberta sample, the 56 lb corn had a
bushel weight that was 93% of the highest sample, but birds grew 103% as fast as the birds on the heaviest corn, they
converted the feed at 99% that of birds on the heaviest corn, and had a DE value of IO I% that of the heaviest corn.
Adapted from Campbell, 1981, Proc. Western Nutr. Conf. pp 238-242.
                                                          79
Dried Bakery Product                                       but like most legume seeds, it is deficient in
   Dried bakery product, as its name suggests, is          methionine. A high level of cystine partly over-
reclaimed waste from bakeries. Because it has been         comes the methionine deficiency and therefore,
cooked during the manufacturing of the original            supplementation of diets containing fababeans with
baked goods, this product is an excellent ingredient       synthetic methionine has not proven to be benefi-
for swine, especially for young pigs whose diges-          cial. The balance of other amino acids appears fine.
tive abilities are not yet fully developed. The
cooking degrades the starches and renders them                Fababeans have a digestible energy content
more available to the pig resulting in fewer prob-         between that of soybean meal and barley. The fat
lems with digestive upset.                                 content of fababeans is low ( 1.5% ), which partially
                                                           accounts for its lower digestible energy. Fababean
  However, because dried bakery product represents         oil contains a high content of unsaturated fatty acids
waste materials, it often lacks uniformity; its con-       which can lead to the early development of rancid-
tents reflect the materials manufactured at the time       ity after the seed has been ground. Therefore,
and place of collection. This lack of uniformity has       processed fababeans should not be stored more than
compromised interest by the swine industry. Also,          a week before use. The relatively high crude fibre
since many bakery products are rich in salt, the by-       content of fababeans (8%) can also account for its
products also contain a lot of salt. Excess salt is not    lower digestible energy content.
desirable in the diet of young pigs. Many bakery
products are also sweet, so dried bakery products            Fababeans are a relatively poor source of calcium
tend to be highly palatable.                               and are low in iron and manganese. In fact, extra
                                                           supplementation of manganese is required if a diet
                                                           contains a large proportion of fababeans. The
          Dried Bakery Product                             vitamin content of the fababean is lower than that
      DE                          3975                     found in soybean meal or canola meal and there-
      Crude Protein                I 0.0                   fore, diets containing a high level of fababeans may
      Calcium                     O .15                    require a specially formulated premix in order to
      Phosphorus                  0.25                     supply a balanced ration.
                                                                           Fababeans
Fababeans
                                                                 DE                          3150
  Fababeans (Viciafaba) arc an annual belonging to
                                                                 Crude Protein               26.7
the legume family and are related to the garden
                                                                 Lysine                       1.62
broad bean. Initially most of the crop was grown
                                                                 Calcium                     0.12
for silage, but in recent years more than 75% of the
                                                                 Phosphorus                  0.49
crop has been grown for grain. The average crude
protein content of fababeans is approximately 24 -
30%. Fababean protein is relatively high in lysine
                                                          80
   Another undesirable factor is hemagglutinin                    The effects of including fababeans in breeding
which has been found to be present in fababeans at              stock diets has received little attention. However,
a level of 2900 to 4200 rabbit RBC units per gram.              Danish workers have reported a significant reduc-
The corresponding values for soybeans, wheat and                tion in litter size both at birth and weaning when
barley are 650, 50 and 5 units. Autoclaving at                  fababeans are included at high levels in gestation
l 20°C has also been shown to reduce the activity of            diets (Table 4-22). Milk protein content and yield
the hemagglutinins in fababeans.                                have also been reported to be reduced when
                                                                fababeans are included at high levels in lactation
   Fababeans contain between 0.3 and 0.5% tannin.               diets. Caution should also be exercised when
The presence of these tannins may lead to a reduc-              feeding fababeans to pregnant sows as fababeans
tion in feed intake when high levels of fababeans               generate stomach gases and may cause constipation.
are fed to swine. In addition, the digestibility of             For these reasons, fababeans should not be used at
protein and energy may be reduced as a result of the            levels greater than 15% in diets for breeding stock.
presence of these tannins.
                                                                Table 4-21. Performance of Grower Pigs (16-45 kg)
  As a result of the anti-nutritional factors present in        Fed Diets Containing Fababeans.
fababeans, it is recommended that fababeans not be
used to supply 100% of the supplementary protein                                         Fababean Level(%)
required to provide a balanced diet for swine.                                       0     10    15    20    25        30
Although there has not been a great deaJ of research
conducted to determine the value of including                   Daily Gain (kg) 0.68 0.66 0.65 0.65 0.61 0.58
fababeans in starter pig diets, the few reports                 Feed Intake (kg) 1.87 1.94 1.89 1.90 I.98 1.95
available suggest that fababeans should not be                  Feed/Gain        2.75 2.92 2.89 2.94 3.25 3.40
included at a level in excess of 15% of the diet
(Table 4-20). At higher inclusion levels, there                 _Aheme et al., 1977, Can. J. Anim. Sci. 57: 321-328.
appears to be problems with palatability and as a
consequence of the lower feed intake, growth rates              Table 4-22. Effect ofFababeans on Reproductive
are impaired.                                                   Performance.
                                                           81
Additional Reading and References                             The fat content of field peas ( 1.2%) is low and
Aherne, F.X., A.J. Lewis and R.T. Hardin. 1977.               crude fibre levels in the field pea (5.8%) are ap-
An evaluation of fababeans as a protein supplement            proximately the same as those found in barley.
for swine. Can. J. Anim. Sci. 57:321-328.
                                                                Field peas contain higher levels of both calcium
Jansrnan, A.J.M., Huisman, J. and van der Poel,               and phosphorus than the common cereal grains.
A.F.B., 1993. lleal and faecal digestibility in piglets       However, field peas are very deficient in vitamin E
offield beans tVicia faba L.) varying in tannin               and selenium and care should be taken to ensure
content. Anim. Feed Sci. Tech. 42: 83-96.                     that the diet is supplemented with these two nutri-
                                                              ents when field peas are included at high levels in
Nielsen, I-1.E. and P.E. Kruse. 1974. Effects of              swine diets.
dietary horse beans ( Vicia faba) on colostrum and
milk composition and milk yield in sows. Livest.
Prod. Sci. 1: 179-185.                                                         Field Peas
                                                                    DE                          3400
Thacker, P.A., 1990. Fababeans. P.A. Thacker and                    Crude Protein                23.4
R.N. Kirkwood eds. Non-traditional Feed Sources                     Lysine                       1.50
for Use in Swine Production. Butterworths Publish-                  Digestible Lysine            1.25
ers, Stoneham, MA. pp. 175-184.                                     Digestible Threonine         0.62
                                                                    Digestible Tryptophan        0.16
Van der Poe), A.F.B., Gravendeel, S., vam Kleef                     Calcium                      0.09
D.J., Jansman, A.J.M. and Kemp, B., 1992. Tan-                      Phosphorus                   O .50
nin-containing faba beans tVicia faba L.): Effects
of methods of processing on ileal digestibility of
protein and starch for growing pigs. Anim. Feed                 Peas, like other pulse crops, contain some anti-
Sci. Technol. 36: 205-214.                                    nutritional factors which will affect pig perform-
                                                              ance if present at high enough levels. The presence
Field Peas                                                    of a trypsin inhibitor is likely the most important.
  Field peas (Pis um sativum. arvense) are primarily          The trypsin inhibitor content of spring seeded
grown for human consumption and used mainly in                varieties of field peas is generally between 2.3 and
soup. Peas, or splits can be used quite effectively as        5.5 TIA (trypsin inhibiting activity) units/mg DM
a livestock feed when available at a sufficiently low         (dry matter) while winter varieties generally average
price. In swine diets, field peas are most often              between 8.9 and 15.9 TIA units/mg DM. Field peas
utilized to replace a portion of the protein supple-          are also reported to contain 80 units of
ment but may also replace part of the cereal grain in         hemagglutinating activity and between 3.8 and 7.0
the diet.                                                     mg/kg of cyanogenetic glycosides. Although these
                                                              anti-nutritional factors can be inactivated by heat,
  At 22 - 29% crude protein, field peas are interme-          they are usually not present at high enough levels to
diate in protein content between cereal grains and            reduce performance and therefore, are not generally
conventional protein sources. They are a good                 considered when formulating diets for swine.
source of lysine and appear to be adequate in all
other essential amino acids with the exception of               There has not been a great deal of research con-
methionine and tryptophan,                                    ducted to determine the value of including field
                                                              peas in starter pig diets. However, the few reports
  The digestible energy content of field peas (ap-            available suggest that field peas can be included in
proximately 3372 kcal/kg) is higher than most                 starter diets up to a level of 15% of the diet without
commonly used feedstuffs. A high concentration of             affecting performance. At higher levels of inclu-
easily digested starch is one of the major factors            sion, feed conversion starts to decline. Inadequate
accounting for this high digestible energy content.           tryptophan levels are suggested to be the cause of
                                                         82
the reduction in performance. Therefore, careful              Table 4-24. Performance of Growing-Finishing Pigs
                                                              (25-90 kg) Fed a Mixture of Peas and Canola Meal.
balancing of the diet for amino acids may permit
even higher levels of inclusion.                                                    SBM      C1      P2    l/3P 2/3 P
                                                                                                           2/3C   1/3 C
  There is much more data available on the effect of
including field peas in the diet of growing pigs.             Daily Gain (kg)        0.82 0.84 0.8 l 0.85 0.88
The results of a feeding trial conducted at the               Daily Feed (kg)        2.32 2.44 2.38 2.53 2.62
Agriculture Canada Research Station in Brandon in             Feed Efficiency        2.83 2.89 2.93 2.94 2.98
which the performance of pigs fed a control diet              P2 Fat Depth
containing 12.3% soybean meal as the protein                  (mm)                   11.7   11.9    12.2   11.7   12.5
supplement was compared with that of pigs fed
diets containing peas at 11, 22 or 33% of the diet
                                                               Canola Meal
are shown in Table 4-23. Pigs fed diets containing            2Field Peas
field peas gained as well as those fed the control            Castell and Cliplef, 1993. Can. J. Anim. Sci. 73: I 29-
diet at all levels of inclusion. No significant               139.
changes were observed in carcass quality as a result
of feeding peas. Therefore, it would appear that                There is limited information on the effect of
field peas can be used as the sole source of supple-          feeding field peas to breeding stock. In one trial,
mentary protein in the diet of growing pigs without           field peas were included at levels as high as 15% in
adversely affecting rate of gain or feed efficiency.          gestation diets and 25% in lactation diets without
                                                              affecting prolificacy or the number and weight of
                                                              weaned piglets. However, other trials have reported
Table 4-23. Performance of Pigs (27-95 kg) Fed Peas
as a Replacement for Soybean Meal.                            reductions in litter size when a level of I 0% of field
                                                              peas was included in diets fed during gestation and
                         Level of Pea Substitution(%)         lactation. Therefore, until further work is con-
                                                              ducted, it may be wise to limit field peas to no more
                          0       11      22       33         than I 0% of diets fed to breeding stock.
Daily Gain (kg)         0.88     0.89    0.83     0.86
Daily Intake (kg)       2.65     2.58    2.59     2.55          In summary, field peas have much to offer as a
Feed Efficiency         3.01     2.90    3.12     2.96        protein supplement and a considerable reduction in
Back.fat (mm)           16. l    15.5    15.7     15.2        feed costs may be achieved by including them in
                                                              swine diets. However, for best results it is recom-
Castell et al.. 1988. Can. J. Anim. Sci. 68: 577-579.         mended that field peas not be used at levels higher
                                                              than 15% for starter pigs and l 0% for breeding
  There has been a considerable amount of interest            stock. For growing pigs, it would appear that field
recently in the use of protein supplements made up            peas can be used to completely replace the soybean
of a blend of field peas and canola meal. Cano la             meal in the diet.
meal is a good source of sulphur containing amino
acids while peas arc a superior source of lysine. As          Additional Reading and References
a consequence, these two ingredients are particu-             Abrahamsson, M., Graham, 11., Dandanell, D. and
larly compatible. Experimental evidence indicates             Aman, P., 1993. Ilea! and faecal digestibility of
that the performance of pigs fed a blend of these             light or dark coloured peas (Pis11111 sativumi in
two protein sources is superior to that of pigs fed           growing pigs. Anim. Feed. Sci. Technol. 42: 15-24.
either ingredient alone (Table 4-24).
                                                              Bell. J.M. and A.G. Wilson. 1970. An evaluation
                                                              of field peas as a protein and energy source for
                                                              swine rations. Can. J. Anim. Sci. 50: 15-23.
                                                         83
Castell, A.G. 1987. Field peas: an alternative               analysis may over-estimate the true feeding
protein source for swine. Feedstuffs (Sept 7). pp.           value. Handling the product tends to be a major
16-17.                                                       frustration because it is bulky and not easily
                                                             conveyed by conventional grain handling equip-
Castell, A.G., 1990. Field peas. Pages 185-197 In            ment. Lack of uniformity is also a concern.
P.A. Thacker and R.N. Kirkwood eds. Non-tradi-
tional Feed Sources for Use in Swine Production.             Table 4-25. Comparison of the Composition of Grain
Butterworths Publishers, Stoneham, MA.                       Dust and Parent Grains.
Castell, A.G. and Cliplef, R.L., 1993. Evaluation                             Wheat     Corn            Soybean
of pea screenings and canola meal as a supplemen-            Nutrient      Grain Dust Grain Dust      Grain Dust
tary protein source in barley-based diets fed to             Moisture(%) 8.3 8.2 9.5 10.4              6.3 7.4
growing-finishing pigs. Can. J. Anim. Sci. 73: 129-          CP (%)      12.7 11.3 10.4 7.9           37.7 12.4
 139.                                                        Ash(%)       l.7 21.2 l.3 4.9             4.9 20.0
Castell, A.G., Neden, L.R. and Mount, K., 1988.              Adapted from Hubbard et al., 1982, Cereal Chem. 59:20-
Potential of field pea (Pisurn sativum) screenings as        22.
feed for market pigs. Can. J. Anim. Sci. 68: 577-
579.                                                           Contamination can also be a problem. Molds
                                                             may become associated with the dust from elevators
Fan, M.Z., Sauer, WC. and Jaikaran, S., 1994.                and other inadvertent contamination from various
Amino acid and energy digestibility in peas (Pisurn          chemicals and solvents used in the grain equipment
sativum) from white-flowered spring cultivars for            must be monitored. This may pose a health threat
growing pigs. J. Sci. Food Agric. 64: 249-256.               not only to pigs eating it, but also to persons work-
                                                             ing in the barn and thus exposed to the dust.
Grosjean, F. and F. Gate!. 1986. Peas for pigs. Pig
News and 1nformation. 7:443-448.                               Except for these very important concerns, grain
                                                             dust would appear to offer opportunities for feed
Ogle, R.B. and Hakansson. 1988. Nordic research              savings if the feed mill or farmer is willing to invest
with peas for pigs. Pig News and Information.                some time and money in developing the expertise
9:149-155.                                                   required to formulate and manufacture appropriate
                                                             diets.
Grain Dust
   Grain dust is normally considered a nuisance              Grain Screenings
if not an outright health and safety hazard in                 Grain screenings are not normally considered for
grain handling facilities. Yet, studies in the               use in swine diets. The product is highly variable
United States suggest it is potentially a very               and thus difficult to characterize sufficiently for
good ingredient, reflecting in nutrient composi-             precise diet formulation. Since screenings are often
tion, the parent material. The possible excep-               available at a very reasonable price, they should not
tions are fibre and ash which tend to be some-               be ignored. However, they should be used with
what elevated relative to the parent grain. Grain            caution.
dust contains slightly more lysine than the
parent material. For example, wheat dust con-                  There are a number of classes of screenings
tains 24% more lysine than the wheat it came                 available, generally from brokers or directly from
from. A summary of one study (Table 4-25)                    terminal elevators. Wheat screenings (No. 1 wheat
illustrates the di ffcrence between dust and                 screenings, wheat feed screenings) are the best
parent grains. One note of caution is the obser-             quality of screenings available. They are obtained
vation that considerable non-protein nitrogen                from the cleaning of wheat, and as such will often
may be present in the dust so that crude protein             contain considerable quantities of broken or small
                                                        84
wheat kernels. These are obviously quite satisfac-           unless adjustment in other ingredients is made.
tory for inclusion in swine diets. They will also            Their use would not normally be recommended in
contain weed seeds such as wild buckwheat, wild              the diets of lactating sows, weanlings or growing
oats, cow cockle, lady's thumb, mustard and others.          pigs.
Table 4-26 compares the chemical composition of
five samples of wheat screenings compared to a               Table 4-26. Chemical Composition of Five Samples of
sample of typical whole wheat.                               Feed Wheat Screenings Compared to Wheat.
                                                        85
Lentils                                                           Lentils are a very acceptable feedstuff for swine.
  Lentils (Lens culinarisy are a pulse crop suited to           Even if slightly frost damaged and infested with
cool temperate regions, exhibiting only moderate                aschocyta, incorporation into diets at 30% will
resistance to drought or high temperature. The                  cause no problems provided they are properly
major varieties grown in western Canada are Laird               formulated with respect to energy and essential
and Esten. Lentils are grown for sale to the human              amino acid content (particularly methionine). On
food market but on occasion, become available at a              the basis limited research, it appears that aschocyta-
competitive price to pork producers due to market               infestation of lentils has little if any adverse effect
conditions or as a result of frost damage, seed                 on pig performance or health.
damage or aschocyta blight. Discoloration during
storage is also a concern, since it results in reduced
grades. Tannins, concentrated in the seed coat,                 Table 4-27. Effects of Feeding Cull Lentils on Digest-
oxidize as the seed ages in storage causing the seed            ibility and Pig Performance (23-100 kg).
coats to darken. It is advised that lentils be stored at
                                                                                            Content of Lentils(%)
a moisture content no higher than 16% to prevent
                                                                                          0      10      20     30
deterioration during storage. If drying is required,
temperatures should not exceed 43°C.
                                                                Apparent Digestibility (%)
   The chemical composition of lentils is influenced            Dry Matter               79       79      78       79
by the cultivar seeded and the growing conditions               Energy                   80       80      78       79
prior to harvesting. The crude protein content of               Nitrogen                 78       78      75       76
lentils (25.7%) is intermediate to cereals and
soybeans. Like all pulse crops, lentils are a rela-             Performance (23 to 100 kg)
tively poor source of sulphur amino acids but are
rich in lysine. Care must be taken in swine rations,            Daily Gain (kg)        0.82     0.83     0.86     0.86
especially those for the young pig, to ensure that              Daily Intake (kg)      2.53     2.48     2.56     2.58
sulphur containing amino acids are not deficient.               Feed Efficiency        3.09     2.99     2.98     3.00
About 7% of lentil protein is non-protein nitrogen
and the percent nitrogen in protein is 5.72%. Raw               Bell and Keith, 1986 Can. J. Anim. Sci. 66: 529-536.
lentils contain trypsin and chymotrypsin inhibitors,
although it appears that these are of a sufficiently
low concentration to offer no difficulties when fed             Additional Reading and References
to swine at up to 30% of the total diet. The lipid              Bell, J.M. and M.O. Keith. 1986. Nutritional and
content is not high, but is rich in linoleic acid, an           monetary evaluation of damaged lentils for growing
essential fatty acid in the diet of the pig. This may           pigs and effects of antibiotic supplements. Can. J.
be particularly important in areas where coarse                 Anim. Sci. 66:529-536.
grains such as wheat and barley are fed. since they
contain much Jess Jinoleic acid than diets based on             Castell, A.G. 1990. Lentils. Pages 205-212 /n
corn.                                                           P.A. Thacker and R.N. Kirkwood eds. Non-tradi-
                                                                tional Feed Sources for Use in Swine Production.
                                                                Butterworth Pub., Stoneham, MA.
                 Lentils
      DE                           3065                         Castell, A.G. and Cliplef, R.L., 1990. Methionine
      Crude Protein                24.6                         supplementation of barley diets containing lentils
      Lysine                       1.63                         (Lens culinaris) or soybean meal: Live performance
      Calcium                      0.08                         and carcass responses by gilts fed ad libitum. Can.
      Phosphorus                   0.33                         J. Anim. Sci. 70: 329-332.
                                                           86
Molasses                                                   lysine, threonine and the sulfur containing amino
  Molasses is a by-product of the manufacturing of         acids. However, in contrast to most cereal grains,
sugar from either sugar cane or sugar beets. Thus,         selection for an increase in protein content will not
molasses is often referred to as beet molasses or          decrease protein quality. In this regard, oats are
cane molasses. Beet molasses is higher in crude            unique among cereal grains since the lysine level of
protein (7-11 % versus 3-4%) than cane molasses.           grain is not inversely related to its protein content.
Beet molasses contain slightly more (4%) energy
than cane molasses but the latter tends to contain           Oats contain approximately l 0% less digestible
more calcium (0.8-1.0% versus 0.1 %).                      energy than barley and about 20% less energy than
                                                           wheat and com. The main reason for the reduction
                                                           in energy content is the fact that oats contain almost
              Molasses - Beet                              11 % crude fibre. This fibre is not digestible by the
       DE                         2475                     pig and its presence also impairs the digestibility of
       Crude Protein               7.1                     other nutrients contained in the grain.
       Calcium                    0.10
       Phosphorus                 0.02                       Oats contain almost twice as much fat as barley
                                                           and wheat. This fat is distributed throughout the
                                                           endosperm, germ and aleurone layer of the oat
  Molasses is becoming a much more difficult               kernel with very little fat present in the pericarp
product to obtain in adequate quantities at a com-         (outer layer). Oats also contain a very active lipase
petitive price and thus is not generally used in swine     (fat splitting enzyme) which is present almost
rations. There are exceptions, however. Some-              entirely in the pericarp of the oat kernel. Fortu-
times, molasses will be used at a rate of 3-5% of the      nately, the enzyme does not normally come in
diet for sows to help prevent constipation and             contact with the fat since this could lead to the
enhance feed intake. Other ways of reducing                development of rancidity. However, when the oat
constipation are available, so using molasses will         kernel is broken or ground, the lipase in the pericarp
depend on the personal preference of the farmer and        is brought into contact with the oat fat and free fatty
his nutritionist.                                          acids are released. As a consequence, ground oats
                                                           are far more likely to spoil than are other cereal
  Typically, molasses contains about 22-25%                grains and care should be taken not to process them
moisture. Energy content is about 12-15% less than         too far in advance of when they are required.
that of oats; therefore, use of molasses in swine
diets will reduce energy and crude protein content
unless other adjustments to the formula are made.
Oats                                                                          Oats
  Oats (Avena sativa) are not widely utilized as an              DE                          2800
energy source for use in swine production. The                   Crude Protein                10.8
main reason for their lack of use is the fact that they          Lysine                      0.40
contain a very high hull content which lowers their              Digestible Lysine           0.26
digestibility and limits their nutritive value for               Digestible Threonine        0.19
swine. However, when economics dictate, they may                 Digestible Tryptophan       0.08
find a place in diets fed to pigs during the various             Calcium                     0.10
stages of the production cycle.                                  Phosphorus                  0.35
                                                          87
  Since oats are low in energy, they are not recom-             The adverse effects of including oats in diets fed
mended as a good source of carbohydrate in starter           to growing pigs are clearly demonstrated by the
diets. For example, when oats replace corn at levels         results of the experiment presented in Table 4-28b.
between O and 50% (Table 4-28a) in diets fed to              It can be seen that as the level of oats in the diet
pigs from 5 - l 8 kg, both growth rate and feed              increases, average daily gains decrease and the
conversion suffer. This reduction in performance             amount of feed required per pound of gain in-
may reflect the pigs' inability to consume sufficient        creases. Clearly, if any reasonable level of perform-
oat diet to meet their energy requirements for               ance is desired, oats should not be included in
optimal growth. The gains are therefore slower and           grower diets.
less efficient.
Table 4-28a. Effect of Substituting Oats for Com on the Performance of Weaner Pigs.
Table 4-28b. Effect of Substituting Oats for Corn on         pigs against ulcers. The protective effect of the oats
Grower Pig Performance.                                      is believed to be due to an alcohol-soluble fraction
                                                             contained in the hulls.
Level of Oats (%)      0      29      60      95
Daily Gain (kg)      0.80    0.70    0.63    0.57            Additional Reading and References
Daily Intake (kg)    2.93    2.95    2.77    2.71            Anderson, D.M., J.M. Bell and G.l. Christison.
Feed Conversion      3.67    4.20    4.44    4.77            1978. Evaluation of a high-protein cultivar of oats
                                                             (hinoats) as a feed for swine. Can. J. Anim. Sci.
Jenson et al., 1959, J. Anim. Sci. 18: 701-709.              58:87-96.
  Feeding oats may also be of value if a swine               Watts, G. and B.D. Moser. 1981. Oats for early-
operation is experiencing problems with gastric              weaned pigs. Nebraska Swine Report. pp. 5-6.
ulcers. Several experiments have suggested that
oats may have some value in protecting growing
                                                        88
Oats: Naked                                                   Table 4-29. Comparison of Naked Oats and Other
  Since the high fibre content of oats is the major           Cereal Grains as an Energy Source for Weane.- Pigs
factor detracting from its use as a livestock feed, a         (25 - 56 days).
considerable amount of effort has been extended to
                                                                                  Naked      Oat Wheat       Corn
try and reduce the crude fibre content of oats. Plant
                                                                                   Oats     Flakes
breeders in several countries have recently devel-
oped varieties of naked oats and these new cul ti vars        Daily Gain (g)       381       434      472     433
would appear to have considerable potential for use           Daily Intake (g)     575       625      651     601
in swine production. The most popular naked oat               Feed Conversion      1.51      1.44     1.38    1.39
currently available in Canada is the cultivar Tibor.
                                                              Fowler, 1985, Pig Farming Suppl. pp 45-52.
  Hulless oats are not really bulless. They do have
a hull but it is much more loosely attached than with
conventional oats and is blown away in the field                Naked oats can be used with a high degree of
during combining. Therefore, the crude fibre                  success in diets fed to growing pigs. The results of
content of hulless oats (2.7%) is much lower than             one experiment in which naked oats were used to
conventional oats (10.7%). As a consequence, the              replace 0, 30, 65 or 97% of the com and soybean
digestible energy content of hulless oats is much             meal in diets fed to pigs from 32 - 98 kg are shown
higher than that of conventional oats. In addition,           in Table 4-30. It can be seen that growth rates were
the oil content is also higher in hulless oats (7 .5%         similar at all rates of inclusion while there was a
versus 4.7%) than conventional oats. Due to the               tendency for feed efficiency to be improved as the
increase in oil, it is wise to supplement diets con-          level of naked oat in the diet increased. Dressing
taining naked oats with higher levels of vitamin E to         percentage was slightly improved in the naked oat
avoid problems with rancidity.                                diets but the carcasses tended to be fatter as the
                                                              level of naked oat in the diet increased.
  The crude protein content of hulless oats is also
higher than conventional oats. Hulless oats have a            Table 4-30. Performance of Market Hogs Fed Naked
good balance of amino acids with lysine and                   Oats (32 - 79 kg).
methionine being the only amino acids not present
in adequate amounts to meet the requirements.                                        0       30      65      97
Vitamins and minerals are generally present at a
slightly higher level than in conventional oats as a          Daily Gain (kg)       0.82 0.84 0.83 0.84
result ofremoving the diluting effect of the hull.            Daily Intake (kg)     2.61  2.59 2.40 2.36
                                                              Feed Conversion       3.18 3.09 2.89 2.81
  Despite the improvement in the nutritional com-             Dressing Percent     77.40 76.90 78.50 78.40
position of hulless oats, it would appear that they           Backfat (mm)         32.60 32.60 32.20 33.40
still cannot be utilized as the sole cereal source in         Carcass Grade          104   104  103   104
diets fed to weaner pigs. The results of an experi-
ment in which naked oats were compared with com               Morris and Burrows, 1986, Can. J. Anim. Sci. 66:
and wheat for pigs from 25 - 56 days of age are               833-836.
shown in Table 4-29. All diets were adjusted with
dried skim milk and fish meal so they provided
26% protein, 1.4% lysine and 17 MJ DE/kg. How-                  Several experiments conducted with naked oats
ever, both daily gain and feed efficiency were                and growing pigs have almost completely removed
poorer for pigs fed the diets containing hulless oats.        the supplementary protein from the diet without
The poorer performance for the weaner pigs fed                significantly affecting performance. This implies
naked oats has been attributed to the presence of a           that the amino acid profile of naked oats is adequate
high level of beta-glucan, a gummy polysaccharide             to meet the requirements of the growing pig. There-
which interferes with digestion.                              fore, the potential exists for a substantial reduction
                                                         89
in feed costs should naked oats be used at high              Oat Groats
levels in swine diets.                                         Oat groats are obtained by mechanically dehulling
                                                             oats. The removal of the highly fibrous hull from the
  Naked oats have been grown successfully in                 oat seed greatly improves the nutritive value of the
Canada. However, in some years, the hulls were               resulting product. Oat groats are considered to be
not as effectively removed by combining as hoped.            one of the most palatable and easily digested
Yields have also been disappointing at times. If the         feedstuffs available for use in swine production.
agronomic problems can be resolved though, naked
oats should have an excellent future in swine diets.           The crude protein content of oat groats is higher
                                                             than that of unhulled oats as a result of removing
Additional Reading and References                            the diluting effect of the hull. The digestible energy
Christison, G.1. and J.M. Bell. 1980. Evaluation of          content of oat groats is also 35 - 40% higher. The
terra, a new cultivar of naked oats (avena nuda)             protein quality of oat groats is considered to be
when fed to young pigs and chicks. Can. J. Anim.             fairly high with lysine and threonine being the only
Sci. 60: 465-471.                                            amino acids not present in sufficient amounts to
                                                             meet the requirements of the growing pig.
Fowler, V 1985. Naked oats, exciting new feed
for pigs. Pig Fanning supplement. November
1985. pp. 45-52.                                                           Oat Groats
                                                                   DE                          3725
Friend, D. W., A. Fortin. L.M. Poste, G. Butler, J.K.              Crude Protein                16.0
Kramer and VD. Burrows. 1988. Feeding and                          Lysine                      0.50
metabolism trials and assessment of carcass and                    Calcium                     0.10
meat quality for growing-finishing pigs fed naked                  Phosphorus                  0.40
oats. Can. J. Anim. Sci. 68:511-521.
Friend, D.W., A. Fortin, L.M. Poste, G. Butler, J.K.           The major factors determining the use of oat
Kramer and V.D. Burrows. 1989. Naked oats                    groats in swine rations are cost and availability. It
(Avena nuda) with and without lysine supplementa-            takes almost 160 kg of oats to produce 100 kg of
tion, for boars and barrows: Growth. carcass and             oat groats. When the initial purchase price of oats
meat quality, energy and nitrogen metabolism. Can.           is combined with the cost of dehulling, the selling
J. Anim. Sci. 69: 765-778.                                   price demanded by oat groat manufacturers is
                                                             higher than can be justified on the basis of their
Morris, J.R., 1990. Oats: Naked. Pages 275-284               nutrient content. Therefore, oat groats are generally
In P.A. Thacker and R.N. Kirkwood eds.                       used only in the higher priced diets such as creep
Non-traditional Feed Sources for Use in Swine                feeds and starter diets. This will depend on local
Production. Butterworth Publishers, Stoneham,                conditions and the pricing policy of suppliers.
MA.
                                                             Potatoes and Potato Products
Morris, J.R. and V.D. Burrows. 1986. Naked oats                 Potatoes (Solanum tuberosumy are grown prima-
in grower-finisher diets. Can. J. Anim. Sci.                 rily for human consumption. However, surplus
66:833-836.                                                  potatoes, potatoes unfit for human consumption,
                                                             and potato by-products (from the production of
Myer, R.O., R.D. Barnett and W.R. Walker. 1985.              potato starch, french fries or potato chips) are
Evaluation of hulless oats in diets for young swine.         available for use as swine feed ingredients.
Nutr. Rept. lnt. 32: 1273-1277.
                                                              Fresh potatoes contain only about 20-22% dry
                                                             matter. Because potatoes are almost all starch, they
                                                        90
are very high in energy but very low in crude                  this is believed to be partly responsible for the poor
protein. More than half of the nitrogen present is in          digestibility and palatability of raw potatoes.
the form of non-protein nitrogen and therefore is              Cooking improves digestibility and removes factors
not protein at all. Essentially then, potatoes are a           that inhibit digestive enzymes. However, the water
source of energy.                                              used for cooking should be discarded and not fed to
                                                               pigs because it contains the water-soluble solanin.
                                                               Cooked potatoes are an excellent energy source,
                  Potatoes                                     containing more energy than corn or wheat. The
      DE                           3350                        digestibility of gross energy in the pig is 96%
       Crude Protein                7.7                        compared to 85%-88% for wheat and corn.
       Lysine                      0.40
       Calcium                     l.20                          Some researchers have developed feeding regimes
       Phosphorus                  0.85                        for swine, allowing free choice access to cooked
                                                               potatoes and Limit-feeding a 19% protein concen-
                                                               trate at the rate of about 1.14 kg per pig per day.
  A major drawback with feeding fresh potatoes is
the low dry matter content (18 to 25%); when large               Raw potatoes can only be fed to dry sows, where
quantities are fed to pigs, bulkiness will limit               6 kg of potato can replace 1 kg of barley with
nutrient intake. In addition, raw potatoes contain             additional protein supplementation. Raw potatoes
anti-nutritional factors (chymotrypsin inhibitors and          have been fed to finishing pigs, but when 25% of
solanine; the latter in green potatoes in particular)          dietary dry matter was supplied by potatoes, signifi-
that make them unpalatable to pigs. Cooking or                 cant reductions in performance were observed.
steaming potatoes effectively inactivates these anti-          Cooked potatoes are accepted by all classes of
nutritional factors and at the same time enhances the          swine. However, due to the water content and its
digestibility of starch.                                       high bulk it is not recommended for starter pigs.
                                                               Recommended maximum inclusion levels ( dry
  The crude protein content of potatoes are low.               matter basis) are 30% for growing pigs. Recom-
More than half of the nitrogen present is in the form          mended maximum inclusion levels (dry matter
of non-protein nitrogen and therefore is not protein           basis) are 30% for growing pigs., 50% for finishing
at all. However, the actual protein that is present in         pigs and 25% and 50% for nursing and dry sows
potatoes is of an extremely high quality. Purified             respectively.
potato protein is widely used in the human food
industry; its high price prohibits its use in diets for           The two main potato by-products that could be
(starter) pigs.                                                used as swine feed ingredients are potato steam peel
                                                               and potato chips. Potato steam peel is a by-product
  On a dry matter basis the feeding value of cooked            of potato processing where the peel is removed after
potatoes is similar to that of cereal grains. The              processing. It contains typically 15% dry matter,
digestible energy is 3370Kcal/kg while the crude               and 3400 Kcal/kg, 16% protein, and .05% lysine on
protein content is I 0.9%.                                     a dry matter basis. This product can be included at
                                                               up to 20% (dry matter basis) in grower-finisher pig
  Potatoes are very low in magnesium. Since                    diets. Substandard potato chips will be high in
magnesium is not usually supplemented in standard              energy (typically 30% fat as they are cooked in oil;
swine diets, special attention should be paid to this          DE content of 5250 kcal/kg) and low in protein
mineral to ensure it is not deficient (ie. Supplemen-          (typically 6.5% protein). It is a highly palatable
tation of the diet may be required).                           ingredient that can be included at levels up to 15%
                                                               in starter diets and 25% in diets for grower pigs, dry
  Potatoes can be used effectively in the diet of              and nursing sows. Because of the high fat content
pigs, although there are notable limitations. Raw              and fat quality (oily) it is recommended that the
potatoes contain solanin, especially in the sprouts;           inclusion level in the finisher diet be restricted to
                                                                10% or less.
                                                          91
Additional Reading and References                             vitamins would also appear to be similar to those
Edwards, S.A. and R.M. Livingstone. 1991. Potato              found in other cereals.
and potato by-products. In (Thacker, P.A. and R.N.
Kirkwood, Ed.): Non-traditional feed sources for                Rye is susceptible to contamination and infection
use in swine production. Butterworths Publishers,             from several undesirable fungi, the most important
80 Montvale Avenue, Stoneham, MA.                             of which is ergot. Ergot may contain IO or more
                                                              toxic alkaloids including ergotamine, ergotoxine
Van Lunen, T.A., D.M. Anderson, A.M. St-Laurant,              and ergonovine (see Ch. 11 ). Although ergot has
J.W.G. Nickelson and P.R. Dean. 1989. The                     been shown to reduce the performance of pigs fed
feeding value of potato steam peel for growing-               rye, it is important to realize that ergot is a fungus
finishing pigs. Can. J. Anim. Sci. 69:225-234.                contaminating rye and not a factor inherent to the
                                                              rye itself. New varieties of rye have recently been
Rye                                                           developed which are much more resistant to ergot
  From an agronomic point of view, rye (Secale                infestation. Examples of these ergot resistant
cereale) is an attractive crop for Western Canada. It         varieties are Puma, Musketeer, Cougar and Kodiak.
produces higher yields of grain than other cereals            When these varieties are fed, ergot is far less likely
particularly on poor quality, sandy soils, and assists        to be a problem than with the more traditional
in better management of the soil by minimizing soil           cultivars of rye.
erosion. Rye is traditionally sown in the fa!J mak-
ing more effective use of water during spring runoff            Recent research has indicated that rye also con-
and allowing for a more equitable distribution of a           tains high levels of soluble pentosans and these
farmer's workload due to its early harvest.                   pentosans may pose a greater problem than the
                                                              presence of ergot. Pentosans are normal constitu-
  Unfortunately, the market for rye has been limited          ents of the cell wall and are related to hemicellulose
due to several toxic factors which can reduce its             but are more soluble. They result in a highly
nutritional value. If the detrimental effects of these        viscous intestinal fluid that interferes with digestion.
anti-nutritional factors could be overcome, it would
make available to the swine industry an alternative             Another compound found in rye which was
feed resource from a previously little used product.          believed to reduce pig performance is a fat-soluble,
                                                              growth inhibiting substance called 5-n-alkyl
  The energy level of rye is intermediate to that of          resorcinol. However, the most recent work on rye
wheat or barley, while the crude protein (N x 6.25)           tends to discount the importance of alkyl resorcinol
content is similar to barley and oats but lower than          as a factor contributing to the poor performance of
wheat. It supplies a reasonable balance of amino              pigs fed rye.
acids and contains roughly the same amount of
lysine as barley or wheat. Relative to the require-             Most recommendations concerning the nutritional
ments of the growing pig (20 - 50 kg), only lysine,           value of rye are based on the older varieties of rye
threonine and the sulfur containing amino acids are           which were often contaminated with ergot. These
not present in sufficient quantity to meet require-           recommendations may no longer be valid since
ments. However, these total values must be inter-             ergot is present at much lower levels in the newer
preted with caution since the availability of the             varieties of rye. As a consequence, the potential to
amino acids in rye are approximately 5 - I 0% lower           use rye in swine diets may be greater than was
than those in barley and wheat.                               previously realized.
  Rye supplies approximately the same amount of                 Rye is relatively unpalatable. Feed intakes of pigs
calcium (0.06%) and phosphorus (0.32%) as other               fed rye-based diets are generally 10% lower than
cereals. A specialized premix should not be neces-            those of pigs fed a barley-based diet. The specific
sary if rye is used as the major energy source in a           factor causing the reduced intake has not been
swine ration. The content of other minerals and               identified although a simple increase in the dusti-
                                                         92
ness of the ration has been implicated. Since                    There is very little information concerning the
weaner pigs are often reluctant to eat solid feed, a          feeding value of ergot-free rye in diets fed to
feedstuff of questionable palatability should not be          breeding stock. Until further research is conducted,
included in a starter diet. Therefore, it is suggested        it may be wise to limit the inclusion of rye to no
that rye not be used in diets fed to starter pigs.            more than 25% of the diet during gestation and to
                                                              avoid feeding rye to sows during lactation.
  Rye would appear to have much more potential
for use in diets fed to growing pigs. The results of            The nutritive value of rye for swine can be im-
one experiment in which grower pigs were fed                  proved by processing. Since dustiness is a problem
either a barley or a rye-based diet are shown in              when rye is fed, producers are advised to utilize a
Table 4-31. Pigs fed diets in which ergot-free rye            coarser grind(< 4 - Smm screen) when processing
was utilized as the sole energy source in the diet            rye for use in swine rations. Supplementation with
gained approximately 5% slower than did pigs fed a            low levels of (0.5 to 2.0%) fat or vegetable oil may
barley based diet. However, it took significantly             also help to reduce the dustiness. In addition, the
less feed to put on a pound of gain using the rye-            nutritional value of rye for pigs has been shown to
based diet. This improvement in feed efficiency               be substantially improved by pelleting.
may provide sufficient incentive to encourage the
incorporation of rye into swine diets, particularly           Additional Reading and References
when the cost of other cereal grains is high.                 Bazylo, R.B., 1990. Rye. Pages 363-372 In P.A.
                                                              Thacker and R.N. Kirkwood, eds. Non-traditional
   The reduction in performance which occurs as a             Feed Sources for Use in Swine Production.
result offeeding ergot-free rye has been attributed           Butterworths Publishers, Stoneham, MA.
to the presence of soluble pentosans. Supplementa-
tion of diets containing rye with an enzyme capable           Thacker, P.A., G.L. Campbell and J. Grootwassink.
of breaking down these pentosans has been shown               1991. The effect of enzyme supplementation on the
to produce a modest improvement in pig perform-               nutritive value of rye-based diets for swine. Can. J.
ance. As a consequence, there is not much differ-             Anim. Sci. 71: 489-496.
ence in the performance of pigs fed enzyme-
supplemented, rye-based diets and those fed barley.           Thacker, P.A., Campbell, G.L. and J. Grootwassink,
Therefore, it would appear that when prices warrant,          1992. Effect of salinomycin and enzyme supple-
enzyme- supplemented rye can be used as 100% of               mentation on nutrient digestibility and the perform-
the cereal portion of a swine diet without a signifi-         ance of pigs fed barley- or rye-based diets. Can. J.
cant reduction in pig performance.                            Anirn. Sci. 72: 117-125.
                                                         93
matcly 70% crude protein on a dry matter basis)
and isolates (containing approximately 90% crude
protein on a dry matter basis). These products have
favourable nutritional characteristics. I lowever
given the high cost of these products, they should
only be considered for use in milk replacer or in
highly complex (pre-) starter diets.
                                                          94
Table 4-32. The Effect of Heat Treatment on the               particular when roasted full fat beans were used,
Digestible Energy Content of Soybeans.                        starter pig performance was in fact reduced when
                                                              heated full fat beans where fed. This may be the
Heat treatment      DE content (MJ/kg dry matter)             result of incomplete inactivation of ANF's, or an
 Extruding                       5020                         insufficient rupturing of the cells in the seed during
 Jet sploding                    4780                         roasting. It should be noted that, given the immatu-
 Micronizing                     4680                         rity of the digestive system, starter pigs will be
 Roasting                        4420                         more sensitive than any other class of swine to
                                                              insufficient processing of full-fat soybeans. When
Derived from Marty et al., Can. J. Anim. Sci. 73:411-         extruded beans are used to replace soybean meal in
419.                                                          starter pig diets, pig performance is generally
                                                              improved. Extruded beans can effectively replace
                                                              all of the soybean meal in starter pig diets.
   A laboratory assay that can be used to routinely
check the adequacy of heat treatment of full fat                 In growing-finishing pig diets heated full fat
soybeans in the urease test. In this test the activity        soybeans can supply all of the supplemental protein
of urease, an enzyme that breaks down urea and that            in cereal grain based diets. Feeding roasted full fat
is present in raw soybeans, is monitored. This                soybeans will result in slight improvement in
enzyme is a protein and is, just like the trypsin             growth rate and feed efficiency as compared to
inhibitors, inactivated when heat treated. The                feeding soybean meal, but not to the same extent as
urease activity, measured in change in pH (acidity)           the feeding of cereal grain, soybean meal and
in a standard solution should not exceed .20 units.           supplemental fat (Table 4-33). Apparently roasted
Unfortunately there is not yet an accepted procedure          beans are not equivalent to a mixture of soybean
that can be used to determine whether beans have              meal and soybean oil, even for growing-finishing
been over-heated. The most reliable method is still           pigs. A concern with using full fat soybeans as the
an objective evaluation of colour and odour. The              sole source of supplemental protein in finishing
heat treated product should be light in colour                diets is its effect on the content and quality of the
(devoid of dark, burned particles) and small "nutty"          fat in the carcass. In particular, in pigs with
rather than burned.                                           unimproved lean growth potential, and increase in
                                                              daily energy intake due to increases in dietary
  If full-fat beans are to be used, one must consider         energy density, may increase carcass fatness and
the extra energy supplied in the diet. Remember               thus reduce lean yield. Carcass fat is certainly
that full-fat beans contain about 12% more energy             softened when full fat soybeans are fed. This has
than high protein (dehulled) soybean meal. Failure            been associated with a paler colour of the meat and
to maintain the proper ratio of essential amino               in several instances, to a reduced acceptability by
acids, in particular lysine, to energy will result in         the consumer of pork. The results of some studies
reduced growth rates, feed efficiency and in the case         indicate that the inclusion level of full fat beans
of finishing pigs in reduced carcass lean yields.             should not exceed 10% in com-based finishing pig
                                                              diets in order to maintain pork quality. As barley
  The inclusion of heated full fat soybeans in starter        and wheat contain less fat than corn, this would be
pig diets has not resulted in consistent improve-             equivalent to approximately 20% in barley or wheat
ments in performance in comparison to soybean                 based diets.
meal based starter diets. In some studies, and in
                                                         95
Table 4-33. Performance of Growing-finishing Pigs                false perceptions about its quality. The presence of
Fed Roasted Beans as the Sole Supplemental Protein               anti-nutritional compounds covered in the previous
Source in Com-based Diets
                                                                 section focused attention on the soybean's limita-
                                                                 tions rather than its potential. However, research
                         SBM      Roasted  SBM                   into the proper handling and utilization of soybeans
                                 Soybeans and oil*               soon overcame the early problems. As a result,
                                                                 soybean meal has been a staple in swine diets since
Daily gain (kg)           .89        .89          .92            the 1940's.
Daily intake (kg)        2.81       2.76         2.64
Feed conversion          3.16       3.10         2.86               Soybean meal gained acceptance in part because it
                                                                 complemented corn so well in the diet of the pig. It
  * The amount of oil and soybean meal added to the diet         is high in energy and is a rich source of the amino
resulted in similar total protein and fat contents to that in
                                                                 �cids such as lysine and tryptophan that are lacking
the roasted soybean meal based diet; derived from
                                                                 in co1:1. The calcium content may vary among
Cromwell et al., 1990. J. Anim. Sci. 68 (suppl. 1 ): 112.
                                                                 suppliers from as low as 0.25% to over 1.00%, due
                                                                 to the use of limestone in some cases to enhance its
  The advantage of including heat-treated full-fat               flowability. This poses no particular problem
beans in sow diets appears quite similar to advan-               provided appropriate adjustments are made to the
tages of alternative high energy sow diets ( chapter             overall formula to ensure that calcium.phosphorus
6). Due to their maturity, sows are better able to               ratios in the final diet are maintained in the accept-
utilize nutrients from full fat soybeans, even if they           able range. Also, as limestone use rises, the con-
are not heat treated properly.                                   centration of other nutrients such as amino acids
                                                                 and energy will fall.
Additional Reading and References
                                                                    Soybean meal is produced from raw soybeans by
Marty, B.J. and E.R. Chavez. Effects of heat
                                                                 removing the oil and grinding the remaining flakes.
processing on digestible energy and other nutrient
                                                                 Different processes may be involved. The most
digestibilities of full-fat soybeans fed to weaner,
                                                                 commonly used are pre-press solvent where oil is
grower and finisher pigs. Can. J. Anim. Sci.
                                                                 literally squeezed from the seed before solvents are
73:411-419.
                                                                 added to remove the remaining fat, or solvent
                                                                 extraction alone.
Herkelman, K.L. and G.L. Cromwell. 1991.
Utilization of full-fat soybeans by swine reviewed.
                                                                       Two types of soybean meal are currently avail-
Feedstuffs. 62(52): 15-17, 22.
                                                                     able: high protein or dehulJed and the lower pro-
                                                                     tein, non-dehulled products. Dehulled meal con-
De Schutter, A.C. and J.R. Morris. 1991.
                                                                     tains less hull and as a consequence, is richer in
Soybeans: full-fat. ln: Non-traditional Feed
                                                                     protein, amino acids and energy than the meal
Sources for Use in Swine Production. Butterworths
                                                                     containing the hull. Because of their widely differ-
Publishers, Stoneham, MA, U.S.A. (Thacker, P.A.
                                                                     ing nutrient composition, it is important to define
and R.N. Kirkwood, Ed.), pp 439-45 l.
                                                                     which meal one is talking about. For example, the
                                                                     protein content of the dehulled meal is about 47%
Soybean Meal                                                         while that of the standard meal is 44.0%. Energy
  Since soybean meal is now used so extensively in
                                                                     levels are approximately 5% higher and lysine
swine diets, it is difficult to imagine that its early
                                                                     about 10-15% higher in the dehulled product.
introduction was fraught with problems due not
only to legitimate anti-nutritional factors but also
                                                                96
                                                         the young piglet's digestive system. However, for
         Soybean Meal: 44%CP                             all other classes of swine a consistent and known
      DE                    3500                         quality of soybean meal can support good levels of
      Crude Protein         44.3                         animal performance if used as the sole source of
      Lysine                2.86                         supplemental protein. In fact, choosing the amount
      Digestible Lysine     2.40                         of soybean meal to use will depend mainly on price
      Digestible Threonine   l.29                        and availability.
      Digestible Tryptophan 0.46
      Calcium               0.29                         Sunflower Meal
      Phosphorus            0.61                            Sunflowers (Helianthus annuusy arc grown in
                                                         southern Manitoba, Saskatchewan and in the north-
                                                         ern Plains States of the U.S., in part for the oilseed
      Soybean meals: 48%CP                               market and, to a lesser extent, the confectionary
      DE                         3675                    market. The two are distinct in terms of the varie-
      Crude Protein              47.5                    ties of sunflowers grown. Confectionary sunflower
      Lysine                     3.18                    seeds will not be used for oil production.
      Digestible Lysine          2.70
      Digestible Threonine        1.44                      Sunflower meal has not gained the same degree of
      Digestible Tryptophan      0.50                    prominence in swine diets as other protein meals,
      Calcium                    0.29                    reflecting not so much nutritional problems, but
      Phosphorus                 0.60                    rather the availability of seed for crushing. It
                                                         appears to be readily accepted by pigs. From a
                                                         nutritional perspective, dehulled sunflower meal
                                                         represents a good product with considerable poten-
  The amino acid content of soybean meal can be          tial in swine diets.
predicted using regression equations based on crude
protein. These equations can be found in chapter 3.
                                                                      Sunflower Meal: DebuUed
  A variety of tests are available to evaluate the             DE                          3115
quality of soybean meal. Moisture, protein and                 Crude Protein               40.5
calcium analyses are recommended to ensure that                Lysine                       1.39
guaranteed minimum (protein content) and maxi-                 Digestible Lysine            1.03
mum levels (hull and limestone content) are met                Digestible Threonine        0.97
and to estimate amino acid content. The urease test            Digestible Tryptophan       0.35
may be used to ensure that the anti-nutritional
factors in the soybean meal are sufficiently inacti-
vated (see previous section on full-fat soybeans).     Three types of sunflower meal are available.
Residual hexane content may also be tested. An       DehuJled sunflower meal contains about 38% crude
objective evaluation of colour will provide a reason-protein; for swine, it is far superior to the standard
able indication of product quality as well. A darker meal which generally contains less than 30% crude
quality suggest that the protein quality is reduced  protein and is very high in crude fibre. A third
and that further laboratory tests may be required    product, partially dehulled sunflower meal, is
(amino acid analyses; protein solubility in potas-   intermediate between the two. Jt contains about
sium hydroxide).                                     32% crude protein. The fibre content varies dra-
                                                     matically between the three meal from about 14%
  Soybean meal is often used as the sole source of   in the dehulled meal to 25 - 30% in the standard
supplemental protein in swine diets. In the diet of  meal. Consequently, the energy content of the
newly weaned pigs, dried milk products such as       various sunflower meals varies widely. Even
whey or skim milk are preferred due to the nature of dehulled sunflower meal has more than twice as
                                                        97
much fibre as dchulled soybean meal, so that it                Additional Reading and References
contains slightly less digestible energy than barley.          Baird, D.M. 1982. Kinds and levels of sunflower
The standard meal contains less energy even than               meal with and without lysine supplementation
wheat bran and much less than oats. Any discus-                for finishing pigs. Proc. Georgia Nutr. Conf. pp.
sion of sunflower meal must differentiate between              74-79.
the three types.
                                                               Dinusson, WE., 1990. Sunflower meal. Pages
   The amino acid profile of dehulled sunflower                465-472 In P.A. Thacker and R.H. Kirkwood, eds.
meal is poor in comparison to soybean meal, largely            Nontraditional Feed Sources for Use in Swine
because lysine is much lower and threonine, another            Production. Butterworths Publishers, Stoneham,
important amino acid in barley and wheat based                 MA.
diets, is somewhat lower. Of additional concern is
the fact that the avai labi I ity of lysine from sun-          Jorgensen, H., W.C. Sauer and P.A. Thacker. 1984.
flower meal is about 15% less than that from                   Amino acid availabilities in soybean meal, sun-
soybean meal; threonine is also slightly less avail-           flower meal, fish meal and meat and bone meal fed
able. The replacement of soybean meal with                     to growing pigs. J. of Anim. Sci. 58:926-934.
sunflower meal in barley-based diets will require
adjustment for both available lysine and threonine
content.                                                       Sunflower Seeds
                                                                  Interest in the sunflower as a crop has increased
  Fortunately it is the dehulled meal that is pro-             tremendously in recent years. This interest is based
duced in Manitoba and it often becomes available at            on the fact that the oil contained in the sunflower
a competitive price. It is produced by the pre-press,          seed supplies a very high level of polyunsaturated
solvent process. It can be used to completely                  fatty acids. Therefore, sunflower oil is highly
replace soybean meal in swine diets, especially                sought after as a vegetable oil for human consump-
those for gestating sows and the late growing stage            tion. However, not all sunflower seeds are suitable
(above 65 kg) where energy concentration is not so             for the production of oil or for use as confectionery
critical. In early growing diets and lactation diets, a        seeds. As a consequence, whole sunflower seeds
drop in energy content will reduce productivity so             are sometimes available for use as a livestock feed.
that care is required in replacing even part of the
soybean meal with sunflower meal. In medium to                   Sunflower seeds can be considered as high in energy
high energy diets, such as starter and lactation               and medium in protein content. They contain on
formulations, added fat and an increase in total               average of about 40% oil. 20% crude protein and 29%
dietary protein or synthetic amino acids to adjust for         crude fibre. The high crude fibre level arises because
lower available lysine and threonine levels, is                of the thick hull on the sunflower seed. Unfortunately,
advised.                                                       there is little information concerning the levels of
                                                               other nutrients in the entire sunflower seed.
  Standard and partially-dehulled sunflower meal
must be used with even greater caution; because of               The results of one experiment in which sunflower
the very low energy concentration and high crude               seeds were fed to weanling pigs from four to eight
fibre content, even partial ly-dehulled sunflower              weeks of age are presented in Table 4-34. In this
meal should not be used to supply more than half of            experiment, inclusion of 6.5 or 13% sunflower
the required supplemental protein in market hog                seeds improved the average daily gain of the weaner
diets. Neither meal should be used in lactation or             pigs while the addition of 26% sunflower seeds
starter diets because the energy content is too low.           depressed performance. The reduction in growth
                                                               rate was associated with a reduced feed intake
                                                               suggesting that the high crude fibre content of the
                                                               sunflower seed limits its usefulness in starter diets.
                                                               In addition, supplementation with synthetic lysine
                                                          98
did not appear to be beneficial. A maximum of                 Sunflower seeds have been fed successfully to
I 0% sunflower seeds is therefore recommended in            sows during the last week of gestation and during
starter diets.                                              early lactation. The most dramatic effect observed
                                                            as a result of feeding sunflower seeds was a linear
Table 4-34. Effect of Feeding Sunflower Seeds on the        increase in the percentage of milk fat. However,
Performance of Weaner Pigs.                                 piglet weaning weight and percentage survival were
                                                            unaffected by dietary treatment. Some palatability
                             Sunflower Seeds(%)             problems were observed at inclusion levels greater
                        0    6.5 13.0 26.0 26.0             than 25% and therefore it would seem wise to limit
Added Lysine                                           +    the level of sunflower seed to less than 25% of the
AD G (kg)             0.61 0.63 0.63        0.55 0.56       diet.
Daily Feed (kg)       l.15 l.l6 1.18        1.03 1.04
Feed Conversion       1.88 l.84 l.87        1.87 1.86          In summary, the high crude fibre content of the
                                                            sunflower seed limits its usefulness in starter diets
Adams et al., 1982, Univ. Ill. Agric. Exper. Station        and a maximum of I 0% sunflower seed is therefore
Report. pp. 1-4.                                            recommended. As the level of sunflower seed in
                                                            the diet of the growing pig increases, carcasses
                                                            become fatter, carcass firmness decreases and bacon
  From an experiment in which sunflower seeds
                                                            sliceability becomes more difficult. Therefore, it
were fed to growing pigs (Table 4-35), it can be
                                                            would appear that market hogs should not be fed
seen that there was a linear increase in growth rate
                                                            diets containing more than l 0% sunflower seed.
as the level of sunflower seeds in the diet increased.
                                                            Finally, palatability problems have been observed at
However, as the level of sunflower seed in the diet
                                                            levels of greater than 25% in diets fed to breeding
increased, carcasses become fatter, firmness de-
                                                            stock.
creased and bacon sliceability became much more
difficult. It appears that market hogs should not be
                                                                Additional Reading and References
fed diets containing above 10% sunflower seeds
                                                                Adams, K.L. and A.H. Jensen. 1985. Effect of
because of deleterious effects on carcass quality and
                                                                dietary protein and fat levels on the utilization of
that results are improved by feeding unprocessed
                                                                the fat in sunflower seeds by the young pig. Anim.
seeds since heating reduces digestibility.
                                                                Feed Sci. Technol. 13:159-170.
Table 4-35. Performance of Growing Pigs Fed                     Adams, K.L., C.C. Lin and A.H. Jensen. 1982.
Graded Levels of Sunflower Seeds.                               Sunflower seeds in diets for young pigs. University
                                                                of Illinois Agriculture Experimental Station Report.
                               Sunflower Seeds (%)
                                                                pp. 1-4.
                              0     2.5    5     10
Average Daily Gain (kg)0.71         0.74    0.76    0.80        Hartman, A.O., R.C. Wahlstrom and G.W. Libal.
Average Daily Feed (kg) 2.36        2.46   2.53     2.60        1983. Sunflower seeds in growing-finishing swine
Feed Conversion         3.32        3.34    3.28    3.30        diets. South Dakota State 27th Annual Swine Day.
Carcass Firmness'       3.70        3.30    3.00    2.50        pp. 8-13.
Bacon Sliceability2     l. l O      l.40    1.65    2.40
IodineNumber3          55.77       58.17   64.18   71.04        Wahlstrom, R.C. 1985. Sunflowers in pig nutri-
                                                                tion. Pig News and information. 6: 151-154.
I Rangeof I to 5 with I the softest and 5 the firmest.
2
 Range of I to 5 with I being the best and 5 the worst.         Wahlstrom, R.C., 1990. Sunflower seeds. Pages
'Hartman et al., 1983, S. Dakota State 27th Ann. Swine          473-480 In P.A. Thacker and R.N. Kirkwood, eds.
Day. pp.8-13.
                                                                Nontraditional Feed Sources for Use in Swine
                                                                Production. Buuerworths Publishers, Stoneham,
                                                                MA.
                                                           99
Tri ti ca le                                                  New varieties of triticale have recently been
  Triticalc iTritlcale hexaploide) is a relatively new,     developed which are much more resistant to ergot
synthetic, small grain crop produced by crossing            infestation than were the older cultivars of triticale,
durum wheat with rye. The goal of plant breeders            Examples of ergot resistant varieties are Welsh and
in developing triticale was to combine the high             Carmen. When these varieties are fed and the ergot
crude protein and digestible energy content of wheat        concentration of the diet is kept below 0.1 %, it is
with the high yields and protein quality of rye. Its        unlikely that any ergot-related problems will de-
name is derived by combining Triticum, the botani-          velop for growing pigs fed triticale.
cal name for wheat, with Seca/e, the botanical name
for rye.                                                      Several other anti-nutritional factors have been
                                                            isolated in triticale. Perhaps the most important is
  Triticale has not been a major crop in Canada and         the presence of a high level oftypsin inhibitor
therefore, large quantities of triticale have not been      which has been shown to tie up the enzyme trypsin
fed to livestock. However, from time to time,               and reduce the digestibility of dietary protein.
significant quantities of triticale become available        There is a great deal of variation in the level of
and can be successfully utilized as an energy source        trypsin inhibitor among varieties of triticale (Table
in swine diets.                                             4-36) and therefore producers should strive to
                                                            obtain cultivars with a low trypsin inhibitor levels.
   One of the most attractive features of triticale is
its relatively high digestible energy content. For
                                                            Table 4-36. Trypsin Inhibition Levels in Spring
pigs, the digestible energy content has been esti-
                                                            Triticale1•
mated to be approximately equal to wheat and com.
A relatively low crude fibre content (2.4%) is one
                                                                Variety%                 Trypsin Inhibition
of the factors accounting for this high energy level.
                                                                Carman                      64.05 ± 7.18
  The crude protein content of triticale has been               Welsh                       79.59 ± 8.71
reported to vary considerably, depending on the                 Tri well                    72.00 ± 3.72
cultivar and growing conditions, with protein                   *75L021                     70.30 ± 0.86
contents ranging from 11.8 - 22.5% being reported.              *79P439                     65.60 ± 1.97
The average is about 16%. Triticale has a better
balance of amino acids for pigs than most other             I
                                                             Acceptable level of inhibition for hogs equals 60%
cereals, although this may be quite variable depend-        inhibition or less.
ing upon variety. Most amino acids tend to be               Adapted from Metzger, I 982. Unpublished Data.
                                                            Corvallis, Oregon.
present at a level intermediate to that of durum
wheat and rye. As is typical of most cereal grain
proteins, triticale is low in cystine, methionine,             Soluble pentosans have also been identified in
threonine and lysine. Lysine is the first limiting          triticale. It is postulated that pentosans form com-
amino acid in triticale while threonine is reported to      plex bonds with the albumen fraction of the cereal
be the second limiting amino acid. The                      protein and thus decrease the digestibility of the
availabilities of amino acids in triticale exceed those     diet. It is also suggested that pentosans increase diet
of barley by approximately 5 - I 0%.                        bulkiness as a result of their water binding capacity
                                                            in the gastrointestinal tract and thus depress feed
  Like rye, triticale tends to be susceptible to ergot      intake.
infestation (see chapter 11 ). Also, as is the case
with rye, it is important to remember that ergot is a          Other factors which have been identified in
fungus contaminating the grain and not a factor             triticale include alkyl-resorcinols and tannins.
inherent to the grain itself.                               However, the levels of these factors are generally
                                                            low and therefore, their presence is usually of little
                                                            consequence in formulating diets for swine.
                                                          100
   In evaluating triticale as an energy source for             There is very little information concerning the
swine, it is important to differentiate between              feeding value of triticalc in diets fed to breeding
problems associated with ergot and problems                  stock. Until more research is conducted to deter-
associated with the grain itself. As stated earlier,         mine the nutritive value of triticale for breeding
triticale with an ergot level above 0.1 % should not         stock, a limit of 25% of the total diet is suggested.
be fed to growing swine. Ergot-infested triticale
should not be fed to the breeding herd. Therefore,           Additional Reading and References
the remaining discussion shall relate to ergot-free          Adeola, 0., Young, L.G. and McMillan, r., 1987.
triticale.                                                   OAC Wintri tritieale in diets of growing swine.
                                                             Can. J. Anim. Sci. 67: 187-199.
   Some scientists have reported that triticale is rela-
tively unpalatable for starter pigs. Others observed no      Jilek, A.F. and R.D. Barnett. 1981. Triticale and
problems. Since starter pigs are often reluctant to eat      wheat as energy sources in swine grower and
solid feed feedstuffs of questionable palatability           finisher rations. University of Florida 26th Annual
triticale should be used with caution. Should it be          Swine Field Day Research Report. pp. 32-34.
used, then it is recommended that initially it not be
included at more than 25% of the total diet.                 King, R.H., 1980. The nutritive value of triticale
                                                             for growing pigs. Proc. Aust. Soc. Anim. Prod.
   Triticale can be successfully utilized as an energy       13:381-384.
source for growing swine. Most of the early work
with triticale indicated that it was relatively unpalat-     Lun, A.K., Smulders, J.A.H., Adeola, 0. and L.G.
able and as a consequence, most nutritionists have           Young, 1988. Digestibility and acceptability of
I imited it to about 50% of the cereal portion of the        OAC Wintri triticale by growing pigs. Can. J.
diet. However, more recent work with ergot-free              Anim. Sci. 68: 503- 510.
triticale indicates that triticale is comparable in
feeding value to both wheat and corn (Table 4-37).           Miller, E.R. and J.P. Erickson. 1980. Triticale as
Therefore. it may be possible to include triticale at        an ingredient for pig diets. Pig News and Informa-
higher levels than was previously recommended.               tion. 1 :207-210.
Producers who wish to feed it are advised to start
with about 25% triticale in the diet and gradually           Radecki, S.V and Miller, E.R., 1990. Triticale.
increase the level. The performance of the pigs              Pages 493-499 In P.A. Thacker and R.N. Kirkwood
should be monitored closely to ensure that no                eds. Nontraditional Feed Sources for Use in Swine
problems arise. Some producers who have fed                  Production. Butterworths Publishers, Stoneham,
triticale have reported problems with excessive              MA.
dust. This problem can be addressed by using a
coarser screen size when grinding or by incorporat-
ing 2 - 5% fat into the ration. Pelleting the ration         Wheat
would also be beneficial.                                      Wheat tTriticum aestiuum) is grown primarily for
                                                             the human food market and is used in livestock
Table 4-37. Performance of Grower Pigs Fed Diets             diets only when market conditions or quality dis-
Containing TriticaJe, Wheat or Corn (18-98 kg).
                                                             counts make it economical to do so. However, feed
                                                             grade wheat, destined specifically for the animal
                           Triticale   Wheat      Com
                                                             feed market, is also grown in various parts of
Average Daily Gain (kg) 0.90           0.90       0.84       Canada. From an animal feed perspective, there is
Average Feed Intake (kg) 2. 78         2.80       2.57       no difference between red or white wheat varieties,
Feed Conversion          3.09          3.11       3.06       nor between those seeded in the spring or fall.
                                                             There are slight differences between hard and soft
Jilek and Barnett, 1981, Univ. Florida 26th Ann. Swine       wheat; the former tend to have more protein, a
Field Day Res. Report. pp. 32-34.                            higher content of essential amino acids, although a
                                                           101
slightly inferior profile, and less energy. However,              Wheat contains approximately I 0% more digest-
both soft and hard wheat support equal perform-                 ible energy (DE) than barley and only slightly less
ance in growing pigs (Table 4-38) and in starter               energy than corn. However, and as discussed in
diets (Table 4-39). Studies of new varieties of feed           Chapter 3, there can be considerable variation in the
wheat, such as the Prairie Spring Wheat variety                DE content of various wheat samples. This is
Biggar or dwarf wheat have not revealed any                    especially the case when wheat is harvested prema-
serious problems, suggesting they can be used in               turely and/or under poor conditions. For example, a
swine diets according to nutrient composition                  recent study at the Prairie Swine Centre indicates
without concern. Differences in feeding value                  that differences in DE content between wheat
between different samples of wheat are more                    samples can be as high as 300 Kcal/kg or I 0% of
attributed to variation in growing and harvesting              the mean DE content. This would result in differ-
conditions that to differences between the various             ences of feed efficiency of I 0%. A means to
varieties of wheat. In general, when amino acids               predict the DE content of individual wheat samples
are properly balanced with energy, wheat makes an              is based on chemical analyses (in particular dry
excellent feed ingredient for swine feed. Dustiness            matter, fibre and protein) and using mathematical
may be a problem, especially for producers more                equations such as the following:
familiar with barley diets. Coarser grinding or the
addition of 0.5 to 2.0% fat to the diet will prove             DE (Kcal/kg of dry matter) = 3584 + 38.3 x Protein
helpful.                                                       content(% in dry matter) - 16.0 NDF content(% in
                                                                                  dry matter).
                                                         102
addition there are important differences in the             The choice of cereal grain in properly formulated
protein quality between wheat, barley and corn. In        pig diets should purely be driven by economical
spite of the differences in protein content between       considerations. In diets for starter, grower and
wheal and barley there are basically no differences       finisher pigs wheat can be used as the sole cereal
in the lysine content. Failure to recognize these         grain. Tn many practical nursing sow rations, the
differences in protein quality, by formulating diets      inclusion levels of wheat can be as high as 50%.
based on crude protein (N x 6.25) rather than amino       There is no need for high energy ingredients such as
acid levels, will result in sub-optimal pig perform-      wheat in dry sow rations. However, provided that
ance. A final point is that the amino acid to energy      attention is paid to feed processing and feeding
ratio should be considered as well. In particular,        management, the inclusion level of wheal can also
when wheat replaces barley in pig diets.                  be 50% in the dry sow diet.
  The actual amino acid content in individual wheat          When wheat is used as a pig feed ingredient close
samples is largely affected by the protein content in      attention should be paid to feed processing and in
individual wheat samples and can be predicted              particular to the fineness of grinding.
based on the mathematical equations presented in
chapter 3. These equations indicate that an increase         Wheat should not be ground too finely since it
in protein does not result in a proportional increase      becomes very floury and palatability may suffer.
in many of the essential amino acids such as lysine.       Finely ground wheat does not flow as well in self-
The changing protein content reflects largely the          feeders and the incidence of stomach ulcers may
change in the amount of gluten present; gluten is          also increase with a fine grind. For growing swine,
rich in non-essential amino acids and is a poor            most studies have demonstrated that rolled wheat
source of essential amino acids such as lysine.            will perform as well as coarsely ground wheat in the
                                                           diets of market hogs. Results of one representative
  The contents of minerals and vitamins in wheat           study revealed that rolling was better than grinding,
are not sufficiently different from those in com and       and that pelleting was the best of all treatments.
barley to warrant major changes in diet formulation.       This particular study revealed that both hard and
                                                           soft wheat responded in the same manner to
   The feeding value of wheat as compared to that in       processing treatment. It may be difficult to obtain
corn and barley. is directly the result of differences     and maintain a uniform feed mixture with rolled
in digestible energy and amino acid contents be-           wheat, so a medium grind (approximately 4.5 mm
tween these cereal grains. The results of many             to 6.4 mm screen) is recommended. If stomach
studies indicate that performance of pigs fed wheat        ulcers become a problem the addition of 5-10%
based diets is similar to that of pigs fed corn based      whole oats in pelleted rations, or coarsely ground
diets, provided that the proper dietary amino acid to      oats in mash rations generally solves the problem.
energy ratios are maintained. When comparing
wheat to barley-based diets, feed efficiency is               Off grade wheat often becomes available lo the
generally close to 10% better in the wheat-based           animal feed market and questions arise about its
diets. Differences in growth rates between pigs fed        suitability for swine. Studies conducted at the
barley and wheat based diets will depend on the            Prairie Swine Centre indicate that the DE content,
body weight of pigs. In starter pigs it is generally       of frost damaged wheat is similar to that in normal
 I 0% higher on wheat based diets and the difference       wheat. This would suggest that the actual feeding
tends to become smaller as pigs grow heavier. As           value of frost damaged wheat is similar to that of
 finishing pigs are able to compensate for changes in      regular wheat as well. The results of a France study
dietary energy density, by adjusting voluntary feed        (Table 4-40) indicate that the feeding value of wheat
intake, difference in growth rate in finishing pigs        that was partially sprouted was the same, or even
fed either barley or wheat based diets are small and       slightly better, than that of regular soft wheal.
can often not be detected.
                                                         103
Table 4-40. Performance of Starter Pigs Fed Either         they may also be called wheatings, thirds or coarse
Sprouted or Regular Soft Wheat.                            middlings.
                     Control' Sproutcd2 Sprouted'             The greatest problem associated with the use of
                                                            these milling by-products in animal feed is the lack
Daily intake (kg)       .87         .90        .87
                                                            of uniformity due to mixing of fractions. Some
Daily gain (kg)         .45        0.50        .51
                                                            bran is always included in wheat shorts, but increas-
Feed Conversion         1.94       1.81        1.71
                                                            ing the extent will decrease energy content and alter
I
  Soft wheat (I idcl)                                       the amino acid profile. Bulk density has been
2 Soft wheat (Fidel), based on visual inspection - 15%      proposed as a gauge of nutritional value, since it
sprouted                                                    would change as the proportion of various fractions
1 Hard wheat (Cando). based on visual inspection - 20%
                                                            changes.
sprouted
Adapted from Gate! and Bourdon, 1989. Pig news and
Information 10 (vol 2): 159-160.                            Additional Reading and References
                                                            Gatel, F. And 0. Bourdon. 1989. Effects of pre-
                                                            harvest sprouting on the feeding value of wheat for
  Scabby wheal, having been infected with the               pigs. Pig News and Information. 10 (vol 2): 159-
mold Fusarium graminearium, has been shown in               160.
feeding trials to reduce growth rate in swine. In
starter pigs, l ppm of vomitoxin in Fusarium-               de Lange, C.F.M., 0. Gillis, L. Whittington and J.
contaminated hard wheat was sufficient to impair            Patience. 1993. Feeding value of various wheat
growth rate in the first week on the trial. There was       samples for pigs. 1993 Annual Research report.
some compensation as the time on test progressed,           Prairie Swine Centre Inc., pp 27-30.
but even after three weeks, vomitoxin at 2.4 ppm
reduced rate and efficiency of gain. Removal of the         Magowan, WI. 1991. Wheat: soft and dwarf. In
contaminated grain from the diet did not result in          (Thacker. P.A. and R.N. Kirkwood, Ed.): Non-
compensatory gain, indicating that losses due to            traditional feed sources for use in swine production.
moldy wheat will permanently increase the number            Butterworths Publishers, 80 Montvale Avenue,
of days to market.                                          Stoneham, MA 02180. U.S.A., pp 501-508.
                                                          104
  Where constipation is a concern, especially in          problem, so that either the proportion of shorts
sows around the time of farrowing, bran at 5-15%          should be reduced or fat added to the diet to settle
of the total diet can be most helpful. Nutritionists      the dust.
must ensure that when adding fibre to the diet and
thus reducing constipation, the sow's need for
energy is not compromised. Lactation may be a             Table 4-41. Typical Specifications for Wheat and its
                                                          By-products
time when constipation is most likely, but it is also
the time when energy demands are very high.
                                                          Ingredient Crude Protein Fat                   Fibre
Consequently, bran should be matched with oil or
                                                                         Minimum(%)                  Maximum(%)
some other concentrated energy source in the diet to
ensure that the sow's energy intake is satisfactory.      Wheat2               13.6          1.7            2.7
                                                          Bran'                14.0          3.5           12.5
Wheat Shorts                                              Shorts'              15.0          4.0            8.0
  Wheat shorts consist of fine bran and parts of the      Middlings'           15.5          2.5            4.5
endosperm from the original wheat kernel. Because         Germ'                25.0          9.0            2.5
of the endosperm fraction, wheat shorts contain less
crude fibre and more energy than bran. In terms of
                                                          I
                                                              Millfeed Manual, Millers' National Federation.
                                                          2
                                                              Appendix I.
energy content, wheat shorts are approximately
equivalent to barley and contain somewhat higher
levels of crude protein, lysine and threonine. (See
Table 4-41.)                                              Wild Oat Groats
                                                            Despite intensive efforts at chemical and cultural
                                                          control, wild oats (Avenafatua) continue to con-
              Wheat Shorts                                taminate a large proportion of the grain produced in
      DE                         3140                     Western Canada. Approximately 40% of the
      Crude                       17.5                    dockage assessed on locally produced cereal grains
      Lysine                     0.70                     and as much as 1 % of the total harvest of grain is
      Digestible Lysine          0.50                     comprised of wild oats. Due to their black-col-
      Digestible Threonine       0.32                     oured hull, wild oats must be removed before the
      Digestible Tryptophan      0.18                     grain can be exported or processed for human
      Calcium                    0.10                     consumption. Over 200,000 tonnes of wild oat
      Phosphorus                 0.85                     seeds are separated annually at commercial seed
                                                          cleaning plants across Canada.
  Wheat shorts are potentially a very good ingredi-         Wild oats have traditionally been marketed as
ent for swine diets; pellet mill operators like them      Mixed Feed Oats and have been utilized almost
because they tend to improve pellet durability.           exclusively in diets fed to ruminants. The presence
Potential concerns in using wheat shorts in pig feeds     of a highly fibrous hull limits their usefulness in
include the variability of the product and the high       diets formulated for monogastrics. However, a
fibre level. Recommended maximum inclusion                technique for dehulling wild oats has been devel-
levels of wheat short are l 0% for starter pigs and       oped recently and the dehulled kernels, commonly
approximately 40% for grower-finisher pigs and            called groats, may have considerable potential to
sows. However, when diets are properly balanced           replace domestic oat groats in diets fed to poultry
for energy and amino acid contents and when a             and swine.
good consistent quality of wheat shorts is used,
animal performance can be maintained at inclusion           Wild oat groats contain significantly higher levels
levels higher than the recommended maximums. If           of crude protein in comparison with domestic oat
the diet is not pelleted, dustiness could become a        groats ( 19.9 versus 15.5%). Despite large differ-
                                                          ences in protein content between the wild and
                                                        105
Table 4-42. Performance of Starter Pigs (7 - 25 kg) Fed Various Oat Products 1.
Thacker and Sosulski, 1994. Anim. Feed Sci. Technol. 46: 229-237.
I
  Diets based on 25% oat product added at the expense of wheat.
domestic oat groats, there would appear to be little         control diet was based on wheat and soybean meal
difference in amino acid composition between                 while the remaining diets contained 25% domestic
common and wild oat protein. With the exception              oat groats, wild oat groats, toasted oat flakes or
of lysine and threonine, all of the essential amino          toasted wild oat flakes added at the expense of
acids are present in sufficient quantity to meet the         wheat. All diets were formulated to contain ap-
requirements of the starter pig (IO - 20 kg).                proximately 20% crude protein and synthetic lysine
                                                             was added so that all diets supplied approximately
   Wild oats contain a higher level of ether extract         0.9% lysine.
than domestic oat groats (7.8 versus 6.3%). Chemi-
cal analysis has shown that almost 90% of the lipid            The growth rate of pigs fed diets containing either
in wild oats is in the form of triglycerides. These          wild oat groats or wild oat flakes was similar to that
triglycerides are highly unsaturated with the oleic,         obtained with domestic oat groats and flakes.
linoleic and linolenic acid levels being 46, 35 and          Toasting and rolling of oat groats or wild oat groats
2%, respectively, of the total fatty acids. The              did not appear to improve their nutritional value.
polyunsaturated fatty acids are desirable nutrition-         The results of this experiment indicate that one may
ally, but they are also subject to oxidative instability     include up to 25% wild oat groats in starter diets
during storage especially after seed grinding.               without any adverse effects on performance. Based
                                                             on the high cost of wild oat groats, there would
  There is an active lipase in the pericarp of the           appear to be little potential for including them in
wild oat. The lipase is not in contact with the lipids       grower diets or in diets fed to breeding stock.
in the intact kernel but crushing or milling the seed
will result in hydrolysis of the triglycerides into free     Additional Reading and References
fatty acids, even at relatively low seed moisture            Sosulski, F.W. and K. Sosulski. 1985. Processing
levels. The free fatty acids are much more suscepti-         and composition of wild oat groats (Avena fatua
ble to oxidation into rancid, bitter breakdown               L.). J. Food Eng. 4:189-203.
products than the intact triglyceride. Therefore, it is
essential to steam the groats thoroughly to destroy          Sosulski, F.W.. K. Sosulski and J.P. Olson. 1985.
the lipases in the bran before proceeding to process         Nutritive value of wild oat groats and flakes. Can.
the groat into feed products. Once the enzyme has            Inst. Food Sci. Technol. 18:220-225.
been inactivated, the dry milled products can be
stored for several months without a significant              Thacker, P.A. and F.W. Sosulski. I 994. Use of
breakdown of lipids, especially if cool temperatures         wild oat groats in starter rations for swine. Anim.
are maintained.                                              Feed Sci. Technol. 46: 229-237.
  The performance of starter pigs fed diets contain-         Thacker, P.A., 1990. Wild oat groats. Pages 509-
ing wild and domestic oat groats and flakes is               515 In P.A. Thacker and R.N. Kirkwood, eds.
presented in Table 4-42. In this experiment, the             Nontraditional Feed Sources for Use in Swine
                                                             Production. Butterworths Pub., Stoneham, MA.
                                                           106
Fats and Oils                                                   single units, they are called free fatty acids. Some
  Fats and oi Is are a very concentrated source of              fats contain only two fatty acids linked together
energy, often containing 2.5-3.0 times the energy of            with the third position occupied by another com-
cereal grains. They also tend to be more expensive              pound such as a phosphorus compound.
per calorie than cereal grains and thus are used only
where higher energy levels are required., such as                 The chemical and nutritional characteristics of all
those for weanling pigs, lactating sows of fast                 fats are dictated by the individual fatty acids. Some
growing market hogs. They are also used to supply               are 'loose structures and thus are soft at room
essential fatty acids, to suppress dust, facilitate             temperature. Vegetable oils are liquid at room
pelleting and reduce wear in feed manufacturing                 temperature because they contain many of these
equipment. Jn fact, as the swine industry has                   'loose' or unsaturated fatty acids. Other fatty acids
developed, and expectations for increasingly higher             are very firm and are hard at room temperature.
standards of performance rise, fat usage in pig diets           They are called saturated fatty acids. Tallow
has increased.                                                  contains a high proportion of saturated fatty acids;
                                                                for this reason, tallow must be heated to liquefy it
   Whenever fat is added to swine diets, careful                so that it can be mixed properly into the diet. Oil is
adjustment of other nutrients, especially amino                 generally the term used to describe fats that exist as
acids, is required to ensure the diet remains bal-              a liquid at room temperature. Tallow and lard arc
anced with respect to nutrient to energy ratio's and            solid at room temperature and thus are not oils.
in particular amino acid to energy ratio's. If,                 Fatty acids are described not only as saturated or
through the addition of fat to the diet, the energy             unsaturated, but also short- or long-chained. This
content is raised without proper adjustments of the             refers to the physical length of the fatty acid mol-
dietary amino acid levels, then animal performance              ecule. The significance of these terms, from a
will be sub-optimal. In market hogs carcass index,              nutritional perspective, is explained below.
growth rates and feed efficiency will be reduced and
nursing sows will not increase their milk produc-                 In some respects, fatty acids are to fat what amino
tion. As fat is added to the diet. daily feed intake is         acids are to protein. However, there are some major
generally slightly reduced, but the pig will consume            differences. Only two fatty acids are essential in the
more energy per day, up to a maximum that will be               diet (linoleic, a member of the w-6 family of
determined by the diet and by such factors as                   polyunsaturated fatty acids and linolenic, a member
genetics and environmental temperature.                         of the w-e family of polyunsaturated fatty acids),
                                                                whereas there are ten essential amino acids. Also,
   Like many branches of nutrition, fats have a                 proteins contain only amino acids while fats contain
specific set of terminology that helps to describe              fatty acids plus other compounds, like glycerol,
their nutritive value. Most fats are present in a               sugar and phosphorus.
compound called a triglyceride (see Figure 4-2). Tt
is made up of three fat chains called fatty acids, that           The requirements for essential amino acids in pigs
are linked together at one end like tines on a fork.            are not well established. For example, there is still
If the fatty acids are not linked together but exist as         considerable debate on the efficiency with which
...
                                                          107
pigs can convert linolenic acid pigs to other polyun-        High grade vegetable fats are not commonly used
saturated fatty acids that belong to the family of w-      in swine diets due to cost. On the prairies, crude
3 fatty acids, and that are essential for normal           canola oil is sometimes used by individual farmers
animal metabolism. Some studies suggest that the           who mix their own feed because it is easy to handle
ratio of w-6 to w-3 fatty acids in the diet is as          in a simple mill. Tallow and lard as well as by-
important for the conversion of these w-3 fatty            product fats are commonly used in swine diets,
acids (see "essential fatty acids" in chapter 3).          although vegetable fats may be preferred due to
                                                           higher energy and essential fatty acid content.
  The term fat is very general, referring to a group
of ingredients including animal fats, vegetable oils          A variety of chemical tests can be carried out on
and related compounds. Feed fats can be divided            fats to determine their nutritional quality. Melting
into six general categories: animal fat, poultry fat,      point refers to the temperature at which the material
blended feed grade animal fats, blended animal and         changes from a solid to a liquid. Vegetable fats
vegetable fats, feed grade vegetable fats and              with a high proportion of 'loose' or unsaturated
soapstocks. Animal fat refers to rendered fats from        fatty acids have a low melting point, meaning they
beef or pork by-products. It includes packing house        are liquid at room temperature. Fats of animal
offal and supermarket trinunings. Because beef fat         origin such as lard and tallow, have a lower propor-
(tallow) tends to be constant regardless of the diets      tion of unsaturated fatty acids, and thus a higher
fed to cattle, it tends to be more consistent in terms     melting point. Consequently, they are solid at room
of fatty acid profile that pork fat (lard) which           temperature and must be heated before being added
depends on the diets fed to pigs.                          to the feed mix.
   Poultry grease is the rendered fat from poultry           Undesirable impurities in fats can be measured.
offal. Blended animal fats may include mixtures of         These include moisture (water, should be less than
pork lard, beef tallow, poultry grease and possibly        l %), insolubles (skin, hair, bone; should be less
restaurant grease. Blended animal and vegetable            than .5%) and unsaponifiable matter (should be less
fats may include proportions of the above men-             than l %). As undesirable impurities rises, energy
tioned animal fats plus fats of plant origin. Vegeta-      content is reduced. Free fatty acid content is not a
ble oils are derived from the refining of vegetable        good criteria of quality. Once triglycerides enter the
fats such as soybean oil, canola oil or corn oil.          gut, they are converted to free fatty acids. How-
Soapstocks, also called foots, contain products not        ever, it the fat source normally has a low free fatty
wanted in oils destined for the human food trade           acid concentration and it suddenly rises, it is a sign
including free fatty acids.                                that deterioration has occurred.
  Good quality tallow or poultry grease may con-              One inexpensive test conducted by some commer-
tain no more than 15% free fatty acids, while              cial laboratories is the iodine value analysis. It
soapstocks contain approximately 50% free fatty            determines the proportion of unsaturated and
acids. They can be used in the manufacture of soap         saturated fatty acids and thus helps to determine
or as a fat supplement in animal diets. Soapstocks         uniformity of fat supply. A change in iodine
are separated from the higher grade of oils by the         number would reflect a change in the composition
addition of an alkali which causes them to settle out,     of the fat and thus indicate a need to discuss product
allowing the still liquid oils to be removed. The          source with the supplier. Table 4-43 illustrates how
precipitate is then re-suspended by adding an acid;        changing fatty acids in the fat can alter iodine value.
for this reason, they are sometimes called acidulated      A hypothetical fat containing 50% oleic acid and
fatty acids. In concentrated form, they are quite          50% linolenic acid would have an iodine value of
corrosive, and must be used with care in the feed           1.8 while another fat, with equal portions of
manufacturing process. Once they are diluted in the        palrnitoleic, oleic and linoleic acids would have an
feed, they have proven to have excellent feeding           iodine number of 1.22.
value. ln terms of fatty acid composition, they are
quite similar to the parent vegetable oil.
                                                         108
Table 4-43. Estimating Iodine Value from Fatty Acid        fat ingredients so errors must be avoided. A
Composition of a Fat.                                      number of factors can influence the digestibility of
                                                           feed fats and thus their contribution to the energy
  Fatty Acid                         Multiply by:          content of the diet. Long chain saturated fatty acids
  C 14: I Myristoleic                    1.10              are not well accepted by the intestinal tract and are
  C 16: I Palmitoleic                    1.00              poorly digested unless mixed with unsaturated fatty
  Cl8:I Oleic                            0.90              acids which make them more compatible with the
  Cl8:2 Linoleic                         1.80              digestive processes. Thus the ratio of
  Cl 8:3 Linolenic                       2.70              unsaturated:saturated fatty acids will help to indi-
                                                           cate the digestibility of a fat source.
  Other factors that are considered include colour            The iodine number, which estimates the ratio of
and odour, A relatively recent concern surrounding         unsaturated:saturated fatty acids, is therefore very
feed grade fats is safety, or more specifically,           useful to the nutritionist. Fat digestibility appears to
freedom from compounds that make the product a             be maximized when the unsaturated:saturated ratio
health hazard. Well-publicized examples of such            is 2.0 or above. Digestibility falls rapidly below
compounds as PCB's (polychlorinated biphenyl:              2.0 and at 1.0, digestibility is reduced by about
poisonous environmental pollutants which tend to           20%. Short-chain fatty acids, whether or not they
accumulate in animal tissues) contaminating feed           are unsaturated are well accepted by the gut and are
grade fats has focused new attention on this subject.      therefore well absorbed. Fibre in the diet reduces
Quality-conscious feed companies know that there           the digestion of fat due to an increase in rate of
is no such thing as a cheap source of fat and avoid        passage. It has been estimated that for every l %
special "deals" that might come along. If farmers          crude fibre in the diet, fat digestion is reduced by
are buying fat ingredients, they would be well              1.3 to 1.5%. In Table 4-44 the digestible energy
advised to follow the same thinking.                       contents of selected fats and oils are summarized. It
                                                           should be noted that in net energy systems the
  All fat supplements, irrespective of quality or          available energy content of fats and oils, relative to
source, must contain an antioxidant to prevent             that in other ingredients is much higher than in
rancidity. Common antioxidants include propyl              digestible energy systems (chapter 3). This is
gallate, BHT (butylate hydroxytoluene), GHA                because dietary fat can be incorporated directly in
(butylate hydroxyanisole) and ethoxyquin. Failure          body fat (even in very lean pigs a substantial
to use properly stabilized fats will reduce palatabil-     amount of essential body fat needs to be deposited)
ity and could compromise the pig's status with             or in milk fat and at low heat increments of feeding.
respect to vitamin E and selenium Since rancid fat         The relative feeding value of fats is thus more
in the diet leads to damage of cell membranes. this        accurately represented in net energy than in digest-
situation must be avoided. If a poor grade of fat is       ible energy systems.
suspected, additional vitamin E and selenium
should be added to the diet. The rancidity of fats           It is difficult to suggest minimum and maximum
can be evaluated by determining the peroxide value.        inclusion levels of fat in pig feeds. With the excep-
This value should not exceed 5.0 milli-equivalent          tion of newly weaned piglets, pigs are able to utilize
per kg. Raw oil in seeds, such as canola or                added fat at inclusion levels that exceed I 0%.
soybeans, contain natural antioxidants such as             However, at inclusion levels that are higher than 6-
vitamin E. However, these may deteriorate during           8% it becomes difficult to process and handle feed
fat extraction procedures and artificial antioxidants      (bridging in bins and feeders). Given the (high)
are required.                                              cost per unit of energy. the economics of using fat
                                                           should be considered as well.
  It is important to use a correct energy value for
fats used in swine diets. In some cases, I 0-15% of
the total energy in the diet may be supplied by such
                                                         109
Table 4-44.   Digestible Energy Content of Selected Fat Sources.
Energy
  The results of studies where the effect of added             A potential concern with the inclusion of large
fat in diets for starter pigs has been evaluated have        quantities of fat in the finishing diet is the direct
been inconsistent. In many studies no or little              relationship between the quality of dietary fat and
change in growth rate or feed efficiency has been            carcass fat. As more unsaturated fat is included in
observed during the first two weeks fo!Jowing                the diet for finishing pigs, carcass fat will become
weaning. This may be attributed to the reduced               softer, more prone to oxidation (to become rancid).
digestibility and the piglets inability to metabolise        This will affect colour. shelf life and consumer
some types of fats, in particular highly saturated,          acceptance of fresh pork products. Some recom-
long chain fatty acids. Fat utilization will be im-          mendations are to maintain the total oil content
proved when medium chain fatty acids (such as                below 5% in finishing diets. In practical terms this
these in coconut oil) or when unsaturated fats (oils         means that in a com-based diet the inclusion level
are used).                                                   of full fat soybeans should not exceed 10% ( or less
                                                             than 2% added oil), and that in a barley based diet
  Fat can be included at high levels in diets for pigs       the inclusion level of full fat canola seed should not
that are older than 6 weeks. In diets for starter and        exceed 7.5% (or less than 3% added oil).
grower pigs the inclusion of fat will increase both
the energy content of the diet and the daily energy            The inclusion of fat in nursing sow diets has been
intake. The value of added fat is thus determined            proven to be beneficial in many studies. In a recent
by improvements in both feed efficiency (basically           comprehensive review of the available literature
proportional to increases in dietary energy density if       conducted by researchers at the University of
diets are properly balanced for nutrient to energy           Minnesota, fat in the saw's diet increased litter
ratios) and in growth rate (basically proportional to        weaning weights in 18 out of 24 studies, and
dietary energy density as well). As finishing pigs           lactation weight loss in the sow was reduced in 11
are better able to adjust for changes in dietary             out of 15 studies (Table 4-45). In some studies the
energy density (by adjusting daily feed intake), the         response to fat in the sows diet was small.
value of added fat in finishing diet should be based
on improvements in feed efficiency only.
                                                         110
Table 4-45. Summary of Responses to Supplemental Fat in Sow Diets.
* When treatment mean survival is less that 80% and when more than I 000 g of fat was fed to the sow prior to
farrowing; derived from Pettigrew and Moser, 1991.
  The addition of fat to the late gestation diet may           Table 4-46. Fat Content - Common Feed Ingredients.
not increase birth weight but will reduce the energy
stores in the piglet at birth. The addition of fat had             Ingredient                         Percent fat'
a positive effect on piglet survivability in 14 out of             Cereal Grains
20 experiments (Table 4-45). This was especially                   Barley                                  2
the case when birth weights were low and pre-                      Corn                                    4
weaning mortalities were high (more than 20%).                     Oats                                    5
The effects are more due to fat per see rather than to             Wheat                                   2
the increase in the daily energy intake prior to                   Fat-rich
farrowing. An additional benefit of fat in the                     Canola seed                            40
gestation diet is that the fat content of colostrum is             Soybean seen                           18
increased. In order to get these benefits, sows                    Meat and bone meal                      9
should consume approximately 1 kg of fat in the
week prior to farrowing. In a direct comparison,               I
                                                                   As defined by ether extraction.
medium-chain fatty acids in the gestation diet
appeared more effective than long-chain fatty acids
from soybean oil in reducing pre-weaning mortality.              Fat can be added to swine diets indirectly, avoid-
                                                               ing the need to install expensive fat handling equip-
  As in growing-finishing pigs, it is extremely                ment. High fat ingredients such as whole seed
important to maintain a proper balance between                 canola or soybeans have been used by many pro-
amino acid and energy intake in nursing sows. A                ducers. Table 4-46 indicates the quantity of fat
deficiency in protein during lactation maybe more              contained in such ingredients, as compared to
detrimental to sow reproductive performance than a             commonly used cereal grains. To add 2% fat to a
deficiency in energy.                                          diet, canola seed containing 40% oil should be
                                                               added to the diet at the rate of 50 kg/t (5%). The
  Due to a more efficient utilization of energy                same quantity of fat would require I 10 kg/t of
derived from digested fat as compared to that from             soybeans ( 11 %). If such high fat, and therefore
cereal grains and protein sources, and due to the              high energy feeds are used. adjustments to the
associated reduction in body heat production, the              amino acid content of the diet may be required. For
inclusion of fat in swine diets tends to reduce the            example, if 2% fat is added to the diet, crude
effect of heat stress on feed intake and animal                protein should also increase by about 0.5%.
performance.
                                                         111
  The amount of added fat required to adequately         Powles, J., J. Wiseman, D.J.A. Cole and B. Hardy.
suppress dust has not been determined. A mini-           1994. Effect of chemical structure of fats on their
mum of 0.5% has been recommended, although               apparent digestible energy value when given to
some producers are adding as much as 2% fat.             young pigs. Anim. Prod. 58:411-417.
Many commercial feed manufacturers have special-
ized fat handling equipment that allows them to add      Cera, K.R., D.C. Mahan and G.A. Reinhart. 1990.
fat to the outside of the pellet. This has proven to     Evaluation of various extracted vegetable oils,
be effective in reducing visible dust in the barn.       roasted soybeans, medium chain triglyceride and an
                                                         animal vegetable fat blend for postweaning swine.
Additional Reading and References                        J. Anim. Sci. 68:2756-2765.
Pettigrew, J.E. and R.L. Moser. 1991. Fat in swine
nutrition. In: (Miller, E.R., D.E. Ullrey and A.J.       Mineral Ingredients
Lewis, Ed.). Swine Nutrition. Butterworth -                Minerals are added to the diet when the basal
Heineman, 80 Montvale Avenue, Stoneham, MA               ingredients fail to meet the requirement of the pig.
02180, pp 133-144.                                       Thus, it is rare to supplement potassium or magne-
                                                         sium since basal ingredients supply more than
Powles, J., J. Wiseman, DJ.A. Cole and B. Hardy.         sufficient amounts of these nutrients. However,
1993. Effect of chemical structure of fats on their      calcium, phosphorus, sodium and chloride are
apparent digestible energy value when given to           routinely added to practical diets. Common mineral
growing/finishing pigs. Anim. Prod. 57:137-146.          supplements are illustrated in Table 4-47a and Table
                                                         4-47b.
Table 4-47. Typical Analysis(%) of Common Mineral Ingredients Used in Swine Diets.
Table 4-47a.
Macro minerals
ingredient                   Calcium     Phosphorus          Sodium   Chloride    Potassium     Magnesium
                                                               -percent-
Dicalcium phosphate            17.0         21.0                                                    0.6
Monosodium phosphate                        26.0              19.0
Disodium phosphate                          22.0              32.0
Mono-dicalcium phosphate       15-23        18-27
Deflourinated phosphate         31.0         18.0             5.0
Limestone                       38.0                                                                0.2
Salt                                                          39.3         60.4
Sodium Bicarbonate                                            27.4
Potassium bicarbonate                                                                39.0
Potassium chloride              0.1                           1.0          47.0      50.0           0.1
Magnesium oxide                 1-6                                                                51-59
Ammonium chloride                                                          65.0
Potassium magnesium sulfate                                                           18.0          11.0
                                                       112
Table 4-47b.
Microminerals
Ingredient                          Iron           Zinc           Copper Manganese             Iodine         Selenium
                                                                     -percent-
Calcium iodate                                                                                   62
E.D.D.I.                                                                                         80
Copper carbonate                                                   55
Copper oxide                                                       75
Copper sulphate H20                                                25
Ferrous carbonate                  36-45
Ferrous oxide                        57
Ferrous sulphate H20               20-22
Manganese carbonate-                                                           45
Manganous oxide                                                               55-65
Manganese sulphate                                                            27-28
Potassium iodide                                                                                 68
Sodium selenate                                                                                                  40
Sodium selenite                                                                                                  45
Zinc Oxide                                        70-80
Zinc sulphate                                      36
Note: Actu� nutrient composition m�y �· depending on the supplier. Nutrient composition should be confirmed by chemical
analysis. Different sources may vary ID mineral availability, especially for copper and iron. Refer to text for details.
                                                            113
pH in the intestine, thereby preventing the prolifera-       The value of adding probiotics to diets fed to
tion of harmful bacteria such as E. coli. The              growing pigs would appear to be questionable
decrease in the number of E.coli bacteria may also         based on experimental data such as that shown in
reduce the amount of toxic amines and ammonia              Table 4-49. Some researchers have speculated that
produced in the gastrointestinal tract. In addition,       probiotics may actually have some negative effects
there are reports which suggest that probiotics may        on pig performance during the growing phase by
produce an antibiotic-like substance and also              competing for nutrients with indigenous organisms
stimulate the early development of the immune              of the digestive tract, decreasing carbohydrate
system of the pig.                                         utilization and increasing the intestinal transit rate of
                                                           digesta. Therefore. although the theoretical concept
  Despite a considerable amount of study, the              of probiotics appears promising, the documented
research conducted to determine the value of               evidence of their therapeutic value suggests that the
probiotics in swine diets has been inconclusive.           search must continue for a workable alternative to
The results of one experiment conducted to deter-          antibiotics.
mine the effects of probiotics in starter diets are
shown in Table 4-48. The results of this experiment        Table 4-49. Performance of Growing Pigs (23 -38 kg)
are typical of most of the research conducted with         Fed Diets Containing a Probiotic.
starter pigs, with most researchers reporting slight
improvements in daily gain and feed efficiency as a                               Level of Probiotic (cfu/kg)
result of probiotic inclusion. However, this is not                               0      10s     ]09     I 010
always the case and several researchers have re-           Daily Gain (kg)       0.61    0.61    0.63     0.65
ported the opposite effect.                                Daily Feed (kg)       1.04    1.02    1.05     1.04
                                                           Feed Efficiency       1.69    1.66    1.66     1.60
Table 4-48. Performance of Starter Pigs Fed Diets
Containing a Problotic.                                    Scheuermann. S.E., 1993. Anim. Feed Sci. Technol. 41:
                                                           181-189.
                                Control     Pro biotic
Average Daily Gain (g)           263           270
Average Daily Feed (g)           599           594         Additional Reading and References
Feed Conversion Efficiency       2.28          2.20        Fralick, C. and T.R. Cline. 1983. The efficiency of
                                                           a commercial available probiotic. Purdue Univer-
Pollmann et al., 1980. J. Anim. Sci. 51: 577-581.          sity Swine Day. pp. 7-10.
Probiotic = Probios (MuLabs Division, Pioneer Hybrid
lntemational).                                             Pollmann, D.S. 1987. Probiotics in pig diets.
                                                           Recent Adv. Anirn. Nutr. 13: 193-205.
                                                         114
Synthetic Amino Acids                                      Some alternatives to pure synthetic amino acids
  Although the most common source of amino               are available or will become available in the near
acids in swine diets is complete proteins such as        future. Examples are methionine hydroxy analogue
soybean or canola meals. some can be provided by         (MHA) and fermentation products that contain high
individual amino acids that are manufactured by          concentrations of several essential amino acids.
bacterial culture. The most common example is L-         MHA can be used to replace methionine. It is
lysine hydrochloride (HCI). Commercial products          available in a dry form (calcium - MILA, 93%
are 98% pure and contain 78.4% lysine in a form          MHA) or in liquid form (88% MHA). There is
that is essentially 100% available to the pig. Other     some controversy about the amounts of amino acids
amino synthetic amino acids that are available, and      that are effectively supplied by these alternative
increasingly at competitive prices. include: L-          products. For example, some research with poultry
threonine (98% pure threonine in the product). DL-       would suggest that MHA is approximately 75%
methionine (99% pure methionine in the product)          effective in supplying methionine as compared to
and L-tryptophan (98% pure tryptophan in the             pure synthetic CL-methionine. Only a limited
product). There has been some data that suggest          number have been conducted to study the effective-
that synthetic amino acids, such as lysine HCI is        ness of these alternative sources of amino acids in
used less efficiently than the lysine from soybean       swine diets.
meal when pigs are fed once a day, but there is no
difference at higher feeding frequencies or when           Interestingly, as these amino acids become more
pigs are fed ad libitum.                                 economical, they provide another alternative to
                                                         balancing swine diets. Rather than only selecting
  Synthetic amino acids, and in particular lysine        among the various protein meals, the opportunity
HCI are sometimes referred to as a growth                will exist to consider the synthetic or free amino
promotant. They are not. They are ingredients            acids as another option. However, as diets change to
which supply the nutrient, lysine. and thus have no      include greater quantities of free amino acids in
growth promoting properties. Growth may im-              place of complete proteins, the need to carefully
prove with the addition of lysine HCl to the diet if     formulate and manufacture diets will increase. The
lysine is deficient. In this case, they merely im-       room for errors will shrink because the over-
prove the nutrient balance of the diet. Lysine HCI       formulation of the past will be replaced by greater
should only be purchased when it is economical. If       precision. Eliminating wasteful excesses will be of
lysine from lysine HCI is less expensive than lysine     great benefit to the industry, but it carries a signifi-
from other sources. they are worth considering. If       cant price tag in terms of quality control.
they are more expensive, the alternative sources
should be chosen.                                        Organic Acids
                                                           Organic acids are widely utilized to inhibit mold
  When large quantities of synthetic amino acids are     activity in stored feedstuffs as well as finished feed.
used it is important to consider the amount of           By treating with organic acids, it is possible to
energy supplied by amino acids as well. Just like in     harvest and store grains at a higher moisture content
protein sources, such as canola meal and soybean         without spoiling. Propionic and acetic acids are the
meal, amino acids do contribute to the digestible        most commonly used acids for this purpose.
energy content of ingredients. It can be estimated
that the DE contents of L-lysine HCL, L-threonine,         Some of the early research conducted to deter-
DL-methioninc and L-tryptophan are 4970, 4250,           mine the nutritional value of acid-treated grains
5750 and 4900 Kcal per kg of product respectively.       observed improvements in pig performance which
The energy contents of synthetic amino will have an      could not be attributed solely to the anti fungal
effect on their economic value in feed formulation       properties of these acids. Therefore, a considerable
systems and on the calculated energy content of          amount of research has been conducted to try and
diets that include synthetic amino acids.                improve rate of gain and feed efficiency through the
                                                         use of organic acids.
                                                       115
  Most of the research conducted with organic acids              between 0.5 and 3.0% of the total diet has been
has been focused on improving their performance                  shown to consistently improve feed efficiency while
of starter pigs using fumaric, citric or propionic               having little or no effect on growth rate. Typical
acid. Supplementation with organic acids at levels               research data is presented in Table 4-50.
Adapted from Giesting and Easter, 1985, J. Anim. Sci. 60: 1288-1293.
  The mechanism by which the beneficial effects of           control pigs. Therefore, if the current consumer
organic acid supplementation are achieved has not            demand for reduced carcass backfat continues,
been determined. However, it has been suggested              propionic acid supplementation of diets fed to
that the reduction in dietary pH may increase the            market hogs may increase in the future.
activity of pepsinogen, a pH-sensitive, protein
digesting enzyme in the stomach. The reduction in
dietary pH may also reduce gastric pH resulting in           Feed Flavours
greater bacteriocidal activity in the stomach, thus
                                                               The use of flavors in animal feeds has increased
reducing nutrient-robbing bacterial loads in the
                                                             considerably in the past decade as more attention is
intestinal tract. Organic acids may also act as
                                                             being paid to ingredient and diet palatability. This
chelating agents which increase the absorption of
                                                             increase in feed flavour usage has been paralleled
minerals in the intestine of the pig. Finally, it has
                                                             by a dramatic increase in the number of commer-
been suggested that the reduction in pH may slow
                                                             cially available products ranging from simple spices
gastric emptying, allowing greater time for
                                                             and tonics to aroma modifiers, sweeteners, flavour
proteolysis ( digestion of protein) to occur in the
                                                             intensifiers and artificial flavors.
stomach.
                                                                Unfortunately, feed flavors tend to be incorpo-
  As pigs age, their ability to produce their own
                                                             rated into swine feeds because of marketing appeal
gastric acid is increased and therefore, there is little
                                                             and consumer preference rather than as a result of
benefit in terms of growth rate or feed efficiency
                                                             proven effectiveness. At present, very little is
from supplementing the diets of growing pigs with
                                                             known about what specific flavors pigs find attrac-
organic acids. However, it may be possible to
                                                             tive and too often flavors are chosen for inclusion in
improve carcass traits through organic acid supple-
                                                             swine feeds because they are attractive to the human
mentation.
                                                             palate rather than that of the pig. Even if a particu-
                                                             Jar flavour has been shown to be preferred by swine
  Recent evidence has suggested that methylmalony
                                                             in free choice or stimulus tests, this preference will
CoA, a breakdown product of prop ionic acid
                                                             not necessarily result in improved performance.
metabolism, inhibits some of the enzymes involved
in fat synthesis. As a consequence, pigs fed high
                                                               The end result is that although there are claims
levels (3-9%) of propionic acid have been shown to
                                                             that such products will stimulate feed intake, very
have significantly lower levels of backfat than
                                                             few studies have demonstrated a consistent
                                                           116
improvement in feed intake or growth rate as a            Additional Reading and References
result of the inclusion of feed flavors in the diet.      McLaughlin, C.L., C.A. Baile, L.L. Buckholz and
One researcher compared 129 different feed flavors        S.K. Freeman. 1983. Preferred flavours and
to determine which specific flavors were preferred        performance of weaniling pigs. J. Anim. Sci.
by pigs. Five of the flavors which were shown to          56: 1287-1293.
be most preferred by pigs were then used in a
feeding trial. None of the flavors significantly
increased the feed intake or growth rate of starter
pigs. Since there is a cost associated with the
inclusion of flavors in the diet, it would be wise to
avoid their use until further research is conducted
and a more consistent response is obtained.
                                                        117
118
5. DIET FORMULATION
   Diet formulation is the process of matching the
pig's nutrient requirements with the nutrients                       FIVE STEPS IN SWINE DIET
supplied by available ingredients. in the most                            FORMULATION
economical manner possible. The process sounds
simple enough, but do not be misled. As explained                            Define Objectives
below, nutrient requirements vary among animals                           Establish Requirements
and farms. Over-formulation may seem to be a
logical way to ensure the animal's requirements are                 Select and Characterize Ingredients
met, but this is costly in both financial and environ-                        Formulate Diets
mental terms. Similarly, defining the nutrients                                Evaluate Diets
supplied to the pig by various ingredients stills lacks
the precision that many of us seek and again, being
too conservative in assigning specifications to
various ingredients wastes money and nutrients.             Define Objectives
finally, linking the process to net income, while              The first step in developing a feeding program is
essential, requires a firm understanding of the             to define the objectives; these will include the
relationship between nutrient supply and animal             objectives of the overall business, of the pig unit
performance. So, while the process of diet formula-         within the overall business and of individual pigs
tion sounds simple enough, it is anything but.              within the pig unit. Consequently, the nutritionist
                                                            and the pork producer must communicate to ensure
  Overall, diet formulation needs to be viewed as a         they are both working towards the same
means to an end. The 'end' is efficient production          objective(s). Following are some typical objectives.
resulting in a final pork product of high quality.          It is clear from the following list that "make the
Following five basic steps breaks ration formulation        most money," as noble (and essential) objective as it
down into simpler components that increases the             may be, is not sufficiently precise!
likelihood of success. Diet formulation, and the
development of associated feeding programs. is       Maximize Net Income per Pig Place
merely one component - albeit an important one - of    A pig place is the barn area occupied by a single
the overall pork production system.                  pig. In a grower barn, a pig place will turn over
                                                     approximately three times per year; in a nursery,
                                                     turnover will occur every 5 to 6 weeks, or about 10
                                                     times per year. By expressing net income on a 'per
                                                     pig place' basis, the importance of growth rate, and
                                                     thus of barn throughput, is recognized. Expressing
                                                     net income on a 'per pig' basis assigns much Jess
                             I.Objectives            emphasis to barn throughput. Therefore, this
                                                     objective will be most appropriate where capital
                             2 .Re�iremerrt's        costs are high and capital debt exists. lf barn
                             3.Iqgredients           throughput increases, fixed costs can be spread over
                                                     more animals.
                             4.Formu/ations
                                                              In a typical commercial circumstance, the only
                                                            instances in which one would not want to maximize
                                                            net income per pig place would be when housing
                                                            space is not limiting, when market prices do not
                                                          119
cover variable costs or when animals are not                  lower animal performance but convert a commodity
available to keep the barn full at all times. In the          that is unsaleable into one which is more easily
vast majority of circumstances, a farm's                      sold.
profitability will be maximized when farrowing
crates, nursery pens and growout facilities are kept          To Minimize the Impact on the Environment
filled to their maximum capacity.                               In Canada, as elsewhere, the impact of pork
                                                              production on the environment is under increasing
Maximize Net Income per Pig Sold                              scrutiny. For example, the spreading of slurry
  Jn some instances, such as when housing space is            places a nutrient load on the land. While this may
not limiting or when the supply offeeder pigs is              be desirable in most cases, due to fertilizing and soil
limiting, the focus of management will shift from             conditioning value, if certain conditions exist, it can
net income per pig place to net income per pig. For           lead to a portion of the nutrients entering nearby
example, feeding programs can be designed to                  lakes and rivers due to runoff. Because diet
move pigs through a barn more quickly. Such diets             composition affects slurry composition, individual
will tend to be more costly, but if there are no              pork producers may manipulate nutrient loading of
additional pigs to replace those which have moved             the soils in a way that is appropriate for their
on to market, the only opportunity to recover the             particular circumstances. Of course, following
extra feed cost is through reduced inventory costs,           good management practices for slurry handling
such as interest on the operating line of credit.             would be assumed to be the most important first
                                                              step in all cases.
To Maximize Animal Performance
  While most commercial farms will focus on                   To Produce a Carcass Possessing Certain Traits
maximizing net income, certain types of farms,                  Production systems that link animal production
notably those involved in genetic selection                   with the packing industry, through either contract or
programs, will seek to maximize animal                        ownership, increases the opportunity to develop
performance. This ensures that nutrient intake is             pork products possessing certain traits. This might
not confounding selection of animals based on                 be related to product uniformity, product quality or
genetic merit. Purebred or nucleus breeding units             possibly even more specific traits such as fatty acid
will typically fall into this category.                       profile or "organic" designation. In some of these
                                                              instances, diet plays a central roll in achieving the
  Under commercial conditions, producers may                  desired final product.
choose to feed some animals a 'non-limiting' diet to
determine the maximum potential of animals under
their particular conditions. For example. the impact
of housing, genetics and health can only be
determined when nutrient supply is not limiting
animal performance.
                                                        120
To Establish a Production System that                       Intake will be discussed in more detail under
Emphasizes Animal Well-being                                chapters dealing with the feeding of specific classes
  Animal well-being is difficult to define, let alone       of swine.
measure, but it is a subject that is attracting greater
interest both within and outside the pork industry.           Nutrient requirements can be established using
Animal well-being is not necessarily impaired by            two approaches. The more traditional system is the
maximizing net income; indeed, profitability and            empirical method which employs experiments to
welfare should not be considered as mutually                measure the pig's response to diets of differing
exclusive. Nutrition is rarely associated with              nutrient supply. This method has the advantage of
violations of animal well-being, other than cases of        employing actual animal response in the evaluation,
outright neglect leading to malnutrition and/or             but often fails to explain why the response was
starvation. However, this may change as we learn            observed. This makes it difficult to extrapolate the
more about basic animal behaviour. Scientists and           results to production circumstances which may
producers continue to work on providing the pig             differ among farms, due to genetics, environment,
with the most positive environment while still              or management.
maintaining a satisfactory profit margin.
                                                              The second approach, the factorial method,
  From this list, it is clear that the nutritionist's       attempts to identify the various functions within the
focus will vary among farms. Depending on the               animal that require a given nutrient and define the
circumstances, it may be the carcass, the pig, the          needs based on each. For example, energy is used
barn, the total agri-business of which pork produc-         for maintenance, growth, pregnancy, and lactation.
tion is only one part, or the environment. In actua1        By expressing the requirement for energy on the
practice, nutritionists will consider all of these          basis of its use for each process, and incorporating
objectives and vary the emphasis depending on the           some relationship between requirement and rate of
individual client and his or her particular situation.      production, one can estimate the pig's requirement
                                                            for energy for various levels of performance. By
                                                            adding factors that address energy needs for
Define Requirements                                         externa1 forces, such as temperature and level of
   Once the nutritional objectives have been set, the       activity, energy requirements can be predicted -
next step is to define the nutrient intake required to      theoretica1ly. The accuracy of such predictions, of
achieve them. Nutrients required by the pig depend          course, depends on the precision of the various
on many factors that are internal (i.e. genotype, age,      prediction equations.
sex, expected level of productivity) or external (i.e.
thermal and social environment, economic condi-                Typically, nutritionists use both approaches, with
tions) to the pig. Consequently, these must be              the factorial method increasing in popularity due to
considered in establishing the nutrients required in a      the need to define requirements under diverse
given diet.                                                 conditions. However, even where the factorial
                                                            method is predominant, animal experiments will
  In addition, total daily feed intake must be              still be employed to confirm the accuracy of the
considered because, in fact, nutrients are required         prediction equations.
on a daily intake basis. This applies to both the
breeding herd and market hogs. Unfortunately,                 Nutrient requirements that are published, for
there is a poor understanding offeed intake. Efforts        example, by the National Research Council (NRC)
to predict intake are limited in scope and few              or the Agriculture Research Council, are by and
operations measure it directly. Consequently, while         large those required to maximize performance.
we recognize that nutrient requirements should be           Clearly, if performance somewhat below maximum
expressed on a daily intake basis, in practice they         is the most economical, then the nutrient levels
are presented on the basis of dietary concentration.        needed may also be somewhat less than those
                                                          121
published. Given the information available, one can           constant from year to year, or even from farm to
develop at least a rough estimate of the                      farm. They need to acknowledge the specific
requirements needed for a given circumstance.                 economic circumstances and respond in an appro-
                                                              priate manner. Simply stated, there is no feeding
   The graph in Figure 5-1 illustrates the general            regime that is best under all circumstances.
approach to defining the pig's requirement for a
nutrient. It shows that as the amount of nutrient
being tested is increased in the diet, the pig
responds in some way, such as by growing faster, or
more efficiently, or by producing bigger litters.
When the diet is very deficient, small additions of
the limiting nutrient give a large improvement in
performance. However, as the requirement is
approached, the response of the pig to each unit of
nutrient becomes less. This is sometimes called the
law of diminishing returns. It is clear from this
figure that the greatest return per dollar spent does
not occur near the peak of the curve. Depending on
the marketplace, it may or may not be economical
to provide full supplementation. Again. the
factorial approach to defining nutrient requirements
is the most adaptable to such changes in expected
levels of productivity.
                                                        122
  Selecting ingredients is not a difficult task, as they     the case of energy, where digestible or
are usually the ingredients at hand. However.                metabolisable energy are always used in place of
advantages may be gained by seeking out less                 gross energy; however, for amino acids and
common ingredients that represent good value and             minerals, availability is considered less often, a
can be successfully integrated into a swine feeding          situation which we believe to be most unfortunate.
program. A review of ingredients in Chapter 4 may            Availability cannot be determined directly on
be helpful in this regard.                                   individual samples without using animal tests.
                                                             Since this is expensive and far too time consuming,
  Determining the nutrients supplied by the ingredi-         indirect methods, such as Near Infrared
ents is much more difficult, involving essentially           Spectroscopy (NTR), where available, or book
two steps: determining the nutrient composition of           values, must be employed.
the ingredient and determining the availability to the
pig of those nutrients. While a more thorough                  There are other considerations as well. First, it
discussion of this subject appears in Chapter 3, a           must be remembered that the grading system for
few of the more pertinent points will be swnma-              grain was designed to satisfy the needs of an export
rized below.                                                 market and for the milling industry, rather than to
                                                             meet the needs of those in the livestock industry.
  The best method of determining nutrient                    Thus, many of the factors that cause downgrading
composition is chemical analysis. The most                   of grains are of little or no consequence to animal
important analyses required on cereal grains and             nutritionists.
protein meals (canola meal, soybean meal) are
crude protein, calcium and phosphorus. If moisture             A case in point is bushel weight. Although very
problems are suspected, dry matter analysis should           low bushel weight may be reflected in lower energy
also be requested. While chemical assays are                 content. a wide range is perfectly acceptable to the
relatively easy and inexpensive for common                   pig. Table 3-5 outlines minimum acceptable bushel
nutrients, they are more expensive and time                  weights for swine diets. Values below the minimum
consuming for nutrients like amino acids and                 are still acceptable, but some lowering of estimated
vitamins. Therefore, in the interest of time and             energy content would be advised. Refer to chapter
economics, amino acid levels are estimated from              3 for details.
protein content (see chapter 3) or book values are
employed, if available. For the most part. these               Visual appraisal, although subjective, should not
approaches are adequate. However, if book values             be underestimated. Freedom from weed seeds,
are used, the variability of the ingredient must be          especially those that are unpalatable or toxic, is
acknowledged, as individual samples rarely reflect           obviously very important. Sprouting, mold or
the average.                                                 contamination are other things to watch for.
                                                           123
1. Linear programs                                            represent the amount of the nutrient in question
   The speed and simplicity of computer formulation           present in each of the four ingredients.
of diets makes it the most effective and popular
method for anyone involved in the process on a              Recently, feed formulation programs have be-
regular basis. Rather than use the more simplistic        come much less expensive for producers to pur-
and slower algebraic methods or Pierson's Square,         chase. These programs are much smaller than those
the computer can handle many simultaneous equa-           used by major feed companies, but they will permit
tions in a fraction of a second. Because of this          simultaneous formulation of a diet considering 50
speed and simplicity, the computer allows nutrition-      or more nutrients supplied by 50 or more ingredi-
ists to consider more ingredients and more nutrients,     ents. This is clearly more than the average pork
and pay far greater attention to cost than would          producer requires!
otherwise be the case. However. it is important to
remember that the information generated by a                There is some question whether the individual pork
computer can be no better than the information            producer really needs such capability. Is it worth the
provided to it initially!                                 money to purchase such programs? The answer lies
                                                          not in the cost of the program, but rather in the time
  Essentially, the computer takes the nutrients           and expertise required to operate it. Errors related to
needed by the pig on the one hand, and the nutrients      ingredient composition or nutrient requirements can
supplied by available feedstuffs on the other, and        lead to disastrous results. For the most part, diet
combines them to develop a balanced ration at the         formulation should be left to professionals with the
least possible cost. Computer ration formulation          training and experience required to develop a feeding
programs use a linear equation (thus, linear              program that best meets the producer's needs.
programming) such as:                                     Ongoing least cost formulation of diets to ensure that
Requirement= aX I + bX2 + cX3 + dX4                       feed costs are truly minimized can be carried out on
                                                          the farm with little risk, provided a nutritionist has
where a, b, c and d are the amounts of each of four
                                                          been involved in defining the restrictions included in
ingredients in the diet and X 1, X2. X3 and X4
                                                          the formulation program.
                                      Formula                                              Price
Ingredients                Actual     Minimum     Maximum                        Low       Actual       High
                                                        124
   Tables 5-1 and 5-2 outline the type of information     Wheat appears to be competitively priced, since it is
provided by a computer-based feed formulation             very close to its lower price limit and much cheaper
program. In Table 5-1, the ingredient summary             than its upper price. Thus, programs of this nature
itemizes the ingredients selected and the amount          can be used to determine the value of certain
required in the diet. Limits (maximums and                ingredients in various diets used on the farm.
minimums) that were set up in the original feed
specifications are also shown. It can be seen in the        Table 5-2 provides a somewhat similar summary
example that canola meal was priced competitively,        for nutrients (as opposed to ingredients). Nutrients
because it went to its upper limit. The premixes are      that are at their lower limit are forcing the cost of
obviously expensive, because they are at their lower      the diet up. For example, digestible energy, lysine,
limits. The output often also tells the formulator        sodium, calcium and phosphorus are all at their
how competitive the price for each ingredient is.         lower limit, meaning that if any of these could be
For example, soybean meal would have to drop to           lowered, the cost of the diet would be reduced.
$252 per tonne in order for more soybean meal to          However, since animal performance might suffer,
come into the formula. Likewise, if the price rose        reducing the diet cost would not necessarily save
to $336, the computer would select less soybean           money in the long run.
meal. There is no lower price limit for canola meal,
since it is already using the maximum amount.
                                                        125
2. Algebraic Formulas                                           Thus, the algebraic equation to solve the problem will be:
   ln the absence of computers, sometimes simple
calculations arc required to balance a simple ration            .0029X + .027(61.5-X) = 0.56
involving only a few ingredients. In such cases,
                                                                The equation will solve as follows:
algebraic methods arc often selected because they
                                                                .0029X + 1.6605 - 0.027X = 0.56
arc more accommodating. However, for those not
                                                                -.0241X=-l.1005
comfortable with algebra, they may be a bit confus-             X =45.7
ing at first. Following is an example:
                                                                Therefore, the amount of barley in the diet will be 45.7
EXAMPLE                                                         percent and soybean meal will make up 15.8 (i.e. 61.5 -
Problem: Formulate a diet to contain 0.78 percent               45. 7) percent. The final formula will thus look as
available lysine using five ingredients: barley,                follows:
wheat, soybean meal, canola meal and premix.
Canola meal is assumed to contain l .45 percent                        Barley               45.7
                                                                       Wheat                25.0
available (ileaJ) lysine and is fixed at IO percent of
                                                                       Soybean meal         15.8
the diet, the premix, containing no lysine, is fixed at
                                                                       Canola meal          10.0
3.5 percent of the diet and wheat, with 0.31 %                         Premix                3.5
available lysine, is set at 25 percent of the total diet.              TOTAL                100.0
The barley contains 0.29 percent available lysine
and the soybean meal 2.7 percent available lysine.              The algebraic method is adaptable to many kinds of
                                                                calculations. However, for many of us, algebra is a
So far, the diet looks like this:                               long forgotten subject; for this reason. the algebraic
                                                                method is not very popular.
  Barley                ?
  Wheat               25.0                                      3. Pierson's Square
  Soybean meal          ?                                       Pierson's square is an established method that is
  Canolameal          10.0                                      popular due to its simplicity. However, its simplic-
  Premix               3.5                                      ity is also its weakness; it functions well only in
  TOTAL              100.0                                      very simple diets. It is best used in diets containing
                                                                only two ingredients. For more complex situations,
Solution:                                                       other methods are preferable.
i) With the wheat and canola meal fixed, this diet already
                                                                EXAMPLE 1
contains 0.22 percentage (.25 * 0.3 l + .10 * 1.45 = 0.22)
points of available lysine. Thus, the barley and the
soybean meal must provide 0.56% available lysine and do
                                                                Problem: Formulate a diet containing 16% crude
so in 61.5 percent of the total mix ( JOO minus 25.0 minus      protein, 0.90% calcium and 0.75% phosphorus,
 10.0 minus 3.5 -61.5).                                         using barley and a 40% hog supplement.
ii)     Let "X" equal the proportion of barley in the diet      Solution:
and let "61.5 - X" equal the portion of soybean                 i)   The barley was analysed, and found to contain I 0%
meal in the diet. The amount of available lysine required       crude protein, 0.05% calcium and 0.29% phosphorus.
will be 0.56 percent (0.78 minus 0.22).                         The hog supplement, according to the label, contains
                                                                40% crude protein. 4.3% calcium and 2.6% phosphorus.
iii) The amount of available lysine supplied by barley will
be .0029X (Xis the amount of barley and .0029 is the            ii) Determine the proportion of barley and supplement
portion of barley that is lysine - that is 0.29%) and the       needed to provide 16% crude protein in the final mix,
amount of available lysine supplied by soybean meal will        using Pierson's Square.
be .027(61.5-X). Thal is, the amount of available lysine
in soybean meal times the amount of soybean meal in the
diet.
                                                              126
Barley                10             24            80%          Fortunately, the diet works out, such that the cal-
                        \/      16
                                                                cium and phosphorus supplied by the supplement
                                                                and the barley provides exactly the amounts re-
                                                                quired in the final mix. This may not always be the
Supplement            40
                        /\           6             20%
                                                                case. However, one cannot formulate the diet solely
                                                                on the basis of protein and select the amount of
                                     30                         supplement so dictated. The feed manufacturer
                                                                assumed a certain rate of use when the supplement
Pierson's Square is used as follows. Place the percent          was formulated. This add rate will therefore supply
protein adjacent to the ingredient and the desired level in     the amino acids, vitamins and minerals required.
the centre of the square. Subtract in the direction of the      However, if less or more of the supplement is used,
arrows (i.e. 16 - 10 = 6 and 40 - 16 = 24). Sum these           errors in other nutrient levels may result. Thus, the
two values to get 30 (i.e. 24 + 6 = 30). Divide 24 by 30        manufacturer's recommendations for inclusion
to get the percentage barley in the diet and 6 by 30 to get     should be followed within 2-3%; if this is not
the same information for the supplement.
                                                                acceptable, a special supplement may be required.
This method can, in fact, be used for any nutrient, not
just crude protein.                                             The Pierson square method can be used for some-
                                                                what more complicated formulations, as shown by
iii) Now that the proportion of barley and supplement           example 2, a diet with three ingredients: barley,
needed to meet the crude protein requirement have been          soybean meal and premix.
determined, the next step is to balance for calcium and
phosphorus. This can be accomplished by completing
Table 5-3.
                                                                EXAMPLE2
Table 5-3. Diet Composition from Pierson Square                 Problem: Formulate a diet containing 16 percent
Calculation.
                                                                protein, 0.90 percent calcium and 0.75 percent
                                                                phosphorus using three ingredients: barley, soybean
Item                       Barley    Supp.      Diet            meal and premix.
                                                              127
Barley            10             30         82.2%            three steps in the process: sampling, mixing and/or
                    �/    16.5
                                                             formulation. It is also possible that an error could
                                                             have occurred in the laboratory. For this reason,
                                                             samples sent to the lab should be split and a dupli-
     Barley                  79.7
     Soybean meal            17.3
     Premix                   3.0
                             100.0
                                                           128
  Mixed feed can be sampled from a number of                    Producers should check the calibration of their
feeders in the barn or as the feed is mixed. The key         mill to ensure that no errors have been made and
is to get a truly representative sample of the mate-         also the formulas for their diets to ensure that they
rial. Note that if feed is collected from the feeders,       are correct. Table 5-5 illustrates how to determine
poor analytical results may be caused by separation          the quantity of a nutrient in a diet. The mix size is
during handling and delivery, rather than by mixing          first corrected to 100. The nutrients supplied by
or formulation errors. These samples should be               each ingredient can then be easily calculated by
thoroughly mixed before submitting them to the lab           multiplying the concentration of nutrient in each
to ensure a good representative mix. If separation           ingredient by the amount of ingredient in the feed.
is being investigated, then they should not be mixed,        The amounts supplied by each ingredient arc added
but sent as individual samples to the lab.                   together to get the total quantity of nutrient in the
                                                             feed.
Table 5-4. Recommended Schedule for Feed Testing on a Farm Manufacturing Three Diets.
Month                                                           Sample
                                    Ingredients                                             Feeds
                       Home Grown                 Purchased                Starter         Grower            Sow
Jan                                                   c                       c               c               A
Feb                                                   c                       c               A                c
Mar                          c                        A                       A               c                c
Apr                                                   c                       c               c                A
May                                                   c                       c               A                c
Jun                          c                        A                       A               c                c
Jul                                                   c                       c               c                A
Aug                                                   c                       c               A                c
Sep                    A (new crop)                   A                       A               c                c
Oct                                                   c                       c               c                A
Nov                                                   c                       c               A                c
Dec                          c                        A                       A               c                c
C: Collect and retain sample; analyse only if problems are suspected. If there are no problems, retain sample in a
cool, dry, dark location for one year and then discard. In this way, if a problem occurs later, these samples will be
available for use.
A: Collect and analyse. In addition to this, twice a year samples should be selected from three or four different
feeders in each of the three to four sections of the barn. Analyse each sample to determine if separation is occurring.
  Table 5-4 provides a recommended schedule for                 The diet in Table 5-5 is well-balanced with respect
feed sampling and testing on a farm where three              to lysine, calcium and phosphorus. Assuming that
diets are being manufactured. The schedule as-               unusual ingredients are not used and that the premix
sumes that no problems are detected. If there is a           is correctly formulated, if these three nutrients
problem, more intensive testing will be needed               (lysine, calcium, phosphorus) are correct, the rest of
until the problem is resolved.                               the diet will likely be acceptable. It would not be
                                                             necessary to check each and every nutrient in the
                                                             diet.
                                                          129
   If problems in the feed analysis are discovered, it           the mill is totally out of calibration and serious
is important to check other sources of error such as             errors will result.
sampling or analysis before adjusting the mill. If
mill calibration is the problem recalculate the diet               In summary, if one obtains poor results from an
and adjust the mill according to the revised formula.            analysis, then the lab, the formula, the mill, the
It is imperative that the mill is not adjusted by 'one           delivery system and the method of sampling can all
or two clicks' to compensate for excess protein. If              be potential sources of error and should be re-
this practice is common, it will not be long before              checked. A flow chart summarizing the above
                                                                 procedures appears in Figure 5-2.
The diet contains the proportions of ingredients shown expressed as parts per I 00. Example: the lysine supplied by
barley will be 0.443 x 0.39 = 0.173.
                 r
                                             I
                                   ARE THE RESULTS
                                    SATISF�CTORY? ------,,
                  '---------
                    -        YES
                                                     �----- NO-------,
                                                           130
Conducting a Feeding Trial                                   7. Sex is another important factor. ln starter trials,
  Producers often need to compare two or more                sex can be ignored because there is little difference
feeds in terms of performance and profitability.             between barrows and gilts at this age. As they grow
The best way to compare feeds is to conduct a                older though, differences in growth rate and carcass
feeding trial in which the growth rate and feed              quality emerge.
efficiency of the pigs on various diets can be
compared. The trial must be carried out properly if            If the comparison keeps these points in mind, any
the results are to be meaningful. The comparison             differences observed in performance will likely be
cannot have biases that might favour one diet over           due to diet. Random variation in the performance
another. Provided these precautions are taken, one           of pigs does occur. Therefore, consistent differ-
can be reasonably assured that the differences               ences between diets arc needed to draw meaningful
observed are due to the diets used in the test and not       conclusions. For example, if pigs on diet A grew an
due to an outside influence. The following are               average of I 0% faster than pigs on diet B, but three
guidelines that will help ensure an effective                out of six individual pens grew fastest on diet A and
comparison:                                                  three grew fastest on diet B, the overall I 0% differ-
                                                             ence in meaningless and no apparent difference in
l. Split litters among the various diets being tested.       the diets exists. If there was an overall difference of
The effect of litter and genetics can be very large.         10% though, and five out of the six individual pens
especially in starter trials.                                grew faster on diet A, then one can be reasonably
                                                             certain that it is a better diet (Table 5-6).
2. Use more than one pen per diet even through it
increases the workload. If only one pen is used, the           Ensure diets contain the same medication, espe-
results in favour of one diet or another could very          cially when studying starter diets. Performance can
likely be due to chance. For a test to be effective , at     be greatly influenced by medication as well as by
least three to four pens per diet are required.              make-up of the diet. To compare only the diets,
                                                             differences due to medication should be eliminated.
3. Spread the pens on each diet throughout the barn          Again, the key to running a proper test is to elimi-
or room. This reduces biases due to pen location.            nate or at least minimize all sources of variabil ity
In almost all barns, there are pens that support better      other than the factor being evaluated.
performance than others.
                                                               Select the pens and the pigs for the feeding trial.
4. Treat all pigs on the diets equally. Factors such         Randomly assign pigs to the test pens assuring that
as feeder type, number of pigs per pen, pen size and         the average initial weight is constant and that litters
shape, pen flooring material, waterer numbers, type          are spread out among pens. If the experiment ends
and location can all have an affect on performance.          before the pigs reach 25 kg, do not separate them by
Therefore, the impact of such factors cannot be              sex. If older pigs are being used, sex differences
allowed to bias the results of your experiment.              start to show up and the number of barrows and
                                                             gilts in each pen should be the same. Record the
5. Be sure to compare diets at the same time.                total weight of pigs within each pen and randomly
Running one diet at one point in time and the                allot diets to the test pens. Add a known amount of
second diet at a later date could result in differences      feed to each feeder. As the experiment progresses,
in performance related to season or variable barn            continue to add feed as required, keeping records of
environment.                                                 how much feed was added to each pen. At the end
                                                             of the experiment, weigh all of the pigs and record
6. The average initial weight should be the same             the amount of feed left in each feeder. The
for all treatments. Heavier pigs will obviously grow         following calculations should be made to help draw
faster but convert feed less efficiently than smaller        conclusions:
pigs.
                                                           131
 I. final weight of pigs - starting weight of the pigs               If differences are small or results are in any way
      = total pig gain                                             questionable, repeat the experiment to strengthen
                                                                   and provide confidence in the results. In all cases,
2. total pig gain + number of pigs                                 remember that bad information is worse than no
     = total gain per pig                                          information at all. If the time, money and trouble is
                                                                   to be invested in an on-farm test, efforts should be
3. total gain per pig+ number of days on test                      made to carry it out properly.
     = average daily gain
                                                                   Common Calculations and
4. total feed added to feeder - feed left at end of
     trial = total feed consumed
                                                                   Conversions
5. total feed consumed+ number of pigs                             Converting 'dry matter' values to 'as fed':
     = feed consumed per pig
                                                                   Multiply the dry matter value by percent moisture.
6. feed consumed per pig+ number of days on test
     = average daily feed                                          Example 1: Convert dry matter to 'as fed'. A
                                                                   sample of barley has 11.2% crude protein (CP) on a
7. total feed consumed+ total gain                                 dry matter (DM) basis and contains 10.5%
     = feed conversion                                             moisture. Express the percent protein on an as fed
                                                                   basis.
8. feed cost per tonne+ 1000
     = feed cost per kilogram                                      Solution:
                                                                   If the barley contains I 0.5% moisture, it contains 89.5%
                                                                   (100-10.5 = 89.5%) dry matter. The amount of protein
9. feed conversion x feed cost per kilogram
                                                                   on an as fed basis then, is 10.0% (11.2 x 0.895 = 10.0%).
     = feed cost per kilogram gain
                                                             132
     6. NUTRITION OF THE BREEDING HERD
     Introduction                                               to service interval, a reduction in conception rates
        Adequate nutrition of the breeding herd is essen-       and a shortened reproductive lifespan. Therefore, it
     tial in order to maximize herd productivity and            is vital that the diet provided contains sufficient
     profit. If one compares the reproductive efficiency        quantities of energy, protein, essential fatty acids,
     of herds obtaining average productivity with those         vitamins and minerals to allow the sow to perform
     obtaining higher levels of performance, consider-          to its genetic potential.
     able room for improvement is evident (Table 6-1 ).
     Improvements in genetic quality and housing can               Since nutrition is a primary factor influencing
     account for some of the variation in reproductive          reproductive efficiency, it follows that the
     efficiency observed. However, maximum reproduc-            establishment ofa successful feeding strategy to
     tive efficiency can never be obtained unless the best      ensure optimum productivity must be based on a
     feeding and management practices are also fol-             sound knowledge of the response of the sow to
     lowed.                                                     specified nutritional inputs. Traditional feeding
                                                                strategies often utilized the body reserves of the gilt
     Table 6-1. Reproductive Efficiency of Average and          and sow to buffer short-term deficits in nutrient
     Superior Swine Herds.                                      intake with minimum effect on the fetus or suckling
                                 Typical                        piglets. However, modern sows, with a lean
                                                Superior
                                                                genotype and superior reproductive performance,
     Baby Pig Mortality(%)        12 - 18             8         must be managed differently from their counterparts
     Weaning to Effective                                       of twenty years ago since they begin their
     Mating (Days)                8 - 10              5         reproductive life with fewer body reserves.
     Farrowing Rate(%)           78 - 85             90         Therefore, nutritional responses must be considered
     Pigs Weaned/Litter          8.5-9.5            10.5        more precisely and a strategy designed to conserve
     Litters/Year                2.0-2.3             2.4        body tissue must be employed.
     Pigs Weaned/Sow/Year        17 - 22            25.2
                                                                Photo 6-1.
       Before defining their desired feeding program,
     pork producers need to establish goals for their
     breeding herd. Typical questions that are relevant
     include the genotype, the emphasis on sow
     longevity, the type of housing employed (indoor
     versus outdoor, group versus individual) and the
     nature of the cun-ent sow herd including factors
     such as appetite and body condition. Once such
     questions are answered, the nature of the feeding
     program can be more accurately defined.
,/                                                            133
Feeding and Management of                                     protein supplements during the developmental
Replacement Gilts                                             period to ensure that puberty is not delayed.
                                                              However, diets formulated to meet the protein and
  Survey data indicate that sow culling rates on
                                                              amino acid requirements of slaughter animals will
most commercial swine operations are somewhere
                                                              typically provide more than enough protein to allow
between 30 and 50%. As a consequence,
                                                              gilts to express their potential in terms of
replacement gilts will constitute a significant
                                                              minimizing age at puberty. Therefore, producers
proportion of the breeding females in most swine
                                                              should not be unduely concerned about protein
herds and any improvement in their productivity
                                                              restriction during this period. A diet formulated to
will have a significant impact on the reproductive
                                                              contain 15% crude protein ( 466 g/day)and 0. 7%
performance of the entire herd. Factors influencing
                                                              lysine (217 g/day) should be adequate.
gilt productivity include age at successful mating,
ovulation rate at the estrus of mating, first litter size
                                                                Diets formulated expecially for replacement gilts
and the ability to be successfully rebred. In far too
                                                              should contain higher levels of calcium and
many herds, the gilt is a neglected member of the
                                                              phosphorus compared with diets fed to feeder pigs.
breeding herd. However, by the adoption of an
                                                              The levels of calcium and phosphorus that result in
appropriate replacement policy and proper gilt
                                                              maximum growth rate are not necessarily adequate
management, the overall productivity of any swine
                                                              for maximum bone mineralization. Feeding of
enterprise can be markedly improved.
                                                              dietary levels of caJcium and phosphorus sufficient
                                                              to maximize bone mineralization in gilts during
Feeding Replacement Gilts During the                          early growth and development have been shown to
Rearing Period                                                improve reproductive longevity in some studies.
  Gilts selected to enter the breeding herd will              Therefore, dietary calcium and phosphorus
typically have superior growth rates and lower                requirements, expressed as a percentage of the diet,
backfat levels than unselected pigs. As a                     are higher for gilts than for barrows and diets fed to
consequence. it is reasonable to expect their                 replacement gilts should be formulated to provide a
nutritional requirements to be greater than those of          minimum of0.82% calcium (25.4 g/day) and 0.73%
pigs destined for slaughter. In addition, diets               phosphorus (22.6 g/day).
formulated to achieve cost-efficient, rapid growth
may not provide sufficient nutrients to prepare the             Genetic programs over the past 10-15 years have
gilt for a long and productive breeding life.                 placed considerable emphasis on selection for
Therefore, where possible, gilts destined to be used          leanness in gilts. The effects of these genetic
in the breeding herd should not be fed the same               changes, combined with earlier mating, mean that
diets as those fed to market animals.                         gilts now start their breeding lives with less body
                                                              reserves than in the past. This reduction in fat
  Nutrition during the rearing period (20-100 kg),            reserves could adversely affect the long-term
through its effects on body weight and backfat                reproductive performance of genetically improved
levels, can influence the age at which puberty is             strains of pigs. Australian workers have observed
attained. Several studies have shown that restrict-           that gilts which entered the breeding herd with
ing the feed intake of young growing gilts (50-85%            greater fat reserves were retained in the herd longer
of ad libitum intake) will delay the onset of puberty         and had a shorter average farrowing interval than
by about IO to 14 days. To ensure that puberty is             gilts with less fat reserves. It may therefore be
not delayed, gilts should be fed ad libitum and               desireable to encourage potential replacement gilts
receive at least 35 MJ DE/day (8361 kcal/day)                 to deposit more body fat. Alteration of the lysine/
between selection and mating.                                 energy ratio could achieve this goal.
                                                            134
age at puberty for non-stimulated gilts is about 200         With proper stimulation, the mean age of puberty
days of age, with a range of 135 to 250 days.              in a herd can be 30 to 40 days earlier than for non-
Canadian data indicate that fewer than I% of gilts,        induced gilts. Unfortunately, many producers who
housed and managed under typical commercial                have tried boar exposure have been unsuccessful in
rearing conditions, will reach puberty by market           inducing early puberty. The reason for their lack of
weight. Therefore, unless special steps are taken to       success is probably related to the fact that attain-
induce early puberty. most gilts will have a               ment of threshold levels of age and weight arc
prolonged interval between the age at which they           prerequisites for successful boar exposure. The
attain market weight and when they are able to be          recommended minimum ages and weights for the
successfully bred. The aim of successful                   commencement of boar exposure are 140 days of
management should be to keep this unproductive             age and 70 kg body weight. With modern geno-
time to a minimum.                                         types of swine, age is more likely to be a constraint
                                                           than is body weight and therefore it is unlikely that
  Although the normal pubertal age of a gilt is            gilts fed under commercial conditions will reach
under genetic control, there are various factors that      puberty much before 90 kg.
have been shown to delay or advance the age at
which a gilt reaches puberty. Factors known to               It is important that if early puberty is induced,
influence the age of puberty include the breed of the      gilts not be bred on their first heat as this practice
gilt, the housing conditions under which the gilt is       will result in a reduction in litter size. Research has
raised, the lighting regime and the degree of stress       shown that an extra one or two pigs per litter can be
experienced by the gilt (mixing and relocation). As        obtained by waiting until the second or third heat
a consequence, most producers could take steps to          before breeding replacement gilts. With induction
reduce the average age of puberty in their herd.           of puberty at 150 days of age, gilts gaining weight
                                                           at a rate of 800 g/day should reach the desireablc
  The most potent stimulus for inducing early              breeding weight of 115-125 kg by their third estrus
puberty in gilts is boar exposure. Fence line contact      (192 days). Target backfat levels at mating are 17-
is not adequate as there must be direct physical           20 mm.
contact between the boar and the gilt. The best
response is obtained by taking the gilt to the boar           A major concern of many pork producers is that if
pen, not vice versa. This is likely due to the fact        gilts are bred at too light a weight and with too little
that boar odors are stronger in the boar pen provid-       backfat, the gilts will have a shortened reproductive
ing a greater degree of stimulation for the gilt.          lifespan. However, recent research has shown that
However, in order to stimulate puberty, the gilt must      this need not be the case. Lean gilts provide a
be placed in the boar pen at least 20-30 minutes a         challenge to the producer, but if managed properly,
day, usually for a minimum of IO consecutive days.         acceptable performance can be expected from gilts
                                                           induced to reach early puberty.
Photo 6-2.
                                                           Feeding Replacement Gilts Prior to
                                                           Breeding
                                                              Ovulation rate is the principle factor limiting litter
                                                           size in gilts and there is a great deal of experimental
                                                           evidence showing that increasing the level of feed
                                                           intake during the rearing period will significantly
                                                           increase ovulation rate at puberty. lt is also well
                                                           established that short-term, high level feeding
                                                           (flushing) during the first estrus cycle increases
                                  [la......                ovulation rate compared with gilts fed restricted
The most potent stimulus for inducing early puberty in     amounts of feed. Dutch researchers have suggested
gills is boar exposure.                                    that ovulation rate increases by about two ova in
                                                         135
response to increased feed intake during the 14-day           Table 6.2. Influence Of Feeding Level Following
period immediately prior to ovulation. As a conse-            Mating On Embryo Survival in Gilts.
quence, should feed intake be restricted for any
                                                                                    Low Energy High Energy
reason during the rearing period, special precautions
                                                                                   (22.9 MJ/day) (40.5 MJ/day)
should be taken to ensure that gilts are ad lib fed (at
least 3 kg/day) for the two week period prior to              Ovulation Rate           12.3            13.8
mating.                                                       Number of Embryos         9.7            10. I
                                                              Embryo Survival (%)      78.3            73.2
  Flushing has been shown to increase plasma
levels of FSH and increase the pulse frequency of             Adapted from den Hartog and van Kempen, 1980, Neth.
                                                              J. Agric. Sci. 28: 211-227.
LI-I suggesting that flushing enhances ovulation rate
by stimulating the secretion of gondotrophins. The
increase in gonadotrophin secretion is thought to be
                                                            The principle mechanism controlling the
mediated through plasma levels of insulin and
                                                          development of embryos and their subsequent
insulin-like growth factor- I (lGF-1). The increase
                                                          survival is the secretion of uterine specific proteins.
in ovulation rate likely occurs either as result of
                                                          These proteins are stimulated by ovarian steroid
increased follicle recruitment or a reduction in
                                                          hormones, particularly progesterone. A rise in
atresia.
                                                          progesterone early in pregnancy enhances the
                                                          uterine environment and makes it more supportive
Feeding Gilts Following Mating.
                                                          of the embyro. Increased gestation feed intakes
   It has been determined that approximately 30% of
                                                          have been shown to be associated with a decrease in
all potentially viable embryos die during the first 25
                                                          plasma progesterone concentration as a result of
days of gestation. High levels of feed intake
                                                          increased metabolic clearance of progesterone
following mating have traditionally been associated       leading to a subsequent reduction in embryo
with an increase in embryo mortality. As a                survival (Table 6-3).
consequence, it is usually recommended that feed
intake be restricted to approximately 2.3 kg/day
 following mating. However, more recent evidence       Table 6-3. Effect of Feed Level in Early Gestation on
suggests that the practise of feed restriction to      Plasma Progesterone Levels and Embryo Survival
reduce embryo mortality may not necessarily result
in an increased litter size.                           Feed Level         Embryo Survival         Plasma
                                                                                               Progesterone
   Most of the research which has shown an increase                                           Concentration
in embryo mortality, as a result of higher feed          (kg/day)                   (%)           (ng/ml)
intakes early in gestation, has involved an increase
in feed intake at or very soon after mating. Such a       1.50                    82.8              16.7
practise would increase ovulation rate and this          2.25                     78.6              13.8
increase per se may lead to an increased embryo          3.00                     71.9              11.8
mortality since higher ovulation rates are commonly
associated with higher embryo loss. The higher         Dyck et al., 1980, Can. J. Anim. Sci. 60: 877-884.
ovulation rate followed by a higher embryo loss
produces a similar number of embryos (and pre-           Recent data from the University of Alberta
sumably piglets) to that obtained from gilts fed at a  suggest  that the first 72 hours after mating may be
lower level (Table 6-2).                               of critical importance in determining the effect of
                                                       feeding level on embryo loss. Increasing feed
                                                       intake from 1.8 to 2.5 kg/day during the first 72
                                                       hours of gestation significantly increased embryo
                                                       mortality whereas increasing feed intake after 72
                                                       hours did not increase embryo mortality. The
                                                        136
increased mortality in the first 72 hours was associ-         during this period of rapid mammary growth
ated with a 10 hour delay in the normal rise in               impairs development of the mammary gland in gilts
plasma progesterone. As noted earlier, a rise in              and should be avoided.
progesterone, early in pregnancy, enhances the
uterine environment and makes it more supportive              Summary of Replacement Gilt Feeding
of the embyro.                                                  Diets fed to potential replacement gilts should
                                                              contain 3100 kcal D.E./kg, I 5% crude protein,
  Clearly more research is needed on the effect of            0.7% lysine, 0.82% calcium and 0.73% phosphorus
feed intake in early gestation on gilt productivity.          and be fed ad libitum from the time of selection
However, given the current state of our ignorance,            (50-60 kg) until needed for breeding. Gilts selected
producers would be wise to continue the practise of           for mating should have experienced at least two
restricted feeding early in gestation, particularly           heat cycles, weigh 115-125 kg and have 17-20 mm
during the first 72 hours following mating.                   of backfat. If feed intake is restricted for any
                                                              reason during the rearing period, special precautions
   High feed intakes late in gestation may also be            should be taken to ensure that gilts are ad lib fed (at
detrimental to gilt productivity as they affect the           least 3 kg/day) for the two week period prior to
development of the mammary gland. In swine, total             mating. For the first 72 hours following mating,
mammary DNA can be used as an estimate of                     feed intake should be restricted to less than 2.5 kg/
mammary cell number and development. Minimal                  day.
development of the mammary gland occurs between
mating and day 50 of gestation. The period of
maximal mammary development occurs between
day 70 and 105. During this time, a threefold
increase in mammary tissue occurs as measured by
total mammary DNA. Recent research at Michigan
State University has shown that a high level of
clietary energy during this critical period of mam-
mary development reduced total marnmary
parenchymal DNA (Table 6.4).
                                                            137
per day (20 to 33 MJ/day) is satisfactory for preg-        that as the level of feed intake during gestation
nant sows housed under reasonable environmental            increases, the level of feed intake during lactation
conditions when free of heavy parasite infestation         decreases (see Table 6-23). Consequently, sows fed
and individually fed. Increasing the feed intake of        high levels offeed during gestation will eat less
sows above this level would appear to be of little         during their subsequent lactation and will start to
benefit.                                                   deplete their body reserves. Restricting feed intake
                                                           during gestation will prevent this loss of body
  The effect of increasing feed intake during              condition and may help to prolong the sow's repro-
gestation on subsequent litter size is shown in            ductive lifespan.
Figure 6-1. It is evident that above a threshold
level, there is a wide range of feed intake over           Methods of Restricting Feed Intake of Sows
which there is little effect on the number of piglets      During Gestation
born. At higher levels of intake the sow may                 Various management systems have been used
become overfat and litter size may decrease.               successfully to limit the energy intake of sows
   11                                                      during gestation. These include:
                                                             -Hand feeding using gestation stalls
                                                             -Computer controlled feeding stations
                                                             -Slow feeding systems
                                                             -Self-closing individual stalls
                                                             -Skip a day feeding
                                                             -Self feeding a high fibre ration
    7
        10        20         30         40         50      Gestation Stalls
        0.8      1.6         2.4        3.2        4.0
                                                             Individual daily feeding, utilizing gestation stalls,
                       MJ DE/Day
                       KG Feed/Day                         provides the greatest control over the feed intake of
                                                           the sow and is the method of choice for most
Figure 6-1. Effects of Feed Intake During Pregnancy
                                                           producers. Use of gestation stalls provides the
on Sow Productivity.
                                                           greatest control over the feed intake of the sow as
                                                           each sow can be fed to condition resulting in a
   Increasing feed intake during gestation will also       reduction in feed costs. In addition, the elimination
dramatically increase sow weight gain but has very         of fighting reduces prenatal losses.
little effect on the birth weight of the newborn pig.
Since these high levels of intake are not improving           Unfortunately, the increased cost of gestation
litter size or bi rth weight, considerable saving in       stalls makes the system unattractive to many pro-
feed costs can be achieved by restricting the feed         ducers and animal welfare groups have directed
intake of sows in gestation.                               criticism at them as well. Unless an automatic
                                                           feeding system is used, the gestation stall system is
  A reduction in feed costs is not the only benefit        very labor intensive. Furthermore. it has been
from restricting the feed intake of sows during            suggested that digestive upset (ulcers, twisted gut) is
gestation. There is some evidence that embryonic           worse in sows housed in gestation crates. There-
survival and thus litter size may be increased by          fore, alternative methods of restricting feed intake
restricting the level of intake during gestation. In       have been sought by some producers.
addition, farrowing difficulties may be reduced and
fewer piglets may be crushed by the sow.                   Electronic Sow Feeders
                                                             The electronic sow feeder has recently been
  Many producers believe that providing high levels        introduced into Canada, following earlier experi-
of feed during gestation will maintain the sow in          ences in Europe. With this technique. 40 sows can
good body condition. However, research has shown           be fed using a single feeding station. The amount
                                                         138
of feed that a sow gets each day is programmed into            Use of an electronic sow feeder does not appear
the computer. When it enters the station, the trans-         to have any detrimental effects on sow perform-
ponder the sow carries causes small portions of feed         ance. In a British study. conception rate, litter size
to be released. While the sow is eating, it is pro-          and the incidence of farrowing problems were
tected from other sows by a special crate which              similar for sows fed using an electronic feeder and
ensures that they cannot steal her feed.                     those housed under more traditional systems of
                                                             management (Table 6-5).
   In essence, this system allows sows to be group
housed but individually fed. The systems most
attractive features are that it allows the use of            Table 6-5. Comparison of Sows Fed with Electronic
                                                             Sow Feeders or in Gestation Stalls.
inexpensive, non-specialized buildings to house
sows during gestation and produces a daily report                                       Gestation      Electronic
on sow feed consumption. Electronic sow feeders
                                                                                         Stalls       Sow Feeders
also find favor with animal welfare enthusiasts
because they are compatible with straw bedding and           Number of Herds                485             27
allow the sow the chance to exercise.                        Litters/Sow/Year               2.3             2.3
                                                             Pigs/Sow/Year                 21.8            21.9
   Sows which are placed on the electronic feeder            Feed/Sow/Year (tonnes)        1.19            1.22
establish a regular feeding cycle. The more                  Sow Replacements(%)           19.3            15.9
dominant sows eat first and the more timid sows              Sow Mortality(%)               2.0             1.9
wait until later in the cycle. The computer system
allows the daily ration to be divided between a              MLC Pig Yearbook, 1989
number of feedings, but most sows consume all of
their daily allotment in one 12-15 minute feeding.
                                                               It must be pointed out that electronic sow feeders
If a sow which has already had its full ration returns
to the station, no further feed is dispensed.                are not completely fool-proof. Some sows have
                                                             learned that repeated banging of the feeding station
  It does not appear difficult to train sows to use the      can dislodge a small amount of feed which will not
                                                             be accounted for on the computer printout. Regular
electronic feeder. One trial conducted in Britian
indicated that 50% of sows required no training,             calibration is also necessary in order to correct for
                                                             differences in density between successive batches of
while a further 27% required only one assisted visit
to the feeding station in order to adapt to the              feed. In addition, collars and ear tags can be lost
                                                             thereby preventing a sow from feeding.
system. The majority of sows seem to learn to use
the electronic feeding system within four days
although there may be a few sows which never                    Many producers have expressed concerns about
adapt and must be culled.                                    the possibility of sows fighting during mixing and
                                                             while queuing at the feed station. However, in most
Photo 6-3.                                                   cases, the large area available for avoidance and
                                                             escape has meant that aggression has not been a
                                                             problem and in fact, sows using the system are
                                                             extremely docile. Vulva biting has been one of the
                                                             more persistent problems, prompting modifications
                                                             in design. If other problems develop, it may be
                                                             because the feeder has been incorrectly located. rt
                                                             is important that the feeding station be sited to
                                                             allow free movement about the entry and exit
                                                             points. The feeder should be placed in an activity/
                                                             dunging area rather than a lying area as incorrect
Sows housed outdoors require additional feed, especially     siting results in sows lying at the station entrance
in cold, damp weather.
               ��������������
                                                           139
and blocking the access of other sows to the feeder.         In the systems tested, feed was supplied twice a
At this time, it is suggested that straw bedding and      day from a volume dispenser and poured into a
not slatted floors be used in conjunction with            container with a small mortar in the bottom. The
electronic feeding.                                       speed at which the mortar turns dictates the amount
                                                          of feed dispensed. The dosing speed has been
  It is important to emphasize that electronic sow        investigated in relation to leftover feed and aggres-
feeders are not a replacement for good manage-            sive behavior (Table 6-6). The faster the feed is
ment. Sow condition must still be monitored and           dispensed, the less aggression develops and the less
adjustments to daily allotment made. Depending on         likely it is that sows will move away from their own
the system employed, heat detection and repeat            troughs. Howver, the amount of leftover feed
breeders can become a serious problem.                    increases. A dispensing speed of JOO to 120 g per
                                                          minute is recommended when pelleted feed is fed.
Slow Feeding Systems (Biofix System)
  Slow feeding systems are not presently available           In general, the slow feeding system works well.
in Canada but have been tested in the Netherlands.        The advantages of the system are that the sows can
Slow feeding systems dispense a continuous supply         be checked during feeding and that they can learn
of a small amount of feed at a predetermined rate.        the system without any assistance. A disadvantage
The feed supply must be fast enough to prevent            is that the sows cannot be fed according to their
sows from looking anywhere but their own troughs          individual requirements.
for feed. On the other hand, the dispensing speed
must be slow enough that even the most reluctant          Self-closing Individual Stalls (Woldrix System)
eater can keep up with the feed dispensed. In this          With self-closing individual stalls, it is possible to
way, all animals in a group will finish eating at the     have sows housed as a group and yet feed them
same time. This prevents the sows from fightng for        temporarily as individuals. The sow pen contains a
leftover feed. This so called biological fixation         sufficient number of feeders to allow each sow to
makes it unnecessary to lock up the sows in the           eat individually as well as communal lying and
feeding station. A simple 40-45 cm wide partition         dunging areas. A slotted floor is situated at the
between the troughs is sufficient to keep the sows        back of the pen and the sows have free access to all
contrained (Figure 6.2).                                  of the feeding stalls at all times except during
 [
     GRID
                 SOLID
                FLOOR
 [   --       - HEATED -     =GRID::::
[ TROUGH=
                                                        140
feeding (Figure 6.3). When feeding starts, the                        These systems have been tested in the Netherlands
entrance to each stall is locked automatically when                 since 1988. During and after feeding, there are very
the sow pushes her head against the trough lid. The                 few signs of aggression between sows. However,
stalls can also be locked manually by the stockman                  reproductive data is not yet available to allow a
as it is important that the entrance to each individual             comparison of the system with more traditional
pen be locked immediately after a sow enters to                     methods of sow housing.
prevent other sows from bullying them and driving
them away from the feeder. The feed required for                The Hurnick-Morris Housing System
each feeding place is stored in a volume dispenser                This system was developed as a result of a
and feed is poured into all the troughs at the same             cooperative study between the University of Guelph
time by the dispenser. Once all sows have con-                  and the Ridgetown College of Agricultural
sumed their feed, the doors can be unlocked and the             Technology. In the system tested, five pens are
sows are free to wander back to the dunging area.               located across the midsection of the barn (Figure
                                                                6.4). Each pen is 1 .6 x 6.6 meters in size and six
                                                                sows can be housed in each pen, providing
                                                                approximately 2 square meters per sow. Computer-
                                      self closing
                                                                controlled entrance and exit gates are located at the
                              1----1sHal�
                                       t                        end of each pen. Two water drinkers are located
                                                                near the exit gates. Two boar pens with space for a
                                                                breeding area are located adjacent to the group sow
walking area
   and    c:::¢>                                                pens. Along the walls of the barn is a 1.2 m
dunging area                                                    alleyway for approaching and departing the feeding
                                           � trough             compartments. The slatted areas extend along the
                                                                walkways and approximately 1.8 m into the pens at
                                                                the drinking/dunging end of the pens. The
                                                                remainder of the pen floor is solid and is suitable
                                                                for use with bedding.
                    � water
                      supply
Table 6-6. The Influence of Dispensing Speed on the Behaviour and Feed Intake of Sows Using the Slow
Feeding System.
                                                              141
                                                          minutes. After this exercise period, the crowd gate
                                                          slowly advances, the sows return to their pen and
                                                          the entry gate closes. The sequence repeats for each
                                                          successive pen of pigs until all sows are fed. Dur-
                                                          ing the exercise period, each sow has the opportu-
                                                          nity to vist the boars at the front partition of the
                                                          boar pen. Interrogating antenna are also located on
                                                          the boar pens to recognize sows that are spending
                                                          time with the boar. Future development of this
                                                          aspect is to use the computer as a tool to assist the
                                                          manager in detecting sows in estrus.
                                                        142
the condition of the sows to ensure that they are not            The size of the sow or gilt is going Lo affect the
being left for either too long or too short a time on          level of feed required. The heavier the sow, the
the self-feeder.                                               greater the maintenance requirement and the greater
                                                               the amount of feed required. Energy requirements
  Sows adjust readily to this system and from all              increase by about 5% for each IO kg increase in
research data, there appears to be no reduction in             body weight. The condition of the sow is another
reproductive performance as a result of interval               factor determining the feeding level required during
feeding. Most research indicates that sows fed on a            gestation. A thin sow will have less thermal
skip-a-day program farrow and wean a similar                   insulation than will a fat sow. Therefore, it will be
number of piglets as sows fed every day. However,              less able to adjust to lower environmental
no data on the success of this system is available for         temperatures. As a consequence, a thin sow will
herds weaning nine or more pigs per litter. For                require a larger increase in feed at a lower
intensive pork production, skip-a-day programs                 temperature than will a sow in good condition.
would not be recommended due to a lack of control
over individual sows and also to concerns about the               The environment in which the sow is housed
welfare of such systems.                                       should also be taken into consideration when feed
                                                               allowances are being set. Sows housed at lower
Diet Dilution                                                  environmental temperatures require more feed than
  A fourth method of restricting energy intake is to           sows housed in their comfort zone. The energy in
dilute the diet with a high fibre ingredients so that          the excess feed is used to produce heat to enable the
the sows may have constant access to the self-feeder.          sow to maintain a constant body temperature. For
Alfalfa hay, alfalfa meal, chopped straw or oat hulls          individually housed sows, the lower critical
have been utilized. This system takes less labour              temperature (temperature below which feed must be
than the other methods of limit feeding but is the             used to produce heat) is about 16- l 8°C. lf the
least acceptable because it costs more to maintain the         environmental temperature drops below this level,
sow and it is difficult to prevent sows from getting           feed intake should be increased by 3-4% for every
fat, even though the energy content of the feed is             I °C below the lower critical temperature. Sows
lower. In addition. there are problems associated              housed in wet or drafty conditions wilJ also require
with grinding high fibre feeds and such feeds tend to          increased feed.
bridge in the self-feeder. Therefore, the use of high
fibre feeds to limit the energy intake of sows during        Sows which are housed and fed in groups com-
gestation is not recommended.                              pete with each other for the available feed. There
                                                           may be considerable inequality in the feed intake of
Factors Affecting the Required Feeding                     individual sows. Therefore, sows which are fed in
Level During Gestation                                     groups should be given feed allowances which are
  Although there are many advantages to restricting        about 15% above that of sows fed individually.
intake during gestation, it must be emphasized that        This will ensure that those sows which are domi-
the actual feeding level will vary according to            nated by others receive feed intakes that are suffi-
individual situations and animals. Factors that            cient to prevent reproductive failure.
should be considered when determining individual
feeding levels include the:                                   The health of the herd is also going to affect the
  - size of the gilt or sow                                feeding level required during gestation. The effect
  - condition of the sow                                   of a disease burden is difficult to quantify in nutri-
  - method of housing                                      tional terms, but represents an important influence
  - environment provided                                   on the utilization offeed. For example, sows
  - method of feeding                                      infected with worms may actually lose weight
  - health of the herd                                     through gestation and produce smaller litters. This
  - productivity level                                     emphasizes the importance of monitoring for
  - standard of management.                                worms and deworming when necessary.
                                                         143
Feeding Pattern Throughout Pregnancy                       weight during pregnancy will enter lactation with
  It is often suggested that the nutrient requirements     low body stores of fat which will adversely affect
of pregnant sows are higher in early pregnancy             the ensuing lactation and increase the interval
when the embryos are being implanted and during            between weaning and successful mating. An
the last third of pregnancy when the fetuses are           example of the components of gestation weight gain
growing very rapidly. As logical as this may appear        is given in Table 6-8.
in theory, several large scale experiments have
shown that the performance of sows fed a constant
                                                           Table 6-8. Components of Gestational Weight Gain.
level of feed throughout pregnancy is as good as
that achieved by increasing feed intake in early or            Piglets ( 11 at 1.3 kg)               13.9 kg
late gestation (Table 6-7). Since a constant level of          Membranes                              2.5 kg
feeding does not impair reproductive efficiency, its           Uterine Fluids                         2.0 kg
simplicity should make it the method of choice.                Uterus                                 3.2 kg
                                                               Udder                                  3.4 kg
                                                               Sow                                   20.0 kg
Table 6-7. Effect of Feeding Level During Last 23
Days of Gestation on Reproductive Performance.                                                       45.0 kg
Cromwell et al., 1989, J. Anim. Sci. 67: 3-14.               Condition scoring uses a combination of visual as-
A cooperative research study involving 8 research          sessment of the sow's general appearance and an esti-
stations and I 080 Litters.                                mation of its body fat reserves to arrive at a number
                                                           or score. The chart in Fiqure 6-5 provides illustra-
                                                           tions as well as descriptions of the five body condi-
Evaluating Your Feeding Program During                     tion classifications into which sows are grouped.
Gestation                                                  Body fat is estimated by applying firm finger pres-
  Since there are so many factors affecting the level      sure over the top-rear (H-bones) of the pelvic girdle
of feed to be fed during gestation, producers are          (Figure 6-5). The amount of fat is judged by estimat-
advised to evaluate their feeding program to ensure        ing the degree of difficulty involved in locating and
that satisfactory levels of intake are being main-         palpating these structures. It should take no longer
tained. Two methods are available, one is to weigh         than three seconds to feel the H-bones with the fin-
sows to ensure adequate but not excessive weight           gertips. If it takes longer than this, the sow should be
gain and the second is to employ condition scoring.        rated either a four or a five.
The former is labour intensive, but objective. The
second requires less labour, but is subjective and           The first step in assessing the adequacy of your
therefore must be managed well to be successfuJ.           feeding program is to obtain an average score for
                                                           the dry sows in the entire herd. If the average is less
  The gilt or sow should gain weight during preg-          than three, then the daily feeding allowance should
nancy to compensate for the weight of the litter and       be increased. If the average score is greater than
fetal membranes as well as to allow for a normal           three, a decrease is necessary. Table 6-9 indicates
increase in sow body weight. Animals losing                how much feed allowances should be altered based
                                                           on the average condition score of the herd.
                                                         144
Table 6-9. Feed Allowances For Dry Sows According          Energy
To Body Condition Score.                                      For the gestating sow, energy is required for
                                                           maintenance, growth of maternal tissue and growth
    Condition Score         Change in Feed                 of the fetus. The maintenance energy requirement
                                (kg)                       is dependant on the size of the sow and has been
                                                           estimated to be approximately 110 kcal DE/kg 75 per
           1.0                   +   0.60
                                                           day and represents in excess of 75% of a sow's
           1.5                   +   0.40
                                                           daily energy requirement. To calculate maintenance
           2.0                   +   0.30
                                                           energy requirements, one must know the weight of
           2.5                   +   0.20
                                                           the sow at breeding and the expected weight gain
           3.0                       0.00
                                                           during gestation in order to calculate the average
           3.5                   -   0.20
                                                           weight of the sow during gestation. The traditional
           4.0                   -   0.30
                                                           thumb rule has been to target for 20-25 kg of
           4.5                   -   0.40
                                                           maternal weight gain and 20 kg of reproductive
           5.0                   -   0.60
                                                           tissue per parity at least up until the 5th parity when
                                                           mature body size is achieved. Table 6-10 converts
Froese, 1987, Manitoba Agriculture Swine Fact Agdex
                                                           actual body weight (BW) to metabolic body weight
No. 440-22-40.
                                                           (BW 75) A calculator that has a yx key can also be
                                                           used.
Nutrients Required During Gestation
  Based on the foregoing discussion, it can be               The energy requirement for maternal growth is set
concluded that the provision of a high quality diet is     by the desired body weight gain during gestation.
essential for sows during gestation. Such a diet is        The energy cost per kg of maternal gain is
necessary in order to provide adequate levels of           approximately 5 Meal of DE/kg. Therefore, the
nutrients to allow for fetal growth, growth of the         energy cost/day for a sow with a net weight gain of
uterus and mammary glands and body growth as               25 kg in a 114 gestation period is 1.10 Meal of DE
well as replenishment of the sow's body reserves           (125 Meal+ 114 days). The daily energy
depleted during the previous lactation. Therefore,         requirement for the growth of the conceptus has
producers are encouraged to conduct a feed analysis        been estimated at 0.19 Meal of DE resulting in a
to confirm that the diet being fed meets the gestat-       total energy cost/day of 1.29 Meal of DE.
ing sow's requirements.
Table 6-10. Metabolic Body Weights of Sows and Maintenance Energy Requirements.
Parity    Body Weight     Body Weight       Average Body        Metabolic Body       Maintenance Energy
          at Start (kg)   at Finish (kg)     Weight(kg)          Weight (kg·75)      Required (Meal/day)
                                                         145
                                        - -
Figure 6-5. Guides to Condition Scoring Sows, Including Photographs of Representative Animals.
                                                   146
Table 6-11. Energy Requirements of Sows (Estimated) During Gestation.
eonceptus gain.
"Assuming diet contains 3, 150 kcal DE/kg. Allowance should be increased by 300 to 400 kcal for every 5°C below
critical temperature ( I 6°C).
  Table 6-11 demonstrates how to calculate the            in order to maintain constant conditions. This is
daily energy requirements under three different           called the maintenance requirement and this loss has
conditions. The first column calculates the energy        been estimated in a number of experiments which
requirements for a gilt bred at 120 kg and gaining        have been averaged to produce the numbers in
45 kg during gestation. The 2nd column shows the          Table 6-12.
calculation for a 4th parity sow being bred at 195
kg and gaining 45 kg during gestation while the         The amino acid requirements for growth of
lastcolumn shows a mature sow bred at 270 kg and      maternal tissue can be calculated from the amino
gaining only 20 kg (reproductive tissue only) during  acid content of pork. The best available estimates
gestation. However, using the values above, pro-      of the protein content of pork suggest a value of
ducers can calculate daily energy requirements for    15.3%. Therefore. a sow gaining 45 kg (25 kg
any sow under any conditions. Dividing these          maternal and 20 kg reproductive tissue) during a
energy levels by the energy content of the feed will  115 day gestation will be depositing 59.86 g of
provide an estimate of the daily feed intake required protein per day (6885 g/115 days). By multiplying
to provide this level of energy (i.e., 5.82 Meal+ 3.2 this value by the amino acid content of lean tissue,
Meal/kg diet= 1.8 kg feed/day).                       one can calculate the required level of a specific
                                                      amino acid needed on a daily basis. It has been
Amino Acids                                           estimated that amino acids used for growth and the
  Amino acids are needed during pregnancy to          products of conception are used at an efficiency of
replace those lost through obligatory sloughing or    63.5% and we can assume that the gestation diet is
metabolism, to develop the pregnant uterus and its    about 80% digestible. Under these circumstances,
contents, to develop the mammary gland as well as     the value derived from multiplying the amount of
to add protein to the maternal body. The amounts      protein deposited daily by the sow by the amino
of the indispensable amino acids needed during        acid content of lean tissue must be divided by a
pregnancy can be estimated by summing the             factor of .508 (63.5% efficiency x 80% digestion)
amounts needed for maintenance and for maternal       to arrive at the daily amino acid pattern required for
protein accretion.                                    growth and reproductive tissue. A mature sow (ie
                                                      one no longer gaining maternal tissue) will only
  During gestation. there is a continous sloughing of gain 20 kg during gestation and therefore the daily
cells from tissues such as the skin and intestinal    protein deposition will be only 26.6 g/day (3060 gl
mucosa. These represent obligatory losses of amino 115 days). For example calculations of the daily
acids from the body and this loss must be replaced    amino acid requirements sec Table 6-13.
                                                        147
Table 6-12. Daily Amino Acid Requirements for Maintenance of Gestating Sows (g/day).
                                     Amino Acids
                                      Required for
                                 Maintenance (mg/kg75)1                         Amino Acid Required (g/day)
Arginine                                      O
Histidine                                     O
Isoleucine                                   30                             1.23              1.69              2.05
Leucine                                      20                             0.82              1.13              1.37
Lysine                                       25                             1.03              1.41              1.71
Total Sulfur Amino Acids                     26                             1.07              1.47              1.78
Total Aromatic Amino Acids                   46                             1.89              2.60              3.14
Threonine                                    39                             1.61              2.20              2.66
Tryptophan                                    5                             0.21              0.28              0.34
Valine                                       21                             0.87              1.19              1.43
'Based on Baker et al ( 1966; J. Nutr. 88: 382-396) and Baker and Allee (1970; J. Nutr, 100: 277-280).
Table 6-13. Daily Amino Acid Reqirements {g/day) for Growth of Gestating Sows (Growth and Products of
Conception).
Arginine                                6.71
Histidine                               4.29                             2.24                            5.03
Isoleucine                              4.32                             2.26                            5.07
Leucine                                 7.38                             3.85                            8.68
Lysine                                  7.90                             4.13                            9.29
Total Sulfur                            3.32                             1.73                            3.89
Total Aromatic                          7.48                             3.91                            9.17
Threonine                               4.16                             2.16                            4.90
Tryptophan                              1.01                             0.51                            1.18
Valine                                  5.10                             2.65                            6.00
                                                             148
   To calculate the total daily amino acid                      The above example likely provides more detail
requirements of sows in gestation, one must sum the          than would be required by the average producer and
maintenance requirement and the requirement for              it would be virtually impossible to provide a diet
growth. Table 6-14 demonstrates how to calculate             that provides precisely the amino acid balance
the daily amino acid requirements under three                shown in Table 6-14. However, since lysine is the
different conditions. The first column calculates            first limiting amino acid, under these conditions, it
the amino acid requirements for a gilt bred at 120           is the amino acid we should be most concerned with
kg and gaining 45 kg during gestation. The 2nd               in diet formulation. Using the estimated daily
column shows the calculation for a 4th parity sow            lysine requirements shown in Table 6-14 and
being bred at 195 kg and gaining 45 kg during                dividing by expected daily feed intakes in Table 6-
gestation while the last column shows a mature sow            11, one arrives at the percentage of lysine required
bred at 270 kg and gaining only 20 kg (reproductive          in the diet to meet requirements (i.e. 0.56%, 0.45%
tissue only) during gestation. However, using the            and 0.24% for the 120, 195 and 270 kg sow,
values shown above, producers can caJculate daily            respectively, compared with 0.43% for NRC).
amino acid requirements for any sow under any
conditions.
Table 6-14. Estimated Total Amino Acid Needs of Pregnant Sows Under Various Conditions (g/day).
  Some flexibility can be obtained by varying the            NRC ( 1988) and the Australian Standing
level of feed intake to meet the lysine requirements         Committee on Agriculture ( 1987). Reasonable
of sows of different weights. However, larger                agreement exists between these two groups in
producers might find it cost effective to formulate at       setting nutritional standards. Where differences
least two diets varying in lysine content and feed           exist, it is suggested that the Australian Standards
one to sows less than five parities and one to sows          be used because these standards tend to be based on
over five parities.                                          ingredisimilar to those most commonly used in
                                                             Canada and the predominant breeds in the two
Minerals                                                     countries (Large White and Landrace) are also
  The most commonly used estimates of mineral                similar. Current recommendations are shown in
requirements of gestating sows are those of the              Table 6-15.
                                                         149
Table 6-15. Mineral Requirements of Gestating Sows           Table 6-16. Vitamin Requirements of Gestating Sows
(% or amount/kg of total diet).                              (Total Supplied in Diet).
Vitamins
  Vitamins have long been recognized as having an          Folic acid has received considerable attention in
essential role to play in reproduction and a consid-     the past few years with several experiments show-
erable amount of research has been devoted towards       ing a response to supplementation at levels greatly
determining the vitamin requirements of the gestat-      in excess of those currently recommended. The
ing sow. Current recommendations are shown in            most common finding has been an increase in litter
Table 6-16.                                              size (Table 6.17). As a consequence, folic acid
                                                         levels of approximately I mg/kg or about three
   Feeding recommendations must continually be           times current NRC ( 1988) levels are recommended.
modified as more research is conducted and we
learn more about the nutrient requirements of
                                                         Table 6-17. Effects of Felic Acid Supplementation of
animals. 1n the past few years, new evidence has
                                                         Diets Fed to Gestating Sows.
come to light which suggests that current recom-
mendations are inadequate for several vitamins and
                                                                                  Control        FolicAcid
it is likely that the next set of requirement tables
                                                                                               ( 1.00 mg/kg)
will show a recommendation for higher levels of
supplementation for these vitamins. These changes        Pigs Born Alive           9.51            10.64
will likely affect folic acid beta-carotene and          Birth Weight (kg)         1.48             1.48
vitamin E.                                               Pigs Weaned (day 29)      8.92             9.24
                                                         Weaning Weight (kg)       7.68             7.44
                                                         Weaning to Estrus
                                                         Interval (days)           7.30            6.66
                                                       150
   The increase in litter size as a result of folic acid     proteins with immunosupprcssive capabilities have
supplementation has been attributed to a decrease in         been identified. These proteins play a key role in
embryo mortality. The rate of cell proliferation             embryo development and could explain the
during embryonic development is extremely high               increased litter size observed with beta-carotene
and the intracellular concentration of RNA, a key            injection. Beta-carotene may also increase the
component, is highly correlated with embryo                  production of progesterone during the initial
survival. The synthesis of the nucleic acids DNA             formation of the corpora lutea. which would
and RNA requires purines and pyrimidine bases, the           provide a more favorable environment for embryo
production of which in tum requires single carbon            survival. Unfortunately, injectable beta-carotene is
units. As folic acid is an indispensable cofactor in         not currently cleared for use in Canada.
the metabolic transfer of single carbon units, it is
logical to assume that an adequate supply is essen-             Vitamin E has also received considerable research
tial for minimizing embryo losses.                           attention and again there are suggestions that current
                                                             recommendations (22 JU/kg) may not be adequate
  Another area of recent research interest has been          for high producing sows. A recent Ohio study
in the use of beta-carotene injections around the            examined supplementation with 16, 33 or 66 JU
time of mating. Beta-carotene is a natural precursor         vitamin E for three parities and observed increased
of vitamin A, but recent evidence suggests that it           litter size at birth and weaning as the vitamin E level
may have a unique role in reproduction independant           increased (Table 6.19). The data suggest that sows
of its function as a precursor of vitamin A. The             housed in less sanitary conditions respond more
most common finding with beta-carotene injection             positively to higher vitamin E levels with reduced
is a modest improvement in litter size (Table 6-18).         incidence of mastiris than those housed in clean
The increase in litter size is suggested to result from      facilities.
a decrease in embryo mortality. Dietary supple-
mentation (as opposed to injections) is unlikely to
                                                             Table 6-19. Effect of Vitamin Eon Sow Productivity.
result in an increased litter size because beta-
carotene is not absorbed intact from the digestive                               Supplemental Vitamin E (fU/kg)
tract of swine.
                                                                                         0      16      33     66
Table 6-18. Effect of Injection of Beta-Carotene on
Reproductive Performance of Multiparous Sows.                Pigs Born Alive        9.85 10.87 11.20 10.04
                                                             Birth Weight (kg)      1.39 1.31 1.37 1.41
                         Control       Beta-carotene1        Pigs Weaned (28 days) 6.73 7.00 7.88 8.14
Days to Estrus              4.6              4.5             Weaning Weight (kg)    6.51 6.27 6.21 6.49
Farrowing Rate(%)          88.7             88.4             Piglet Survival(%)    68.30 64.30 70.30 81.00
Pigs Born Alive            10.0             10.6
                                                             'Basal diet contained 8.4 mg/kg vitamin E. Experiment
Birth Weight (kg)           1.5              1.5             conducted over 3 parities.
                                                             2Mahan, J 991, J. Anim. Sci. 69: 2904-2917.
'Sows were given i.m. injection of200 mg of Beta-
carotene on the day of weaning, breeding and day 7 of
gestation.
2Coffey and Britt, 1993, J. Anim. Sci. 71: 1198-1202.        Feeding the Sow Around Farrowing
                                                             Time
                                                             Feeding Levels
  The mechanism by which beta-carotene enhances
                                                               Opinions as to the level of feed to be provided
embryo survival is not certain. However, it has
                                                             immediately prior to and for the first few days after
been shown that beta-carotene can increase the
                                                             farrowing arc divided. Excessive restriction can
production of uterine specific proteins which
                                                             cause excessive sow excitement due to hunger
support embryo survival. A basic glycoprotein with
                                                             resulting in an increase in piglet deaths due to
iron binding capacity and a groups of acidic
                                                           151
crushing. There is also concern about feeding too          result in a reduction in preweaning mortality.
much. Therefore, a balance has to be kept in               Furthermore, the increased energy level in the sow
relation to the feeding level around farrowing.            diet may decrease sow weight loss during lactation.
Before the sow farrows, it is probably best to
maintain the same level of feed intake as that               The response to fat inclusion is variable and a
normally fed during gestation (2 - 2.5 kg). Follow-        beneficial response has not been reported in all
ing farrowing, a gradual increase in feed intake is        cases. The amount of fat fed prior to farrowing
recommended, with the objective of getting the sow         affects the response obtained. ln general, the
to maximize feed intake as soon as possible into her       greater the amount of fat fed, the greater the re-
lactation.                                                 sponse. It appears that sows must be fed at least
                                                           one kg of fat prior to farrowing in order to obtain
Use of Laxatives Prior to Farrowing                        any benefit. Therefore, it is usually necessary for
  Many producers add wheat bran, beet pulp or              fat to be included in the diet at a level of at least
some other bulky feed to the sow's diet before and         10%. In addition, fat must be fed for at least five
for a few days after farrowing. It is a believed that      days prior to farrowing to obtain a positive re-
this practice helps prevent constipation, reduces the      sponse. This time period is required in order to
incidence of mastitis and may prevent death loss           allow the sow time to make the physiological
due to twisted gut. The most recent research on this       adjustments necessary to absorb the fat and trans-
subject does not support the need for the addition of      port it to the mammary glands.
such bulky feeds as a means of preventing mastitis
or improving sow productivity. However, the                   Unfortunately, the inclusion of a high level of fat
addition of bulking agents may improve sow                 in the diet is relatively expensive and may result in a
comfort and produce a softer stool. If sows are            very greasy feed. The potential to obtain an eco-
experiencing problems with constipation, producers         nomical response is greatest when mortality is high
may wish to consider the addition of 5 to 15% bran         and birth weights are low. Most producers will find
or 5 to 7% beet pulp to the prefarrowing diet.             it uneconomical to routinely include fat in their sow
Potassium chloride or potassium magnesium sulfate          diets unless preweaning mortality is greater than
have been used as laxative agents at the rate of0.5        25% or the incidence of low birth weight piglets is
to 1.5% of the total diet. However, the longterm           inordinately high. However, producers should not
effect of such salts on the health of the sow are          underestimate the benefits of high fat feeding when
unknown.                                                   environmental temperatures rise above 25°C.
                                                           Under these circumstances, high levels of dietary fat
Feeding Fat in Late Gestation and Lactation                are beneficial in maintaining energy intakes of
  A topic of recent research interest has been the         lactating sows, increasing sow milk production and
addition of fat to sow diets during late gestation and     thereby increasing survival and weaning weights of
lactation in an attempt to improve the survival of         piglets.
baby pigs. Baby pig losses are often high during
lactation, with the greatest losses attributed to             Most of the early work on fat supplementation of
crushing by the sow. Pigs which are crushed are            sow diets involved the use of fats made up pre-
often hypoglycemic due to low glycogen stores.             dominantly of long chain (>Cl6:0) fatty acids (i.e.
These low energy stores result in weak piglets             tallow or soybean oil). An area of recent research
which may be unable to move out of the way of the          interest has involved the study of fats made up
sow and are subsequently crushed.                          predominately of medium or shorter chain fatty
                                                           acids ( <C: 12:0). Coconut oil is one readily avai 1-
  It has been hypothesized that the incorporation of       able source of medium chain fatty acids. These
fat into diets fed during late gestation and lactation     shorter chain fatty acids are known to more easily
will increase both milk yield and milk fat content,        digested by swine and have found use as energy
thus increasing the energy supply available to the         sources in creep and starter diets. However, it is
nursing pig. This increase in available energy may         only recently that they have been studied as an
                                                         152
Table 6-20a. Examples of Gestation Diets
Ingredients, %                                           2             3                 4        5                6
Barley                                   86.0          87.2                                      78.8
Corn                                                                  82.0          72.5                       48.5
Peas                                                                                             10.0
Wheat shorts                                                                                                   40.0
Soybeanineal-47°/o                       4.5            7.8           13.0                                      6.5
Canola rneal                             4.5                                        22.5         6.2
Premix                                   5.0            5.0           5.0            5.0         5.0               5.0
Nutrients, minimum %
D.E., kcal/kg                           3,000          3,000         3,390          3,300       3,000         3,250
Crude protein                            l3.0           13.0          13.1           14.8        13.1          14.3
Digestible lysine                        0.43           0.43          0.49           0.48        0.43          0.46
Digestible methionine                    0.16           0.16          0.18           0.18        0.16          0.17
Digestible TS.A.A.                       0.20           0.20          0.23           0.22        0.22          0.22
Digestible threonine                     0.27           0.27          0.31           0.30        0.27          0.29
Digestible tryptophan                    0.06           0.06          0.07           0.07        0.06          0.07
Calcium                                  0.85           0.85          0.85           0.85        0.85          0.85
Phosphorus                               0.75           0.75          0.75           0.75        0.75          0.75
Sodium                                   0.15           0.15          0.15           0.15        0.15          0.15
Chloride                                 0.15           0.15          0.15           0.15        0.15          0.15
Ingredients, %                             l             2             3                 4        5                6
Wheat                                   50.00                        50.00                      49.67
Barley                                  26.30                        21.90                      25.20
Com                                                    76.00                        71.70                     70.65
Soybean meal - 47%                       I 1.70        19.50         15.80          23.30        15.10        24.20
Canolameal                               7.50                         7.50                       3.20
Fat/oil                                   1.00          1.00          1.30              1.00      2.50         l.00
LysineHCI                                                                                        0.23          0.10
Threonine                                                                                        0.10          0.05
Premix                                   3.50          3.50           3.50          3.50         4.00          4.00
Nutrients, minimum %
D.E .. kcal/kg                          3,250          3,460         3,300          3,415       3,350         3,460
Crude protein                            18.6           15.6          19.3           17.2        18.0          17.5
Dig. lysine                              0.65           0.65          0.75           0.75        0.85          0.85
Dig. methionine                          0.16           0.16          0.19           0.19        0.21          0.21
Dig. T.S.A.A.                            0.33           0.33          0.38           0.38        0.43          0.43
Dig. threonine                           0.41           0.41          0.47           0.47        0.54          0.54
Dig. Tryptophan                          0.10           0.10          0.11           0.11        0.13          0.13
Calcium                                  0.85           0.85          0.85           0.85        0.85          0.85
Phosphorus                               0.75           0.75          0.75           0.75        0.75          0.75
Sodium                                   0.15           0.15          0.15          0.15         0.15          0.15
Chloride                                 0.15           0.15          0.15           0.15        0.15          0.15
These are sample diets only and are for illustrative purposes only. While every attempt has been made to present
examples that reflect successful commercial formulations, these examples arc not intended for actual use without
assistance from a qualified nutritionist.
All amino acid concentrations are expressed as apparent ileal digestible amino acids.
                                                             153
ingredient in sow diets. The results of a recent            synthesis of lipid and to spare glucose, they have
study conducted at the University of Georgia are            the potential to improve fetal energy stores at birth
shown in Table 6-21.                                        and thereby improve the piglets chances of survival.
Table 6-21. Effect of Type of Fat Fed During Late           Feeding the Sow During Lactation
Gestation and Early Lactation on Sow Productivity.          Introduction
                                                              Lactation is a particularly important stage of the
                         Control     Long     Medium        reproductive cycle. Its primary purpose is to
                                    Chain Triglycerides
                                                            successfully rear a large number of healthy, heavy
                                                            piglets. A lactating sow may need to draw on body
Pigs Born Alive      11.33           10.79      11.59
                                                            fat reserves to obtain the energy she needs for
Birth Weight (kg)      l.27            1.31       1.21
                                                            maintenance and milk production. This drain on
Pigs Weaned (day 21) 9.06             8.89      10.12
                                                            her body reserves causes a significant loss of body
Weaning Weight (kg) 5.48              5.62       5.51
                                                            weight resulting in an extended weaning to remating
Mortality(%)         20.03           17.61      12.68
                                                            interval, poor conception rates and premature
Azain, 1993, J.Anim. Sci. 71: 3011-30 J 9.
I                                                           culling. Therefore, particular care must be taken to
Fat sources fed at a level of 12% during the last three
2                                                           ensure that sows are properly fed during lactation.
weeks of gestation and the first week of lactation.
                                                              The importance of maintaining high feed intakes
                                                            during lactation has been clearly demonstrated.
     The results of this experiment suggests that           The results of a study in which lactating sows were
medium chain triglycerides are superior to long             fed between 1.5 and 5.0 kg of feed per day are
chain triglycerides in reducing preweaning mortal-          shown in Table 6-22a. Sows receiving low levels of
ity. The difference in performance would appear to          feed during lactation lost significantly more weight
be a reflection of the different ways in which these        and depleted their backfat reserves to a much
fat types are metabolized. Medium chain                     greater extent than did sows fed higher levels of
triglycerides are rapidly metabolized to ketone             feed. Consequently. sows fed low levels of feed
bodies. Because ketone bodies can readily cross the         during lactation had longer weaning to conception
placenta and are used in the developing fetus for the       intervals than sows well fed during lactation. In
                                                            addition, significantly fewer sows fed low levels of
                                                            feed exhibited estrus within eight days of weaning.
Table 6-22a. The Effect of Sow Feed Intake During Lactation on Subsequent Reproductive Performance.
Lactation Weight Loss (kg)                      44.5       30.8        27.4        19.6        15.8         9.0
Backfat Loss During Lactation (mm)               8.9        7. L        6.4          5.7        4.2         4.0
Weaning to Conception Interval (day)            29.8       32.4        23.6        16.4        15.5        11.4
Number of Eggs Ovulated                         12.2       13.3        10.9        l 3.3       11.7        12.0
Sows in Estrus Within 8 Days of                  8.3       33.3        50.0        58.3        58.3        83.3
 Weaning(%)
                                                          154
  The level of feed provided to the sow during                 Methods of Increasing Feed Intake During
lactation will also affect piglet performance. Re-             Lactation
search has shown that there is a direct relationship         One method of increasing feed intake during
between the amount of feed the sow consumes                lactation is to ensure that the sow is not overfed
during lactation and the amount of milk produced           during gestation. Research has shown that there is
by the sow (Table 6-22b). As the level of feed             an inverse relationship between the amount of feed
intake increases, higher levels of milk production         consumed during gestation and the amount
are achieved. This increased milk production also          consumed during lactation (Table 6-23). As the
increases the growth rate of suckling piglets.             level of feed intake during gestation increases, the
                                                           level of feed intake during lactation decreases.
                                                           Therefore, feed intakes during gestation should be
Table 6-22b. Effect of Feed Level on Milk Yield at 21
                                                           reduced if excessive, since the higher the feed
Days (kg/day).
                                                           intake during pregnancy the lower the appetite will
                                                           be during lactation.
                         Feed Intake (kg/day)
                        4.5   5.3    6.0       6.8
                                                           Table 6-23. Effect of Feed lntake During Pregnancy
  Parity 1              5.9   5.4    6.7       6.1         on Feed Intake During Lactation.
  Parity 2              5.4   6.0    6.6       6.6
  Parity 3              5.5   6.8    7.3       8.0                             Pregnancy Feed Intake {kg/day)
                                                         155
Table 6-24. Effect of Feeding Various Levels of Protein During Lactation on Sow Feed Intake and Body
Condition.
  Sows will consume more feed if fed twice rather           It has been well demonstrated that sows will
than once daily. Consumption is likely to increase        consume more of a wet feed than they will of a dry
further if they are fed even more frequently. ln a        one. The results of one experiment are presented in
study conducted by the NRC-89 Committee on                Table 6-25 and it can be seen that sows fed a wet
Confinement Management of Swine, sows were                feed consumed approximately I 0% more feed than
allowed unlimited access to feed either once or           sows fed a dry one. Although it would be impracti-
three times a day. Sows fed three times a day             cal for most producers to convert their operations to
consumed a total of I 08.4 kg of feed during lacta-       a wet feeding system, the simple act of mounting
tion while those fed only once a day consumed             the water nipple in the farrowing crate directly over
101.6 kg of feed. In addition, weight loss during         the feed hopper can help to stimulate sow feed
lactation was reduced for the group of sows fed           intakes. However, care should be taken to clean the
three times a day (22.5 versus 28.5 kg of body            feeder on a regular basis in order to prevent molds
weight lost).                                             from developing on the wet feed.
                                                        156
sufficient. The only difference between the two              The effective environmental temperature of the
flow rates is a large waste of water at the 2.0 L per      farrowing room is one of the most critical factors
minute flow rate. Waterer type may affect feed             affecting feed intake in lactating sows and many
consumption. University of Minnesota studies               producers maintain their farrowing room at too high
revealed that daily feed consumption was about 0.3         a temperature. Table 6-27 shows the results of an
kg less when lactating sows were provided water via        experiment in which sows were housed in farrowing
nipples compared with a drinking cup.                      rooms maintained at 27°C or 2 I °C. Sows main-
                                                           tained at the lower temperature consumed more
  The use of flavors in animal feeds has increased         feed, lost less weight and weaned heavier piglets
considerably in the past decade as more attention is       compared with those housed at the higher tempera-
being paid to palatability. Many experiments have          tures.
been conducted using feed flavors in the hope of
increasing the feed consumption of sows during               In order to stimulate feed intake, attempts should
lactation. The productsused have ranged from               be made to cool down the farrowing room. If
simple spices and tonics to aroma modifiers, sweet-        building a new barn, producers might consider the
eners, flavor intensifiers and artificial flavors as       installation of snout coolers or drip cooling. Snout
well as certain natural feed ingredients.                  coolers with an airspeed of 0.3 m/sec have been
                                                           shown to increase feed intake by 250 g/day while
  The results of two experiements in which sow             drip coolers, providing 2 litres per sow per hour
diets were supplemented with a flavor enhancer are         (running for 3 minutes in a JO minute cycle), have
summarized in Table 6-26. Sows fed diets supple-           also been effective in increasing feed intake. How-
mented with a flavor enhancer consumed approxi-            ever, it must be remembered that if the temperature
mately 7% more feed and lost 30% less weight               of the farrowing room is to be decreased, supple-
during lactation than did sows fed the control diet.       mental heat must be provided for the piglets.
In addition, there was a slight reduction in piglet
mortality as well as an increase in piglet weaning            An additional consideration regarding lowering
weight as a result of including the feed flavor in the     the temperature of the farrowing room is to ensure
sow's diet. Although these results indicate that           that the building is properly insulated. The standard
flavor additives can be used to attract sows to their      insulation specification for the roof is 0.4W/m2/°C
diet, increase feed intake and reduce sow weight           and for walls 0.5W/m2/°C. The maximum ventila-
loss during lactation, producers should be aware           tion rate of the farrowing room should also be
that very little is known on what specific flavors         checked to ensure adequate airflow. Although this
pigs find attractive. Since flavours tend to be            may not necessarily cool the sows, fresh, non-
expensive feed ingredients, considerable caution           gaseous air is beneficial in stimulating appetite.
should attend their use in sow diets.
                                                             The farrowing floor surface also influences heat
                                                           loss and therefore could affect feed intake. Sows
Table 6-26. Performance of Sows Fed Artificial             housed on plastic-coated, expanded metal or woven
Flavours During Lactation.
                                                           wire floors consumed about 0.5 kg less than sows
                                    Control Flavour        housed on highly conductive floor surfaces such as
                                                           metal (ie Tri-bar) or concrete.
Pigs Born Alive                        9 .9       9.9
Pigs Weane                             8.4        8.9         Energy intake can be affected by increasing the
Mortality(%)                          14.9        9.9      caloric density of the diet. Use of high density diets
Sow Feed Intake (kg/day)               4.8        5.2      is particularly effective in maintaining energy
Lactation Weight Loss (kg)            15.6       10.9      intakes during hot weather. Incorporating fat or oil
Weaning to Estrus Interval (days)      7.5        8.3      as a partial substitute for carbohydrate energy will
                                                           reduce the heat increment of the diet and thus allow
Moser et al. 1986. Minnisota Swinea Research Report.       sows to consume more energy in hot weather.
pp 53-56.
                                                         157
Reducing the protein level of the diet and using           lation for producers to make and most nutritionists
synthetic amino acids to provide the proper balance        use a thumb rule of I% of a sow's body weight to
of amino acids can also achieve this purpose.              estimate maintenance requirements. Therefore, a
                                                            165 kg sow requires 1.65 kg of feed for mainte-
                                                           nance. If we assume a energy value of the feed of
Table 6-27. Effect of Environmental Temperature on
Feed Intake and Weight Loss of the Sow and Weight          3200 kcal then this level offeed intake would
Gain of the Litter,                                        supply 5.28 Meal of energy which comes reason-
                                                           ably close to the 5.06 Meal of energy we would get
                                  Temperature              by using the above equation (See Table 6-1 I).
                                27°C        21°C
                                                             The energy level required for milk production is
Sow Feed Intake (kg/day)       4.6           5.2           estimated to be 2 Meal of DE per kg of milk pro-
Sow Weight Loss (kg)                                       duced. This is calculated by assuming that the gross
I IO Days to Weaning          21.0          14.0           energy of milk is 1.3 Meal DE per kg and that the
Piglet Weight at 28 Days (kg) 6.2           7.0            milk is produced with a 65 percent efficiency of
                                                           utilization ( 1.3 Meal+ 0.65 = 2.0 Meal). The milk
Lynch, 1978.                                               production of a sow can be estimated from piglet
                                                           weight gain. Since it takes about 4 g of milk to
  Recent evidence has suggested that sows exposed          produce 1 g of piglet gain, total milk production can
to longer periods of lighting during the day may           be calculated by multiplying the daily weight gain
have higher feed intakes. Increased weaning                of a litter by four. As an example, a litter of 9
weights and improved rebreeding performance have           piglets gaining 240 g/day would suggest a daily
been observed when sows were housed in farrowing           milk production of 8.64 kg (9 x .24 x 4). This level
rooms with 16 hours of light per day in comparison         of milk production would require an energy intake
with 8 hours of light per day. Automatic timing            of 17.28 Meal DE (8.64 kg x 2 Meal/kg milk
devices are currently available which allow produc-        produced).
ers to control the duration of lighting in the farrow-
mg room.                                                     It is evident from the above discussion that no
                                                           single energy level will apply to all sows under all
  To properly monitor lactation feed intakes, it is        conditions. The following table indicates how
recommended that producers keep track of the               daily energy requirements are altered by factors
individual feed intake of sows using a card similar        such as sow body weight, litter size and piglet
to that shown below (Fiqure 6-5). Use of such              growth rate (Table 6-28).
cards creates an easily interpreted, graphic display
of the pattern of intake of each sow. lt is only              Table 6-28 indicates that a 142.5 kg sow nursing
through the use of a proper recording scheme that          12 piglets, gaining 240 g/day will require 8.34 kg of
producers will be able to detect when alterations in       a diet containing 3300 kcal of DE in order to meet
feed intake occur and take steps to intervene when         its energy requirements. Unfortunately, our modern
reductions occur.                                          genotypes of sows will often not consume this
                                                           amount of food. As a consequence, such sows with
                                                           a high level of productivity may not consume
Nutrients Required During Lactation
                                                           sufficient feed to meet their energy requirements
Energy
                                                           and will have to "milk off their backs" in order to
  The daily energy requirement of the sow during
                                                           meet the energy needs of the nursing litter.
lactation includes a requirement for maintenance
and a requirement for milk production. The mainte-
nance energy requirement of the lactating sow is
assumed to be the same as that for the gestating sow
and is estimated to be I IO kcal of DE per kg of
body weight 15• This is sometimes a difficult calcu-
                                                         158
Table 6-28. Estimated Daily Energy Requirements of Lactating Sows.
Sow Body Weight (kg)                                                                         142.5               142.5               217.5             217.5       280     280
Number of Suckling Piglets                                                                     8                   12                  8                 12         8       12
Daily Gain of Piglets (g/day)                                                                 200                 240                 200               240        200     240
Estimated Milk Yield (kg/day)'                                                                     6.4            11.5                6.4               11.5        6.4     11.5
Energy for Maintenance (Mcal/day)2                                                                4.53            4.53                6.23              6.23        7.52    7.52
Energy for Milk Production (Meal/day)?                                                           12.80           23.00               12.80             23.00       12.80   23.00
Daily Energy Requirement (Meal/day)                                                              17.33           27.53               19.03             29.23       20.32   30.52
Daily Required Feed Intake (kg/day)'                                                              5.25            8.34                5.76              8.85        6.15    9.25
1
 Milk yield estimated from litter gain/day x 4
2
  Sow energy needs for maintenance estimated from BW75 x I IO kcal per kg (see Table 6-10)
3
  Energy for milk production estimated from milk yield x 2 MCal/kg milk produced
"Feed intake required based on 3300 kcal DE/kg diet
  Using the above example, assume that the sow is                                                            Amino Acids
only consuming 6.5 kg of feed per day. In this                                                                 The amino acid requirements of lactating sows
situation, the sow will have an energy deficit of                                                            have been the subject of a considerable amount of
6006 kcal/day ( 1.82 kg x 3300 kcal). It has been                                                            research during the past decade and it is now
calculated that for every kg of weight loss by the                                                           evident that the current estimates of requirements in
sow, 11,200 kcal of DE equivalent are released.                                                              the NRC tables are inadequate to allow high pro-
Therefore, to make up the 6006 kcal/day deficit will                                                         ducing sows to perform to their genetic potential.
require a maternal weight loss of 536 g/day or about                                                         This is likely a reflection of improvements in sow
15 kg during a 28 day lactation. As a rule, it is                                                            milking ability due to increased selection pressure
desireable to keep maternal weight loss during a 28                                                          on this trait by breeding companies.
day lactation to less than 10 kg and as a
consequence, steps would have to be taken to                                                                   An estimate of the amino acids needs of the
increase lactational energy intake by the sow or                                                             lactating sow can be obtained by examining the
reduce the energy demands for lactation through                                                              amino acid composition of sow's milk. By using
cross fostering.                                                                                             the daily milk yield of the sow (based on piglet
                                                                       LACTATION FEED
                                                                       INTAKE RECORD
                        SOWID:                /Q,J(i                 PARITY:             _!/___ DATE:Farrowed'#s/O�/Q/               Weaned��
                                                        l2.
                                                                                . . .Ll .. . . . . •� ..;-r...• L::.....,. ...._..,, ..., . �. ..::,.1
                                                                      w.�                                                        �I Per day             7.3/r...
                        .. I .                           . .
                        NO: Born Alive                                                                      FEED INTAKE: To1al
                        ...
                            I                                                   ti
                                                                                     I  .. I ;
                                                                                             I
                                                                                                 -
                         .. II I                          t/1          ti                        i
                                                                                                            l
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                                                                ti
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                       ..... ,
                       I'll"                                                         I                       I
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                            I 2 J
                                     '"s s
                       u, 3.l!_'' .!J� ,, , 'l•l4• 't' 't' ,, "P "'
                                                                                                 �SS5'.                              S" (' S"            ,_ ,_
                         M
                                                                                                                                                   I-
                        .. ii I I 1 J "1¥                           rs r r r r           f
                                                                 ,,
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                                                                                                                                                         ,_.._
                                                                                                          159
weight gain shown above) one can calculate the                  The most important animo acid for sows in lacta-
amounts of the essential amino acids which are               tion is lysine. Dividing the daily lysine requirement
secreted in milk on a daily basis. The efficiency of         by the expected feed intake of the sows will show the
utilization of amino acids for milk production has           required level of lysine in the diet. For example, a
been estimated at 80% and most lactation diets                142.5 kg sow with a litter gaining 2250 g/day (i.e.,
would have a digestibility coefficient of                    producing 9 kg of milk per day) will require 55.9 g of
approximately 80%. Therefore, by dividing the                lysine per day. If the sow is consuming 7 kg of feed,
amounts of amino acids secreted into milk by 0.64            then the diet will need to provide 0.79% lysine (55.9
(80% efficiency x 80% digestibility), one arrives at         g + 7000 g = 0.79%).
the total amount of amino acids required to support
this level of milk production. Examples are shown              Obviously, it is not possible to formulate one diet
in Table 6-29.                                               that will meet the lysine requirements for each and
                                                             every sow in a herd. In addition, since the objective
  The total dietary amino acid needs of the lactating        oflactation feeding is to maximize feed intake, vary-
sow can be determined by adding the amino acids              ing the level of feed provided is not available as a
required for maintenance (see table 6-11) to those           means of varying the lysine intake of individual sows.
need for lactation (Table 6-29). Examples are given          Under practical feeding conditions, it is suggested that
for three weights of sows and four daily milk yields         producers formulate two rations to be fed during lac-
but can be calculated for any situation using the            tation. One to be fed to first and second parity sows
figures provided.                                            (these sows typically cannot consume adequate feed
                                                             and therefore this diet would be formulated with higher
                                                             lysine levels) and one for the remainder of the herd.
Table 6-29. Estimated Daily Amino Acid Requirements for Milk Production by the Sow (g/day).
              Composition
              of Milk (%)1                               Daily Milk Production (kg/day)2
                                         5                      7                   9                     11
                                  S3         R4          s          R         s         R           s          R
Arginine        5.1               12.7       19.9      17.8         27.9     22.9       35.8      28.0      43.8
Histidine       2.8                7.0       10.9       9.8         15.3     12.6       19.7      15.4      24.1
Isoleucine      4.5               11.3       17.6      15.7         24.6     20.2       31.6      24.7      38.7
Leucine         8.7               21.7       34.0      30.4         47.6     39.1       61.2      47.8      74.8
Lysine          7.8               19.5       30.5      27.3         42.6     35.1       54.8      42.9      67.0
Sulfur AA       3.3                8.3       12.9      11.5         18.0     14.8       23.2      18.1      28.4
Aromatic        8.3               20.7       32.4      29.0         45.4     37.3       58.4      45.6      71.3
Threonine       4.2               10.5       16.4      14.7         22.9     18.9       29.5      23.1      36. l
Tryptophan      1.2                3.0        4.7       4.2          6.6      5.4        8.4       6.6      10.3
Valine          5.6               14.0       21.9      19.6         30.6     25.2       39.4      30.8      48.1
I
  Adapted from Speer ( 1990). Partitioning nitrogen and amino acids for pregnancy and lactation in swine: A review. J.
Anim. Sci. 68: 553-56 l.
2
  Daily milk production calculated by multiplying daily weight gain of litter x 4.
3Secreted
"Required
Based on sow producing milk with 5% protein. Assumed absorbed amino acids used at an efficiency of 80% for
milk production and diet had 80% digestibility.
                                                          160
Table 6-30. Estimated Total Daily Amino Acid Requirements of Lactating Sows (g/day).
Arginine                   19.9   27.9    35.8    43.8      19.9     27.9   35.8   43.8      19.9   27.9   35.8   43.8     21.2
Histidine                  10.9   15.3    19.7    24.1      10.9     15.3   19.7   24.1      10.9   15.3   19.7   24.1     13.2
Isolcucine                 18.8   25.8    32.9    39.9      19.3     26.3   33.3   40.4      19.6   26.6   33.7   40.7     20.7
Leu cine                   34.8   48.4    61.9    75.6      35.1     48.7   62.3   75.9      35.4   48.9   62.5   76.1     25.4
Lysine                     31.5   43.7    55.9    68.0      31.9     44.0   56.3   68.4      32.2   44.4   56.6   68.7     31.8
Sulfur Amino Acids         13.9   19.1   33.7     29.4      14.4     19.5   24.7   29.8      14.7   19.8   24.9   30.I     19.1
Aromatic                   34.3   47.3   60.2     73.2      35.0     47.9   60.9   73.9      35.6   48.5   61.5   74.5     37.1
Threonine                  18.0   24.6   31.1     37.7      18.6     25.2   31.7   38.3      19.1   25.6   32.2   38.7     22.8
Tryptophan                  4.9    6.8    8.6     10.5       4.9      6.8    8.7   10.6       5.0    6.9    8.8   10.6      6.4
Valine                     22.7   31.5   40.2    48.99      23.0     31.8   40.6   49.3      23.3   32.0   40.8   49.5     31.8
I
    Based on a 165 kg sow consuming 689 g/day crude protein.
Vitamins and Minerals                                              will dry the sow off more rapidly, leading to a shorter
  There has been surprisingly little research conducted            interval between weaning and rebreeding. However,
on the vitamin and mineral requirements of sows dur-               others feel that moderate to high levels of feed will
ing lactation alone. The available data suggests that              give the best results.
the lactating sows requirements do not differ appreci-
ably from those of the gestating sow and therefore                   A Texas study investigated the effects of feed and
the nutrient levels suggested in Tables 6-12 and 6-13              water deprivation prior to weaning, on the number of
are recommended.                                                   days to successful service. The results of this trial
                                                                   indicate that feed and water deprivation at weaning
Feeding the Sow at Weaning                                         prolonged the weaning to service interval (Table 6-
  There is a great diversity of opinion as to how sows             31 ). Therefore, it would appear that the most effec-
should be fed and managed from weaning to                          tive way to dry off a sow is to maintain them on a
rebreeding. Many producers believe that withhold-                  moderate level of feed and allow milk to accumulate
ing feed and water for 24 to 48 hours after weaning                in the udder. The resulting increase in intramammary
                                                                   pressure will very effectively and rapidly stop milk
                                                                   secretion and stimulate a rapid return to estrus.
Table 6-31. Effect of Feed and Water Deprevation, Prior to Weaning, on Days to Successful Service After
Weaning.
                                                                                          Hours of Deprivation
                                                                                           Prior to Weaning
Feed                                                                    0                 48.0              0             48.0
Water                                                                   0                  0               24.0           24.0
Days to Successful Service                                              6.0               11.9              9.1           12.8
Number not Showing Estrus Within
21 Days Postweaning                                                     0.0                5.0              3.0            5.0
Orr et al., 1981, Anim. Prod. 15: 259-264.
                                                               161
Feeding the Sow Between Weaning                              reasonable condition to achieve high conception
                                                             rates and good litter size.
and Rebreeding
   The level of feed intake selected from weaning
                                                                 During hot weather, conception rates are often
and mating should allow the release of sufficient
                                                             reduced. There is some evidence to show that
ova, their successful fertilization and implantation.
                                                             feeding vitamin fortified diets to the breeding herd
Many producers find that gilts which have just
                                                             may improve conception rates during periods of
weaned their first litter are difTicult to rebreed. This
                                                             elevated temperatures. In a recent study, sows were
is generally due to the poor body condition of first
                                                             fed higher than recommended levels of vitamin A
litter sows and the stress of competing with older
                                                             ( I 50% NRC), vitamin B-12, pantothenic acid and
sows after weaning. The best solution is to feed
                                                             choline during periods of hot weather. The concep-
sows better during lactation than to try to catch up
                                                             tion rate of the vitamin fortified group increased
later on. However, supplying high levels (3.5 - 4.0
                                                             from 52.8% to 76.0%.
kg/day) of feed to gilts in poor condition from
weaning to rebreeding will improve conception
rates and reduce the number of days to rebreeding
                                                             Feeding Boars
(Table 6-32). After breeding, feed levels should be          Introduction
reduced to maintenance levels in order to prevent a            The boar is often the forgotten member of the
high incidence of embryo mortality.                          breeding herd and there has been surprisingly little
                                                             research conducted to determine their nutritional
                                                             requirements. Most tables of nutrient requirements
Table 6-32. The Effect of Feed Intake During the             group boars together with dry sows and assume that
Weaning to Remaining Interval on the Reproductive            a diet that meets the requirements of sows during
Performance of Gilts and Sows.                               gestation will also be satisfactory for the herd sire.
                                Feed Intake(k;g/day)
                                                             Nutrition of Boars During Rearing
                                    1.8    2.7   3.6         (20 -100 kg)
                                                                 Young boars are usually selected to be herd sires
Weaning/Estrus                                               according to an index which includes such charac-
Interval (days)             Gilts   21.6 12.0 9.3            teristics as growth rate. appetite, feed efficiency,
                            Sows     4.9 4.7 5.0             lean tissue growth rate and carcass quality. As a
Conception Rate (%)         Gilts     58 75 100              consequence, young boars need to be fed high-
                            Sows     100 87 100              energy and nutrient dense diets ad libitum in order
Subsequent Litter Size      Gilts    9.4 10.1 11.6           to allow for the accumulation of meaningful per-
                            Sows    12.6 11.8 12.2           formance data for use in selection programs. Dur-
                                                             ing the period from birth to 50 kg body weight, the
Adapted from: Brooks and Cole, 1972, Anim. Prod. 15:
259-264, and Brooks et al., 1975, Anim. Prod. 20: 407-
                                                             boars potential for lean tissue growth appears to be
412.                                                         beyond the upper limit of appetite. This means that
                                                             high energy diets can be fed ad libitum without
                                                             excessive fat deposition or decline in feed effi-
   Flushing does not work with older sows. The               ciency and energy levels of at least 14 MJ DE/kg
purpose of a high level of feeding after weaning is          (3344 kca1/kg) diet are recommended. From 50 to
to increase ovulation rates and thereby increase the          I 00 kg liveweight, the boars potential for protein
subsequent litter size. However, since ovulation             deposition lies within the limits of appetite and
rate is not usually a limiting factor for sows, even if      excess energy levels may affect performance.
the ovulation rate is increased, an improvement in           Australian data indicate that the maximum rate of
litter size is not usually forthcoming. A daily feed         protein deposition occurs with energy levels of 33
intake of approximately 2.7 kg between weaning               MJ DE/day (7883 kcal/day). If the potential ad
and remating is generally sufficient to allow sows in        libitum feed intake of the boar is known, then the
                                                           162
required energy content of the diet can be calculated      Table 6-33. Nutrient Requirements of Developing
by dividing this energy level by the potential feed        Boars (20-100 kg).
intake.
                                                                                         Weight of Boar
  The protein requirement of growing boars is                                         20-55 55-100        120-
greater than that of barrows or gilts since boars gain                                                  Breeding
faster, are more efficient and have less backfat.
Average daily gain and feed efficiency are maxi-           Energy (MJ/day)          29          33         25
mized for growing boars at levels of 20% protein           Protein(%)               20           18        14
during the growing period (20-55 kg) and 18%               Lysine(%)                1.0         0.8      0.65
crude protein during the finishing period (55-100          Tryptophan (%)          0.17        0.14      0.12
kg). In addition, developing boars require 0.15% to        Threonine(%)            0.65        0.60      0.50
0.25% more lysine than barrows. Data from the              Methionine+ Cystine (%) 0.45        0.35      0.40
University of Kentucky indicate that the lysine            Calcium(%)              0.75        0.70      0.65
requirement of 35-60 kg boars is 0.86% while that          Phosphorus(%)           0.60        0.55      0.50
of 60-100 kg boars is 0.74%. These levels are              Salt(%)                 0.25        0.25      0.50
higher than those recommended by the NRC.                  Zinc (mg/kg)             75          50         75
                                                           Iron (mg/kg)             75          50         75
  The developing boar may require higher levels of         Manganese (mg/kg)        20          20         20
calcium and phosphorus than the levels fed to              Copper (mg/kg)             6           5         6
market animals since the tendency to show leg              Selenium (mg/kg)        0.15        O.IO      0.10
weakness may be exacerbated by high growth rates           Vitamin A (lU/kg)       3000        2000      4000
and by feeding to appetite. Calcium and phospbo-           Vitamin D (ill/kg)       300         200       400
ms levels of0.75% and 0.60% respectively are               Vitamin E (IU/kg)          18           15       20
recommended during the growing period and drop             Vitamin K (mg/kg)         l.5         1.0       2.0
to 0.65% and 0.50% by the time the boar is ready           Riboflavin (mg/kg)       4.0          3.0       5.0
for breeding. A summary of nutrient requirements           Niacin (mg/kg)            30          25        35
for developing boars is presented in Table 6-33.           Pantothenic Acid (mg/kg) 15            12       20
                                                           Choline (mg/kg)          400         300       500
   The experimental evidence suggests that nutrition       Vitamin B12 (ug/kg)       15           10       20
during rearing can influence both the age of puberty       Biotin (ug/kg)            50          50       100
and the rate of sexual development but these are
                                                           Adapted: Wahlstrom, 1991. Swine Nutrition pp 517-
unlikely to be impaired under current feeding
                                                           526.
regimes and practices. Under normal breeding
conditions, boars attain puberty between 5 and 8
months of age when they weigh 80-120 kg body               Nutrition of the Working Boar
weight. Age is more important than body weight in
                                                             The daily energy requirements for working boars
determining the onset of puberty. Although a 30%
                                                           can be assessed as the summation of the following
reduction in feed intake has been shown to cause a
                                                           components: maintenance, body gain, semen
42 and 30 day delay in puberty for purebred and
                                                           production, mating activity and the requirement for
crossbred boars. unless severely undernourished,
                                                           extra heal production when kept below their lower
this restriction does not appear to impose any long
                                                           critical temperature. Estimates have been derived
lasting, damaging effects upon reproductive capac-
                                                           from the factorial principle of nutrient partitioning
ity other than the obvious effects on the growth and
                                                           and range from 29 to 41.5 MJ DE/day with an
body size of the animal. In practise, most young
                                                           additional 3% for each I °C decrease in temperature
boars are offered feed to appetite during rearing and
                                                           below an environmental temperature of20°C.
at this feeding level there is unlikely to be any
negative effects upon sexual development or subse-
quent reproductive capacity.
                                                         163
  The energy requirements for maintenance can be            The results of these calculations indicate that
calculated in a similar manner to those of the            working boars should be fed between 2.6 and 3.0
gestating sow using a value of .4 LS MJ DE per kg         kg per day. The exact amount given to the boar will
body weight:". The energy requirements for growth         depend on breeding load, climatic conditions, body
are more difficult to calculate because there has         weight and condition. Boars should be individually
been very little research conducted to determine the      fed, twice a day. This allows for a daily check of
optimum growth rate of working boars.                     the health and vigor of the boar and an adjustment
Overfeeding is undesireable because overfeeding           in the amount of feed provided where necessary.
may reduce I ibido and may increase the risk of leg       Fresh water should be provided at all times.
weakness. Ln addition, overfeeding can increase the
size and weight of the boar to the point where the           Several studies have indicated the importance of
boar is no longer compatible with the sows in the         protein and more specificaUy the amino acids
herd and must be culled. Given their high purchase        lysine, methionine and cystine on the number of
price, early culling of boars is clearly detrimental.     sperm cells produced. There appears to be a mini-
However, underfeeding may affect reproductive             mum level of protein neccessary to ensure maximal
characteristics such as number of sperm cells and         sperm production since low protein levels have
the fertilizing capacity of the sperm cells.              been shown to reduce the number of sperm cells
                                                          ejaculated. A crude protein content of 14% with
  One of the few attempts to suggest an optimum           lysine and sulfur containing amino acid levels of
growth rate for working boars comes from recent           0.65% and 0.44% are recommended.
research at the University ofWageningen. Dutch
workers recommend a moderate growth rate (400 gJ             Calcium and phosphorus are the most important
day) for young boars (150-250 kg) and a reduced           minerals to consider in the context of the nutrition
weight gain (200 g/day) in mature boars (250-400          of the working boar as they are crucial not only to
kg). Their research suggests an energy cost for           optimal growth rate but also to bone mineralization
growth of 32.8 MJ per kg weight gain. The energy          and hence overall soundness of the limbs. It is
requirements for reproduction (semen production           generally accepted that higher levels of calcium and
and mating activity) amount to about 18 kJ per kg:"       phosphorus are required for optimum bone miner-
which amounts to less than 3% of the maintenance          alization as opposed to maximizing growth rate.
cost of the boar. Therefore, this energy cost is          Soundness oflimbs is an important index to con-
usually ignored in calculating the daily energy needs     sider as foot-related problems are a major contribu-
of the working boar. The energy requirements of           tory factor to loss of libido and the inability of the
boars of various body weights are calculated in           boar to mount a sow. It has been reported that
Table 6-34.                                               boars which were fed 150% of NRC recommenda-
                                                        164
tions for dietary calcium and phosphorus and                fibrous foods. For example lesions of the
thicker-walled metacarpals which had greater                esophagus are common in cereal fed animals
strength than boars fed I 00% of recommendations.           resulting in a reduced feed intake and poorer repro-
                                                            ductive performance. Increasing the fibre content
  Zinc has an established role in spermatogenesis           of the diet has been shown to reduce the severity of
since zinc deficiencies are implicated in the retarda-      such lesions.
tion of the leydig cells. a reduced response to
leutinizing hormone and a reduction in testicular              In summary, the energy and protein needs of a
steroidogenesis. A level of I 00 mg/kg is recom-             typical boar (i.e., one of 18-24 months and 175-250
mended. Boars show no additional requirements                kg liveweight maintained in a thermoneutral envi-
for the other major minerals in relation to sows.            ronment) are satisfied by a daily feed allowance of
                                                             2.6-3.0 kg of a diet containing around 13 MJ DE/kg
  It has also been suggested that the working boar           and 140 -160 kg of crude protein. For every 1°C
has no additional requirement for vitamins above             that the house temperature drops below 21 °C a boar
those of the breeding sow. However, the role of              on a solid unbedded floor should be offered an
biotin in the diet of boars is becoming increasing           extra I OOg of food daily. For a boar well bedded
important as a result of its association with foot           on straw, this rule would apply below 17°C.
lesions and the attendant implications this bas for
reproductive performance in the boar. Biotin is an
                                                             Table 6-35. Composition of Diets for Adult Boars.
essential water soluble sulfur containing vitamin.
Supplementation of the diet with biotin has been                                          150-200 kg 200-350kg
shown to significantly strengthen the hoof. The
precise mode of action of biotin in the prevention of        Dietary Energy, MJ DE/kg         13.0         13.0
foot lesions is still uncertain. It is known that biotin     Protein,%                        15.0         14.0
increases the compressive strength and hardness of           Lysine,%                          0.7         0.55
the hoof wall which decreasing the hardness of the           T.S.A.A., %                      0.47         0.40
heel bulb tissue. A soft heel bulb presumably acts           Calcium,%                        0.80         0.75
as a cushion, minimizing stresses and absorbing              Phosphorus, %                    0.70         0.60
strain energy. There is also the suggestion that
vitamins E and C may be of special importance in             Adapted from Aherne, F. 1995. Pig Letter 15:16.
the diet of stress susceptible breeds.
                                                           165
Additional Reading and References                           Kemp, B. and den Hartog, L.A., 1989. The influ-
Aherne, F.X., 1991. Feeding the gilt and sow for            ence of energy and protein intake on the reproduc-
optimal production. Pages 455-473 ln J.M. Naylor            tive performance of the breeding boar: A review.
and S.L. Ralston eds. Large Animal Clinical                 Anim. Reprod. Sci. 20: 103-115.
Nutrition, Mosby Year Book, St Louis, Missouri.
                                                            Kemp, 8., den Hartog, L.A. and Grooten, H.J.G.,
Azain, M.J., 1993. Effects of adding medium-chain            1989. The effect of feeding level on semen quan-
triglycerides to sow diets during late gestation and        tity and quality of breeding boars. Anim. Reprod.
early lactation on litter performance. J. Anim. Sci.        Sci. 20: 245-254.
71: 3011-3019.
                                                            Mahan, D.C., 1990. Mineral nutrition of the sow:
Brooks, P.H. and Cole, D.J., 1972. Studies in sow           A review. J. Anim. Sci. 68: 573-582.
reproduction. I. The effect of nutrition between
weaning and remating on the reproductive perform-           National Research Council, 1988. Nutrient Require-
ance of primiparous sows. Anim. Prod. 15: 259-              ments of Swine. 9th Revised Edition, National Acad-
264.                                                        emy Press, Washington, D.C.
Brooks, P.H., Cole, D.J., Rawlinson, P., Croxson,       Noblet, J., Dourmad, J. Y. and Etienne, M., 1990.
V.S. and Luscombe, J.R., 1975. Studies in sow           Energy utiliztion in pregnant and lactating sows:
reproduction. 3. The effect of nutrition between        Modelling of energy requirements. J. Anim. Sci. 68:
weaning and remating on the reproductive perform-       562-572.
ance ofmultiparous sows. Anim. Prod. 20: 407-
412.                                                    Pettigrew, J.E., 1993. Amino acid nutrition of ges-
                                                        tating and lactating sows. Biokyowa Technical Re-
Close, W.H. and Roberts, F.G., 1993. Nutrition of       view, 18 pp.
the working boar. Pages 347-371. In D.J.A. Cole,
W Haresign and P.C. Garnsworthy, eds. Recent            Speer, V.C., 1990. Partitioning nitrogen and amino
Developments in Pig Nutrition 2, Nottingham             acids for pregnacy and lactation in swine: A review.
Univeristy Press, Sutton Bonington, UK.                 J. Anim. Sci. 68: 553-561.
den Hartog, L.A., Backus, G.B.C. and Vermeer,           Standing Committee on Agriculture, 1987. Feeding
H.M., 1993. Evaluation ofhousing systems for            Standards for Australian Livestock: Pigs. CSIRO
sows. J. Anim. Sci. 71: 1339-1344.                      Printing Centre, Collingwood, Victoria, 226 pp.
Froese, C., 1987. Assessing sow body condition.         Takach, M.D., and Dial, G.D., 1992. Managing the
Manitoba Agric, Swine Fact Agdex No. 440-22-40.         lactating sow for optimal weaning and rebreeding per-
                                                        formance. Pages 559-573. In R.C. Tubbs and A.O.
Hoofs, I.A., 1990. Group housing of pregnant sows       Leman eds. The Veterinary Clinics ofNorthAmerica:
in the Netherlands. D. Kellliher, B. Lynch, M.          Swine Reproduction. W.B. Saunders Company, Phila-
Martin, F. Tobin and N. Walsh eds. Pig Production       delphia.
Trends for the l 990's.Proceedings of the 19th
Annual Syrnp., University College, Dublin, Ireland.     Wahlstrom, R.C., 1991. Feeding developing gilts
pp. 14-23.                                              and boars. Pages 517-526 in E.R. Miller, D.E.
                                                        Ullrey and A.J. Lewis eds. Swine Nutrition.
Kemp, B., Bakker, C.M., den Hartog, L.A. and            Butterworth-Heinemann, Stoneham Mass.
Yerstcgcn, M. W., 1991. The effect of semen
collection frequency and food intake on semen
production in breeding boars. Anim. Prod. 52: 355-
360.
                                                      166
7. FEEDING THE SUCKLING PIG
    Young piglets have a great capacity for growth,        Photo 7-1.
much greater in fact, than is commonly recognized.
Growth in the suckling piglet is primarily limited by
insufficient intake of milk or of alternative sources
of nutrients. Because practical approaches for
transferring more nutrients from the sow to the
piglet are not yet available. it is the producer's job
to get more nutrients into the piglet so that growth
can be maximized.
                                                          167
   To ensure all piglets acquire colostrum,                   Photo 7-2.
producers should take the time to observe litters
shortly after farrowing and assist weak piglets to
suckle by giving them access to the sow's udder.
One way to give access is called "split suckling''.
Shortly after birth, half the piglets are removed
from the sow and are kept in a warm, dry box. The
two halves of the litter are rotated on and off the
sow to give individual piglets maximum
opportunity to suckle and receive colostrum.
   Another way to ensure piglets receive some                 Iron injections should be administered in the neck to
colostrum is to keep a supply of cow colostrum in             avoid staining the ham at the injection site.
the freezer and give weak and unthrifty piglets an
oral dose (using a small syringe) if they have had
inadequate suckling of their mother's colostrum.              Although iron supplementation is crucial to the
                                                          suckling piglet's we11-being, excess iron may
The Need for Supplementary Iron                           increase the incidence of scouring and susceptibility
    Based on nutrient content and availability, sows'     to bacterial infections. Iron is generally found
milk is generally accepted as the nearly perfect food     tightly bound to protein in the suckling pig's body.
for the very young piglet. The only nutrient that is      This binding limits how much iron is accessible to
present in sub-optimal levels is iron. Because of         bacteria. However, if iron is present at a level
this deficiency, piglets require supplementary iron       beyond the capacity of these proteins to bind it,
or they will become anemic and die.                       pathogenic bacteria may proliferate. Therefore,
                                                          take care to ensure that piglets are provided with
    The iron deficiency in sow's milk can be over-        adequate, but not excessive, iron supplement. Lron
come by the administration of iron dextran either         injections occasionally result in the rapid death of
orally or by an intramuscular injection. The amount       some piglets. which may indicate a vitamin E and
of iron required to meet their requirements largely       selenium deficiency.
depends on the age at which the piglets are to be
weaned. Researchers have calculated that piglets              Water Requirements
require approximately 10 mg of iron per day to                Piglets require water immediately after birth and
allow growth at their maximum rate. Therefore, if         should be offered water from their first day,
piglets are to be weaned at three weeks, a single         especially in warm conditions. Providing drinking
injection of 150-200 mg of iron dextran is suffi-         water to very young piglets does not reduce their
cient. However, if piglets are to be weaned later, a      motivation to suckle; rather, they benefit from
booster shot may be required between days 14 and          supplemental drinking water, especially if the sow
21. A means to check the adequacy of iron injec-          milks poorly and they do not consume enough
tions is to monitor haemoglobin levels in the blood       water in the limited amount of milk they receive.
of the young piglets. The levels should be higher         Litters that gain poorly during the first one to three
than 8 ml/I OOml. Iron injections should be admin-        days after farrowing drink more water than faster
istered in the neck rather than the ham to avoid          growing litters. Provision of a fresh water source is
staining of the most valuable cut in the pig carcass.     also beneficial in reducing stress that piglets
                                                        168
undergo at weaning. Pigs accustomed to drinking                     Artificially reared piglets should be fed milk
from a watering system prior to weaning will                     replacer frequently (4 to 6 times daily) and in small
experience fewer problems with water intake after                doses ( 10 ml of milk replacer per pig at first).
weaning. Make the water source easy for newborn                  Daily consumption of milk replacer may be
piglets to find. Piglets take too long to find and use           gradually increased to 300 to 400 ml per piglet at
a nipple drinker so a dish drinker located in the                which time dry feed should be introduced. Prevent
creep area is a better choice, at least until the piglets        over-consumption of milk replacer and take care to
are accustomed to consuming water. If water                      ensure the piglets do not get wet during feeding.
nipples are to be used, then these should be pointed
downward to discourage piglets from playing with                    The successful use of milk replacers to reduce
the nipple and wasting water.                                    pre-weaning mortality in runt pigs has prompted the
                                                                 development of several mechanical devices or
Use of Milk Replacers                                            "artificial sows" which are designed to automate the
    Milk replacers have been used quite successfully             delivery of milk replacers. The manufacturers of
to raise orphan piglets or surplus piglets from large            these devices claim that a piglet can be fostered
litters. Milk replacer may also be offered to piglets            onto an artificial sow as early as 3 days of age.
while they are with the sow in the farrowing crate to
increase weaning weights and reduce variation in                    The need and use of milk replacers in the
weaning weight in early weaning programs                         commercial swine industry is anticipated to
(weaning age 21 days or less). A point to                        decrease as more complex creep and early weaning
remember: if milk replacers are used, piglets must               feeds are developed. Some of the early weaning
still obtain colostrum from the sow or another                   dry feeds can support good levels of performance in
source, such as cow colostrum.                                   piglets weaned at 7 days or earlier (see chapter 8).
                                                                 Creep Feeding
Photo 7-3a.                                                          The sow generally reaches her maximum milk
                                                                 production during the third week oflactation;
Photo 7-3b.
                                                                     .
                                                                           8
                                                                     "ti
                                                                           ..
                                                                     >
                                                                     �2
                                                                     aJ
                                                                     0
                                                                           O+---,.�.---.---.�...-������---��
                                                                                0   2           4           6       8
                                                                                        Week of Lactation
Piglets should have ready access to water.
Photo courtesy: Centre for Food and Animal Research. Ottawa.     Figure 7-1. A lactation Curve for a SO\\..
                                     -                     -
                                                               169
Benefits of Creep Feeding                                    quality diet. Such a diet would contain at least
   The benefits derived from creep feeding have               l.25% (up to 1.70%) lysine and 3400 - 3600 kcal
been amply demonstrated. The most significant                DE/kg. This was demonstrated by recent studies
benefit is that piglets are heavier at weaning. In a         conducted at the Agriculture Canada research
typical four week weaning program, weaning                   station in Ottawa. In these studies, piglets were
weights may be as much as I 0% higher if high                weaned at four weeks of age. Litters on high
quality creep feeds are used. For piglets weaned at          complexity, palatable diets ate more creep feed
an earlier age, the intake of creep feed is generally        prior to weaning, tended to gain more during the
insufficient to stimulate heavier weaning weights.           week before weaning, and converted feed more
                                                             efficiently. These piglets also gained more weight
   Tn addition, the variation in weaning weights             in the first two weeks after weaning as compared to
within a litter may be smaller, i.e., fewer runt pigs in     litters that were fed a low-complexity, corn-soybean
the litters at weaning. The response to creep feed           meal based diet.
will also depend on litter size, the individual sow's
milking ability and the quality of creep feed.                   During the first two or three weeks of life, up to
                                                             approximately five kilograms body weight, a
    Creep feeding can also minimize the setback that         piglet's digestive enzymes necessary for the diges-
piglets undergo immediately following weaning.               tion of starch, sugar (sucrose), and non-milk pro-
An abrupt change from a liquid diet to a meal diet is        teins are present at relatively low levels. A piglet's
a nutritional stress that generally results in a growth      digestive tract is best suited to digest milk proteins
check during which time pigs consume very little             (casein), milk sugar (lactose), glucose, and specific
feed, gain little weight, and often suffer from              fats so its diet should include ingredients such as
scours. A gradual introduction to solid feed during          skim milk. high quality whey products, high quality
the suckling period will develop the enzymes                 blood products, added fat (preferably fats with
required for digestion, allow piglets to learn how to        medium chain fatty acids, such as coconut oil, or
consume dry feed, and minimize the growth check.             unsaturated fats, such as com oil, canola oil, or soy
However, in order to achieve a reduction in setback          oil), cooked cereals (rolled oats or flaked corn), and
after weaning through enhanced creep feed con-               glucose. In addition, small amounts of non-milk
surnption, substantial amounts of creep feed must            proteins should be added to encourage development
be consumed. According to some studies, this may             of the piglet's digestive enzymes. High quality pre-
be as much as one kilogram per piglet.                       starter (Phase T or MEW) diets may be used as
                                                             creep feeds for suckling piglets weighing less than
    Creep feeding has also been shown to be                  five kilograms (see chapter 8).
beneficial to the sow. When creep feed is provided
piglets tend to suckle less intensively so sows tend
to be in better body condition at weaning. As a
result, these sows have shorter weaning to                              a
rebreeding intervals and higher conception rates                 >      7
than sows whose litters have not been provided with              >
                                                                 ....   6
                                                                                            ., ,
creep feed.                                                      fi     5        � ......
                                                                 <      4
                                                                 w
Creep Feed Formulation                                           �      '2
                                                                                                                                 La-(·-
                                                                 �
    Creep feeds are designed to supplement the                   w
nutrients contained in sows milk. Therefore, creep                           L-......,..-.....-�..----...-----.--,                  ...,,....ug,u,
feed does not have to provide all the nutrients                              t                     2     3    4      5   6   7
                                                           170
    At approximately three weeks of age, or when               before being cleaned and/or replaced. Little and
body weight exceeds five kilograms, this largely               often is the rule for creep feeding, with fresh creep
milk-based diet can be replaced by a less expensive,           feed supplied each day. Not only does this practice
pre-starter diet based on cereals and high quality             ensure the creep is always fresh, but the daily arrival
proteins. In general, high energy grains such as               of fresh feed serves to stimulate the piglets' inherent
corn or wheat as well as readily digested protein              curiosity in the new material, which also helps to
supplements such as soybean meal form the basis of             encourage consumption.
most pre-starter diets. The pre-starter diet should
still contain some of the high quality products that               Consumption can be stimulated by feeding a
are included in creep feeds. Ingredients such as oat           pelleted or crumbled creep feed. Piglets fed pellets
groats, flaked corn. fish meal. dried skim milk,               with a small diameter (0.4cm or smaller) or crum-
dried whey, sugar, and fat are commonly added to               bles tend to eat more and waste less as compared to
increase palatability and digestibility. Pre-starter           piglets fed pellets with a large diameter (0.5cm or
(Phase I or high quality Phase II) diets may be used           larger). If small pellets or crumbles are not avail-
as creep feeds for suckling pigs weighing more than            able, a short-cut pellet (2 - 3 mm long) is preferred.
five kilograms as well (see chapter 8).
                                                                   Even though piglets have access to the sow's
Factors Affecting Creep Feed Intake                            milk while eating creep feed, they must have access
   Creep feed consumption is affected by many                  to fresh water if feed intake is to be maximized. If
factors. These factors include: the feeding system,            pigs are weaned after three to four weeks of age,
feed palatability, the environment in which the                creep feed intake will be reduced significantly
piglet is housed the sow's milk composition and                without readily available fresh water. Pigs accus-
yield, the piglets' health, vigor, and growth rate, and        tomed to drinking from a nipple waterer while in
the amount of water available. A reasonable target             the farrowing crate will experience fewer problems
creep feed intake for litters weaned at 4 weeks is             in adapting to the watering system at weaning.
about 2.5 kg of feed per litter. If litters are not            However, water nipples should be pointed down-
achieving this level of intake, take steps to increase         ward to discourage piglets from playing with the
their creep feed consumption.                                  nipple and wasting water.
                                                             171
Starting Pigs on Creep Feed                                    cases, the hypersensitivity can result in an immune
    Piglets should be started on creep feed when               response in the gut, damaging the intestinal tract
they are about 7 days old by offering a little feed on         lining The damage can cause diarrhea in the newly
a clean, dry section of the floor (use a board in              weaned pig, and poor post-weaning performance,
farrowing crates with a slatted floor). Piglets are            However, if creep feed is introduced early and large
more likely to show interest in feed initially offered         amounts are consumed prior to weaning, the im-
in this manner. Never offer creep feed within a                mune system may develop a tolerance to feed
couple of hours of the sow being fed; the litter will          antigens prior to weaning, and post-weaning scour-
be suckling or sleeping and may not notice the feed.           ing can be controlled. The effect of feeding strate-
ln addition, never offer more than 20 g/litter/day             gies on changes in the piglet's immune system and
until the piglets are obviously consuming the feed.            hypersensitivity in the gut remains an active area of
                                                               research.
    Floor feeding should continue for the first three
to four days or until the piglets are definitely eating        Additional Reading and References
the feed. When the litter is obviously eating the
                                                               Aherne, F.X., Danielsen, V. and Nielson, H.E.,
feed, a small feeder may be used. Since piglets
                                                               I 982. The effects of creep feeding on pre and post
prefer to feed together, the feed trough should be
                                                               weaning pig performance. Acta Agric. Scand. 32:
large enough to allow as many piglets as possible to
                                                               155-160.
feed at the same time. A fairly heavy, shallow,
circular trough is suitable for this purpose because it
                                                               Azain, M.J., and Jewell, D.E. 1991. Impact of early
makes the creep feed obvious and easily accessible
                                                               nutrition on pig performance. Proc. 1991 Georgia
to the piglets, as well as accommodating a fairly
                                                               Nutrition conference. pp 99-107.
large litter.
                                                               Etheridge, R.D. and Seerley. R.W., 1984. Aspects
                                                               of dietary stresses on the digestive system of the
                                                               baby piglet. Proc. 1984 Georgia Nutrition Confer-
                                                               ence. pp 32-43.
                                                             172
8. FEEDING THE WEANED PIG
  The period immediately after weaning is an                         In addition, it is almost impossible to prevent
extremely important time in the life of the pig, one               some decline in nutrient intake as the pig changes
which has a significant impact on future perform-                  from sow's milk to a starter diet. This decline in
ance. For example, on farrow-to-finish farms                       intake results in a proportionate decline in meta-
where age at market weight has been identified as a                bolic body heat generation, making the pig more
problem, one often finds the root cause in the                     susceptible to chilling at the time of weaning.
nursery, rather than in the growout barn. Indeed,                  Control of the thermal environment is critical, and
studies at the Prairie Swine Centre have demon-                    in some respects, will be related to diet composition
strated that about 30% of the variation in age to                  (see "the pig's environment" later in this chapter).
market can be attributed to the time it takes a pig to
reach a body weight of 23 kg.                                        It is self-evident that exposure to disease-causing
                                                                   organisms compromises the ability of the pig to
  Healthy, vigorous pigs that are eating and growing               adjust to weaning. However, management, as
well adjust more easily to weaning and suffer little               opposed to treatment of disease, has proven to be
or no effects of "post-weaning lag." Poor-doing,                   the most effective strategy. Attaining minimum
lethargic pigs are slower-growing, do not adjust                   disease status in a herd offers many overall advan-
well to weaning and suffer the greatest drop in                    tages. particularly for the weanling pig. However,
performance at weaning. Clearly, the key is to get                 not all diseases can be eliminated from the herd.
pigs off to a good start to ensure ongoing success                 Depending on the diseases that are a concern in a
(Figure 8-1).                                                      given herd, segregated early weaning (SEW) offers
          184
                                                                   advantages in terms of reducing pathogen transmis-
          182                                                      sion from the sow to her litter (Table 8-1 ). In the
                                                                   instance of other diseases, minimizing the age
    s
          180
    a:
          178                                                      spread of piglets within the farrowing room and
    � 176
                                                                   within the nursery is highly desirable.
    f?
    (I)
          174
    � 172
    0
          170
                                                                     In addition to the stresses of nutrition and disease,
          168                                                      the impact of a sub-optimal environment and
          166                                                      disturbances to the previous social order must not
                             0· 150       150 • 225     >225
                A.D.G. (grams) DURING FIRST WEEK AFTER WEANING
                                                                   be underestimated. A holistic approach to
                                                                   managing the newly-weaned pig, particularly as the
Adapted from Tokach et aJ. 1992. Proc. Kansas State                industry moves to weaning at an earlier age,
Univ. Swine Day. Manhattan, KS. pp. 15-17.
                                                                   becomes paramount. Modification of diet cannot
Figure 8-1.                                                        overcome the effects of a poor environment, just as
                                                                   creating an ideal environment will not compensate
                                                                   for a poor diet. All the pieces of the puzzle must
Stresses at Weaning
                                                                   be in place and integrated.
  At the time of weaning, the pig is exposed to a
series of stressors that, if left unchecked, can lead to
                                                                      However. the process of weaning, no matter how
poor performance and increased mortality. For
                                                                   carefully managed, results in changes in the gut of
example, due to removal from the sow, the piglet is
                                                                   the pig that make it more susceptible to digestive
forced to change from an almost ideal liquid diet,
                                                                   upset, diarrhea and impaired appetite. The objec-
based on lactose, fat and milk protein, to a dry diet
                                                                   tive, then, in developing diets for the weanling pig
based on different proteins, fats and carbohydrates.
                                                                   is to recognize these unique needs and facilitate a
Sow's milk offers other advantages, containing
                                                                   smooth transition to the piglet's new lifestyle,
'non-nutritive' components that facilitate digestion
and provide protection against disease.
                                                                 173
resulting in rapid and efficient growth and Limited        Photo 8-1.
health problems. This must all be accomplished
with a keen eye to cost and economic efficiency.
Table 8-2. Performance Targets for Pigs from 7 to 25          Water containing high levels of minerals,
kg Body Weight.                                            especially sulphate, can contribute to diarrhea, but
                                                           have surprisingly little impact on feed intake. Each
                           Good      Better     Best       situation where water is a concern needs to be
Average daily gain (kg)    460       520        585        evaluated closely.
Average daily feed (kg)    740       780        820
Feed conversion             1.6      1.5        1.4          Much is often said about the importance of fresh
Mortality,%                 1.0      0.5        0.2        feed. While caked, stale or fouled feed is clearly
                                                         174
                                       Q
                                       0
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                                                                                     C)
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                                                                                                                                            LO
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                                                                                                                                            C\I
                                                                                                                                            '<t
LO
                                                                                                                                                   -
                                                                                                                                            er:>
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                                                                                                                                                   -
                                                                                                                                                   �
                                                                                                                                                   "O
                                                                                                                                                   w
        -c
        ·-                                                                                                                                         �
                                   z
                                                                                                                                            <X)
                                                                                                                                            C\I
        as
        �                         �g�
                                  z�o
                                  �;s
                                  CZ)�c,
                                                                                                 ........ .
                                                                                                      l
                                                                                                   . .......
                                                                                                      . ..
                                             �
                                             z                                                          --
                                                                                                         ---.....
                                                                                                               ------·u,
                                                                                                                  -·
                                                                                                                               .........
                                                                                                                     II:   I
                                                                                                                                            0
(sweJ50l!)f) .lH813M
Figure 8-2. Growth Curves for Pigs Demonstrating Good, Better and Best Perfromance to 1 O Weeks of Age.
                                                                   175
undesirable, there is little data to suggest that feed              Other important factors that determine feed intake
needs to be added fresh daily. Indeed, limited research           are addressed later in this chapter. These include
suggests barn odours have little effect on voluntary              ambient temperature, floor space and feeder design.
feed intake in weanling pigs. However, regularly
adding new feed to the feeder appears to attract the
pig's attention and thus may stimulate intake.
                                                                  Managing the Early-Weaned Pig
                                                                     Although feed composition is critical to success in
                                                                  early weaning (<21 days), proper animal and feed
  Under some circumstances, limiting daily feed
                                                                  management is also essential. Pigs should be
intake in the newly-weaned pig might help to
                                                                  weaned in groups of 10 - 15 pigs per pen into an
control diarrhea. While such a practice may be
                                                                  all-in-all-out nursery. At the time of weaning, the
effective as a short-term solution, ultimately the
                                                                  temperature should be approximately 32° C. Heat
underlying cause of the problem needs to be
                                                                  lamps should also be used for the first two or three
addressed, as the objective in the nursery is to get
                                                                  days, especially for smaller pigs. If heat lamps are
pigs onto full-feed as quickly as possible. Only
                                                                  not available, the temperature should be increased
then will growth rate be maximized. A full
                                                                  by 2° C. The temperature can be reduced by 1 ° C
discussion of limit feeding appears later in this
                                                                  every two to three days until the room temperature
chapter.
                                                                  is 28° C. At this time, the temperature should be
1
 Not a "requirement" per se, but a suggested D.E. level for pigs of this weight (age) range. Actual D.E. will depend on
available ingredients and relative cost.
2
    All amino acid requirements expressed as "available" using ilea! digestibility as the standard.
3Lcvels    for animals destined for slaughter. Levels for animals intended for breeding stock are provided in parenthesis.
                                                               176
lowered by 1 ° C each week. These are estimates                Some people recommend minimum lighting to
only, as conditions due to drafts, building insula-          reduce tail biting and other vices. This may actually
tion, flooring material, humidity, etc., will affect the     have the opposite effect. Turning on the lights to
exact needs of the pig. Pig behaviour is a good              check the pigs then creates a high level of activity
indicator of comfort. Lying in piles on their bellies        that in some cases has led to increased vices. It is
is generally an indication of chilling. Panting is           best to create a pleasant, warm, dry, draft-free, and
indicative of heat stress. Ideally, pigs should be           uncrowded environment for the pigs. These
lying on their sides, well separated from their              conditions are the best way to avoid behavioural
neighbours. Remember, high temperatures suppress             problems.
feed intake and may contribute to illness.
                                                             Nutrient Requirements of the Weaned
  Initially, feed should be provided on pans placed
                                                             Pig
in the centre of the pen. Heavy metal pans with 12
                                                               The nutrient requirements of the weaned pig, like
to 25 millimetre edges are desirable as they are easy
                                                             other classes of swine, depend on many factors,
to work with, difficult to tip, and help reduce
                                                             including their genetic potential for lean and fat
wastage. Various feeding regimes are recom-
                                                             gain, the environment and health status. For the
mended, but we like offering 50 - 75 g per pig on
                                                             purposes of this chapter, when defining nutrient
the pan. It should be replenished three times a day.
                                                             requirements and diet composition, we assume
By the third day, pigs should be starting to use the
                                                             superior genetics, an optimurn environment and a
self-feeder. By the third to fifth day, the feed pan
                                                             reasonably high health status. There are three
can be removed.
                                                             reasons for taking this approach.
  Feeders should be adjusted to ensure ready access
                                                               First, our understanding of the interrelationship
to feed while avoiding wastage. Plenty of feeder
                                                             among health status, the environment and nutrient
space is required, as pigs at this age are group
                                                             requirements is not well understood at this time.
eaters. Single space feeders are not recommended.
                                                             Because of the importance of performance at this
To avoid fouling, the feeders should be at least 15
cm from a corner, since pigs Like to defecate in             stage in production, one will typically err on the
                                                             side of over- as opposed to under-formulation.
corners. Finally, feeders should be designed with
spacers in the trough to prevent pigs from sleeping
                                                                Second, the economic "risk" of over-feeding
in them.
                                                             nutrients in the nursery is small compared to the
                                                             potential benefit of improved performance. This is
  While nipple drinkers are easiest to keep clean,
                                                             due to the fact that pigs in the nursery eat a rela-
dish drinkers are more conducive to water intake at
                                                             tively small amount of feed; indeed, Jess than l O
the time of weaning. Most importantly, water must
                                                             percent of the total feed required to carry the pig
be readily available at all times.
                                                             from weaning to market will be consumed during
Photo 8-2.                                                   this period. In particular, during the early post-
                                                             weaning stage, diet cost will be influenced more by
                                                             selection of ingredients than levels of nutrients.
                                                             This will be discussed in the next section.
                                                           177
(Figure 8-3 ). In pigs weaned at 19 days of age, less                 Recommended nutrient requirements for the
than 5% of the total starter feed is consumed prior                weanling pig are summarized in Table 8-3. The
to 7 kg body weight.                                               recommended energy levels are expressed as a
                     18.0 to 23.0 kg
                           30%
                                                                   range and are quite arbitrary. The exact levels will
                                                                   depend on the cost and availability of high energy
                                                 3.4 to 7.0 kg
                                                                   basal feedstuffs. The ability to include moderate
                                                      13%          amounts of fat into diets based on wheat must also
                                                                   be considered. Lower energy values are possible in
                                                                   all cases, although growth rate will decline propor-
                                                                   tionately.
                                           7.0 to 11.5 kg
                                                22%
                                                                     There may, however, be an upper limit to crude
                                                                   protein in these diets, but that has not been firmly
                                                                   established. There are concerns that if crude protein
                                                                   levels are too high. scouring may be encouraged. In
              39%
                                                                   addition, the metabolic cost of eliminating excess
Figure 8-3b. Starter Feed Usage: three week weaning.               nitrogen should be minimized, allowing available
                                                                   nutrients to be directed towards more productive
  Third, during the weanling period, daily nutrient                purposes.
intake is generally the most critical factor limiting
animal performance. While genetics. as well as the                 Feeding Programs
environment and health may influence feed intake,                    For a variety of reasons, including a rapidly
few pigs are fed diets that allow full expression of               maturing gastrointestinal tract and the cost of
genetic potential because the nutrient density of the              providing an effective diet to the newly-weaned
diet is limiting for both practical and economic                   piglet, the optimum diet for weanling pigs changes
reasons (Figure 8-4).                                              almost weekly. Consequently, a phase feeding
                                                                   program is necessary to balance animal perform-
                                                                   ance and feed costs. The nature of the program will
                                                                   depend on a variety of factors. the most important
                                                                   of which is the age at weaning.
                                                                 178
result from offering small amounts of creep feed            production cycle make up for early losses in animal
containing soybean meal, or from piglets eating             performance. The presence or absence of true
small quantities of the sow's feed. This allergic           compensatory gain in pork production remains
response results in damage to the intestinal villi,         controversial.
which in tum impairs digestive and absorptive
capacity of the associated epithelial cells. Over              It is generally safe to conclude that in production
time, pigs develop a tolerance.                             systems requiring rapid animal throughput or in
                                                            early weaning systems, complex diets are essential.
  The problem is less acute in pigs weaned at four          It is difficult to envisage getting pigs to market in
weeks of age or later, due to the larger intake of          less than 150 days on simple starter diets, or
creep feeds and the resulting tolerance to soybean          weaning 12 to 17 day old pigs onto a cereal-based
proteins that develops. It is more acute in younger         feed. Conversely, with older weaning and less
pigs because their consumption of creep diets wiU           emphasis on days to market, less expensive diets
be Jess and tolerance will not yet be established.          may suffice. To restate, phase feeding allows
                                                            producers to optimize their feeding programs, using
  A variety of products derived from soybean meal,          more complex diets at the time of weaning,
such as soy protein isolates or concentrates, have          followed by increasingly simple and less expensive
been developed to overcome this problem. In                 diets later.
addition, based on research from Kansas State
University, various kinds of heat treatment, includ-          The solution, as in other controversies, remains
ing moist extrusion, help to alleviate the problem.         the same: producers should monitor performance on
                                                            their farm, consider all options and determine the
Simple Versus Complex                                       best choice under their particular circumstances.
  An age old topic of discussion in nursery diets           The final decision will depend on economics and
revolves around the use of complex versus simple            animal well-being. Without information specific to
diets. Complex diets involve a number of different          their farm, producers must depend on
ingredients, including cereal grains, proteins from         recommendations based on "typical" or "average"
both animal and vegetable sources as well as fats           results.
and/or oils. Simple diets, in contrast. generally
contain cereal grains, vegetable proteins and a small       Ad libitum Versus Restricted Feed Supply
quantity of oil. In reality, a continuum exists from          Diarrhea is an all-too-common problem in many
highly complex diets, with a dozen or more ingredi-         nurseries. Diet change, environmental stress and
ents, to a very simple diet containing no more than         reduced resistance to disease combine to increase
4 or 5 ingredients. Complex diets are more expen-           the likelihood of illness, often related to the
sive to manufacture and require more skill in               digestive system. Typically, at the time of weaning,
obtaining the large number of ingredients. They             a period of low feed intake is followed by gorging
also tend to be more difficult to manufacture and           which overloads the intestinal tract, resulting in
often are difficult to peUet. However, one cannot           scouring.
dispute the much improved performance observed
with early-weaned pigs fed such diets as compared             One effective way to address this situation is to
to the more simple formulations.                            limit feed the pigs for the first 5 to 7 days after
                                                            weaning to prevent this period of overconsumption.
  The discussion arises from the relative economics         While the practice is effective, it is also labour
of the two types of diets, and in particular, whether       intensive and requires careful management to ensure
early improvements in performance are reflected in          that feed intake is not restricted excessively,
overall gains in productivity and profitability for the     resulting in slower growth. Consequently,
farm. The answer, as one might suspect, is not a            producers seek ways to provide diets ad libitum that
simple one. Some proponents of simple diets                 avoid diarrhea and thus eliminate the need to restrict
suggest that compensatory gain later in the                 feed intake.
                                                          179
  The solution is to provide a high quality physical       Diet Composition
environment which includes good control of                   While meeting the nutrient requirements of the
temperature and embraces the all-in-all-out                weanling pig is essential in providing a balanced
management system. These two factors, along with           diet, the composition of the diet, in terms of
minimizing drafts and keeping piglets dry, will go a       ingredient content, is also very important. This is
long way towards eliminating most of the major             due to the fact that the gastrointestinal tract of the
contributors to ill health.                                young pig is not fully matured and is influenced by
                                                           the presence of anti-nutritive factors more so than in
   The other solution is diet composition. The very        older animals. It is also assumed, at least, that
high quality diets recommended for the newly-              palatability is more important at this stage of
weaned pig are designed to be highly palatable and         production.
digestible thus avoiding the scenario that leads to
gut overload. Such diets are costly, but support             The primary focus of composition of weanling
maximal performance. It is up to each producer to          diets is providing nutrients in a package that
decide whether a lower cost diet, limit-fed for the        recognizes the unique needs of the immature
first five to seven days after weaning, is preferable      gastrointestinal tract, avoids anti-nutritive factors
in their operation.                                        and maximizes early feed intake. Cost is an
                                                           important consideration, but needs to be addressed
Diet Form                                                  in the context of the total production system.
  There is not a great deal of published information       Generally speaking, diets for the newly-weaned pig,
on the relative merits of pellets, crumbles and mash       especially those weaned at less than 4 weeks of age,
in starter diets. Generally speaking, large pellets        represent a compromise between what is ideal, from
should be avoided in the very young pig as it does         the pig's perspective, and what is economically
reduce feed intake. For creep feeds and early-phase        feasible, from the producer's perspective. The
starters, crumbles, small pellets or short-cut pellets     greatest success occurs when this compromise is
are generally preferred. Further discussion on             managed most effectively.
pelleting pig diets appears in chapter 10.
                                                             While ingredients can be selected based on
Phase Feeding Programs                                     "typical" results, it is important to realize that
  Whether one is considering the breeding herd,            quality standards are particularly important in
market hogs or weanling pigs, phase feeding                weanling diets. Following are a few guidelines to
programs are employed to closely match an                  consider.
animal's changing requirements with diet
composition in the most economical manner. In the          Fats and Oils
case of the weanling pig, these changes are rapid             Young pigs, particularly those immediately after
and profound. This is due not only to nutrient             weaning are sensitive to the type of fats used.
requirements changing, but also the ingredients            Short-chained fatty acids are more digestible than
most suited to the two to three week old piglet            medium-chained fatty acids and long-chained fatty
differ in a significant fashion from those necessary       acids are the least digestible. The differences are
in the five week or eight week old pig.                    greatest in the 3 week old piglet and decline almost
                                                           weekly thereafter. As the pig ages, its ability to
  The newly-weaned pig, especially those weaned            utilize the energy from fats increases. During the
at less than 4 weeks of age, have a unique                 first 2 to 3 weeks after weaning, vegetable oils are
requirement for a diet of high nutrient density, high      clearly preferred over tallow and lard. However, as
digestibility and high palatability. Meeting these         the pig ages, animal source fats are used with
needs requires very expensive ingredients.                 increasing effectiveness.
However, as the pig ages, its needs become less
stringent and a broader array of ingredients will
support optimal performance.
                                                         180
  To maximize the utilization of energy from fats,                       dried blood cells are produced in a similar manner.
excess calcium should be avoided to reduce the                           The whole blood is treated with an anticoagulant to
formation of poorly digested soaps. For this                             maintain it in liquid form. It is then centrifuged to
reason, calcium levels must be carefully controlled                      separate the cells from the plasma. Once separated,
in starter diets.                                                        the cells are spray-dried in a manner similar to that
                                                                         applied to whole blood. With increased interest in
Blood Products                                                           plasma proteins and blood cells, blood meal is
  Only spray-dried blood products should be used                         becoming a less common ingredient.
in the diet of the young pig; flash and ring dried
blood products are of lesser nutritive value and                           The manufacture of these products has
should generally be avoided in starter cLiets.                           contributed greatly to the success of early-weaning
                                                                         programs. However, when formulating diets
  Spray-dried blood meal is collected from packing                       containing blood products, particular attention must
plants. The moisture is removed in a very                                be directed to methionine levels, as these tend to be
controlled manner to prevent damage to the                               quite low. For example, in whole blood, the ratio of
proteins. Spray-dried plasma proteins and spray-                         methionine to lysine is about 0.12, less than half of
                                                                         what is required in the final pig diet.
I
    Feed until the pigs reach 7 kg bodyweight; for later weaning. feed for at least 7 days after weaning
NB. When ranges are provided, consideration can be given to economic conditions as well as the level of stress imposed on the
animal by housing conditions.
Whey and Skim Milk Powder                                                  Skim milk powder may also be used; however, it
   Whey powder is used extensively in weanling                           tends to be a much more expensive source of
diets, especially immediately after weaning when                         lactose and casein. If skim milk powder is priced
lactose and milk proteins are preferred over starch                      competitively, it is a highly desirable product in
and vegetable proteins. However, for greatest                            starter diets.
success, high quality whey powder must be used.
For early-weaned pigs at least, spray-dried, edible-                     Dehulled Cereal Grains
grade, high-lactose whey is preferred, even though
                                                                           Oat groats, when available, tend to be a highly
it is more expensive.
                                                                         palatable ingredient for use in starter diets. In
                                                                         regions where other cereal grains are more common
                                                                    181
Table 8-5. Sample Diets for Weanling Pigs.
Ingredients
Wheat                             37.55               39.39             40.73             51.68              69.37
Com                         36.69               39.01             41.37             52.52              66.38
Soybean meal - 47%          15.00 15.00         20.00 20.00       25.00 25.00       25.00 25.00        28.65 25.00
Spray dried whey            25.00 25.00         20.00 20.00       25.00 25.00       15.00 15.00
Lactose                      5.00 5.00           5.00 5.00
Select menhaden fish meal    2.50 2.50
Spray-dried plasma proteins 6.00 6.00            5.00   5.00       2.50    2.50
Spray-dried blood meal       2.00 2.00           2.50   2.50       1.25    l.25       2.50     2.50
L-lysine HCI                 0.21 0.15           0.11   0.03       0.10    0.04       0.08              0.12     0.13
L-threonine                  0.14 0.11           0.11   0.08       0.13    0.10       0.08     0.04     0.10     0.10
DL-methionine                0.16 0.09           0.17   0.10       0.15    0.08       0.07     0.03
Fat/oil                      3.80 3.60           4.10   3.90       1.00    l.80       1.00     2.00     0.50     l.15
Premix                       3.50 3.50           4.00   4.00       3.50    3.50       3.75     3.75     4.25     4.25
Antibiotic                   +     -r            +      +          +                  +        +        +        +
Nutrients. minimum
D.E., kcal/kg                 3600 3550         3600 3550         3450 3450         3450 3450          3400 3400
Crude protein, %              22. l 23.9        22.0 23.8         22.2 24.1         20.7 23.l          19.5 21.5
Lysine:D.E., g(Mcal            3.60 3.60         3.60 3.60         3.60 3.60         3.20 3.20          2.90 2.90
Dig. lysine,%                  l.30 1.28         1.30 1.28         1.24 1.24         1.10 l.10          0.98 0.98
Dig. methionine,%              0.36 0.36         0.36 0.36         0.36 0.36         0.31 0.31          0.28 0.28
Dig. TS.A.A., %                0.72 0.72         0.72 0.72         0.72 0.72         0.61 0.61          0.55 0.55
Dig. threonine, %              0.85 0.85         0.85 0.85         0.85 0.85         0.72 0.72          0.65 0.65
Dig. tryptophan, %             0.22 0.22         0.22 0.22         0.22 0.22         0.19 0.19          0.17 0.17
Calcium,%                      0.90 0.90         0.90 0.90         0.90 0.90         0.80 0.80          0.70 0.70
Phosphorus, %                  0.70 0.70         0.70 0.70         0.70 0.70         0.65 0.65          0.60 0.60
NB. These are example diets only and should not be employed without consulting a professional nutritionist. Exact
formulations will require integration of the nutrient profiles oflocal ingredients, as well as completion of least cost
procedures. Also. the technology associated with starter diet formulations is changing very rapidly.
The Phase ll diet can be fed from a body weight of7 to 12 kg. Under some circumstances, producers will find that
such a diet wi II provide satisfactory performance for pigs weaned at 4 weeks of age, avoiding the need for the more
expensive Phase I diet. Success will depend on such factors as quality of the nursery, health status of the pigs and
overall management skills of the herdsman.
The Phase rI1 diet is the simplest in design, requiring no expensive plasma proteins, whey powders, etc. Blood meal
would only be used if economical. Whey powder could be used if problems are observed, but under reasonable
conditions, whey powder will not be required if the previous feeding program has been adhered to.
                                                            182
Table 8-6. Effect of Medication on the Performance          The Phase I diet is designed to be fed from the
of Weanling Pigs in a High Health Herd.                   time of weaning until the pig reaches 7 kg, or for at
                                                          least one week, whichever is longer. lt may also be
                             Control     Medicated        used as a creep feed. A variety of Phase 1 diets are
Average daily gain (kg)       0.37         0.47           provided in Tables 8-4 and 8-5, depending on the
Average daily feed (kg)       0.58         0.70           age at weaning. In general, the younger the age at
Feed conversion               l.56         1.48           weaning, the lower the level of starch and vegetable
                                                          proteins and the higher the desired levels of lactose
NB. Four week trial, commencing immediately upon          and milk proteins. For pigs weaned at less than two
weaning                                                   weeks of age, lactose levels should be in the range
                                                          of 20% or more. Spray-dried plasma proteins,
Source: Patience and Christison, 1988.
                                                          spray-dried blood proteins and spray-dried red
                                                          blood cells have all proven beneficial. The exact
than com, groating oats represents a method to            levels required will depend on conditions within the
enhance the digestible energy content of what might       barn and will, of course, be influenced by economic
otherwise be lower energy feeds. Naked oats and           considerations.
dehulled barley represent another alternative to
increase the energy content of the feed, but their          As the weaning age rises, the Phase I diet will
value in starter diets has not been fully defined.        become less expensive as a greater portion of the
                                                          energy and protein can come from plant sources.
Diet Formulations                                         If the pigs are weaned at 21 days of age or older,
  Many alternatives are available in developing           they should receive the appropriate diet in Table 8-4
effective weanling feeding programs. Diets                for a period of at least one week, before switching
presented in this chapter are offered as examples         gradually to the Phase Il diet.
only.
                                                            While such diets are expensive, they are conswned
  Producers are strongly encouraged to discuss their      in relatively small quantities. so the cost per pig is
specific needs with a qualified nutritionist as the       quite small. For example, at the time of writing, feed-
formulation of starter diets is a highly specialized      ing the "later wean" Phase I diet for one week after
science, one which is advancing at a rapid rate.          weaning would cost less than $1.75 per pig. The fi-
                                                          nal decision on the use of such diets rests with its
                                                          impact on the overall profitability of the farm. The
Photo 8-3.                                                cost per tonne of feed tells only a part of the story.
                                                          Feed Additives
                                                          Antibiotics
                                                            There is little doubt that antibiotics employed for
                                                          growth purposes are economical in diets for pigs up
                                                          to 25 kg body weight. Research at the Prairie
                                                          Swine Centre, a high health herd. confirms this
                                                          conclusion (Table 8-6). In this particular experi-
                                                          ment, the medication increased net income by about
                                                          one dollar per pig.
                                                        183
time. Such guidelines also make economic sense.                 compared to complex formulations. As with all
ln addition, there are legal implications, as such              feed additives, selecting the right conditions for
high levels of zinc cannot be added to the feed                 their use is critical to their success.
without a veterinary prescription.
                                                                The Pig's Environment
Zinc Oxide                                                      Temperature
   A number of years ago, research in Denmark                     The weanling pig presents a unique set of circum-
revealed that adding zinc to the diet of newly-                 stances in terms of establishing an ideal thermal
weaned pigs reduced the incidence of diarrhea,                  environment. In particular, low post-weaning feed
improved growth rate and reduced mortality. Not                 intake elevates the pig's thermal requirements; for
surprisingly, the use of zinc supplements has grown             example doubling the newly-weaned pig's feed
rapidly in the interim. Generally, the recommenda-              intake reduces its lower critical temperature from
tion is to provide 2.5 to 4.0 kg of zinc oxide per              32°C to 26°C! This is due to the body heat gener-
tonne of feed for l O to 20 days post-weaning.                  ated by normal digestion and metabolism. Conse-
                                                                quently, room temperature must be linked to the
Copper Sulphate                                                 piglet's feed intake.
  Copper sulphate is a relatively common growth
promotant added to many starter diets. It is                  Various components oftbe nursery facilities may
relatively inexpensive, and for this reason, its use is     contribute to the overall thermal environment.
almost automatic in many situations. However,               While the effect of moisture and draughts on pig
concerns about the environment, recognition that            comfort are well known, the impact of cold walls,
the response to copper sulphate is not universal and        floors and ceilings, in fact, any cold surface, which
the appearance of "sticky" droppings from treated           increase radiant heat losses from the pig, lower the
pigs has tempered its use in recent years. If               effective ambient temperature relative to measured
included in the diet, the exact amount will depend          temperature. Clearly then, in addition to warm air
on local legislation, but is typically in the range of      temperatures, well-insulated walls and ceilings are
125 ppm copper as copper suJphate.                          critical to piglet comfort.
  The observation that organic acids are not always           Temperature variation is also a particular concern
beneficial may be due to the fact that pH is not            in the nursery. Twenty-four hour fluctuations in
always excessively high. Some research has                  excess of 2°C have been shown to reduce both
suggested that mixtures of acids will be more               growth rate and feed conversion efficiency. It
beneficial than single additions. The response to           should be noted that so-called high-low thermom-
organic acids will generally be greater in simpler as       eters may underestimate the extent of temperature
                                                          184
Table 8-7. Recommended Floor Space Allowances for Weanling Pigs.
Adapted from Recommended Code of Practice for the Care and Handling ofFarrn Animals.Pigs, Ag. Canada, 1993.
fluctuation in the nursery, because they do not           Additional Reading and References
record rapid changes very effectively. For example,       Nelssen, J.L., R.D. Goodband, S.S. Oritz and M.D.
in mercury thermometers, the response to tempera-         Tokach. 1995. The effect of weaning age on
ture change is quite slow and may underestimate           nursery pig behavior and growth performance. In.
fluctuation by 50% or more.                               (Foxcroft. G., ed), Advances in Pork Production,
                                                          Vol. 6, pp. 95-112. University of Alberta, Edmon-
  While too low a temperature is undesirable,             ton, AB.
failure to lower room temperature as the pig's
thermal needs change will suppress appetite and           Pettigrew, J.E. and J.T.F. Stairs. 1991. The impact
lower overall animal performance. Managing room           of starter diets on the subsequent growth of swine.
temperature in the nursery is not a simple task,          In: Proceedings of the Minnesota Nutrition Confer-
because the consequences of error are severe.             ence, pp. 163-170, Minneapolis, MN.
                                                        185
186
9. FEEDING MANAGEMENT OF MARKET
HOGS
   Approximately 60% of the total cost of pork                    control, pig density, and diet composition. The goals
production is associated with growing out the feeder              defined in Table 9-1 are a reasonable guide for most
pig. Traditionally, more attention is paid to                     feeder barns. These goals arc identified as good,
managing sows and starter pigs than growing-                      better, and best to denote the ease with which they
finishing pigs. The breeding and farrowing areas are              can be achieved. The values in the 'good' column
often considered more challenging and therefore,                  are being achieved consistently by many producers.
according to many people, deserve more of                         Those in the 'better' column are being met by some
management's attention. The indifference shown to                 producers so are currently possible. The goals in the
the feeder barn probably stems from the presence of               'best' column are not all being achieved consistently
apparent problems in the sow-related areas and from               at the present time, but the potential exists to meet or
a lack of appropriate records in the feeder barn.                 exceed them under the right combination of
                                                                  management. The tremendous gains made in the
                                                                  recent past provide confidence that the goals in the
  Records are very important in identifying both                  'best' column will be met or surpassed in the near
production problems and opportunities to improve                  future.
performance in the feeder barn. Fortunately, as better
record keeping systems and performance monitoring
systems become available, and as potential                        Table 9-1. Goals for the Feeder Barn.
improvements in profitability become more apparent,
more attention is paid to the feeder barn. Feed is the                                             Goal Level
largest single expense in any swine operation.
                                                                                          Good        Better       Best
Although there must be an absolute minimum, the
variation in feed costs among farms is proof that                 Age at 105kg             160         150         140
major savings can be achieved in this area. Even on               Days in Feeder Barn      110         100          90
farms with high management standards, differences                              Feeder Barn (20 - 105 kg)
in feed costs (up to 25% per pig) still exist.
                                                                  Avg Growth Rate (g)      775         850         950
                                                                  Feed Conversion          3.20        2.85        2.60
Goals for the Feeder Barn                                         Mortality(%)             2.0         0.5         0.1
  Establishing production objectives for the feeder               Carcass Index            107          110        112
barn is important. These objectives will vary from
farm to farm depending on a variety of circumstances
including pig genotype, health status, environmental                These goals are to be used only as guidelines. The
                                                                  overall objective is to maximize profit. Establishing
Photo 9-1.                                                        the proper goals for your operation is just as
                                                                  important, if not more so, than your capital outlay.
                                                                  For example, although superior pig performance is a
                                                                  desirable goal, setting a goal of the highest standard
                                                                  of performance may not be profitable because the
                                                                  extra expense needed to produce the high standard
                                                                  may not be regained from the market. Setting
                                                                  arbitrary goals without records to back them up may
                                                                  also be costly. A strict capital invested to goals
                                                                  accomplished ratio does not exist, either. In some
Approximately 60% of the total cost of production is              cases, the overall goal of profitability can be attained
associated with the feeder barn. Feed is the majD.r_npe11st>._    with a low capital approach that frees the operation
                                                                 187
from heavy debt and cushions it against depressed                In young pigs, growth consists largely of bone and
prices and high interest rates. Establishing your              muscle, but as the pig reaches maturity, fat deposition
goals will help you determine the areas important to           occurs at an increasingly rapid rate. For example, in
you, and how much you are willing to spend in each             the new-born pig, muscle and bone represent 29 and
of those areas.                                                19%, respectively, of total empty body weight while
  Once production goals have been established, the             fat represents less than 4% of the total. By 28 weeks
next step is to develop feeding and management                 of age, muscle and bone represent 32 and 7%,
strategies to accomplish these goals. Finally, a record        respectively, and fat has risen to 35% of the total
keeping system should be implemented to monitor                (Figure 9-2).
                                                                                            BIRTH
important production parameters, such as days in the
barn, feed usage, carcass lean yield, and mortality.
Lean Tissue Growth in Growing-
Finishing Pigs
  ln feeder pigs, both the growth rate and the
composition of growth should be considered.
Growth is the increase in body weight with time or                                    • FatH%
age of the pig. A typical growth curve is shown in                                    •     Bone3U%
                                                                                      CJ    MJsde 56.2%
Figure 9-1. This curve, a sigrnoida1 growth curve,
                                                               Figure 9-2a.
represents a pig's theoretical growth pattern from
                                                                                           28WEEKS
fetus to adulthood. It is clear that at an early age, the
pig's gain is at an increasingly rapid rate; later, rate of
gain is essentially constant and later still, decelerates
as the pig approaches full, mature weight. This curve
represents a pig's potential but under normal farm
conditions Jess than optimal growth may occur due to
limitations in genetics, nutrition. housing, disease, or
other management aspects. Furthermore, the curve is                                   • Fat46.9%
influenced by sex - boars and barrows grow faster                                     • Bone 10.1%
                                                                                      D     �e43,0%
than gilts. Pigs are generally slaughtered at a weight         Figure 9-2b.
which is under half their mature body weight and at a
time that growth rates are just starting to decline.
                                                               Figure 9-2. Relative Proportions of Muscle, Bone and
                                                               Fat in Pigs at Birth and 28 weeks of Age (From
                                                               McMeakan, 1940).
                                                            189
Table 9-2. Saskatchewan Carcass Grading Grid (effective March 6, 1995).
                                                t'-C"'IO\t'-\OViM0\00
                                                                   r-- I"--
                                                0,. 0,. 00 00 00 00 00 00
                                                0 00         °'°'
                                                ViC"'IOOO\O...;t'NO,....;t-0
                                                             o,. 0,.00 0000
                                   °' °'
                                                                                    ·�
                                    °'°'
                              <o r- °'                                 °'°'
                                                ...;t'MNOl'-...;t'Noor-o
                                                          000          o,.          �
                                                                                     ....
                                   00 00                                            .....
                                                                                     0
                                                                                      E
                                                                                      E
                                                                                    00
                              Vi
                                   0 00
                                       °'
                                       o-
                                       '°
                                   00
                                                N Ooo r- Vi MN 00 MO
                                                O Oo,. o,. a, o,.o,. oo 00 00
                                                OOOOOOOOViC"'IO
                                                00 00 00 00 00 00 I'- I'- r- I"--
                                                                                      E
                                                                                      :::,
                                                                                      E
                                                                                    .5
                                                                                      E
                                                                                      �
                                                                                      oI
                                                   00 \0 Vi Vi \0 C; 0- 0           ...._...E
                                                   01....:oo;oor-\O\O°'
                                                00 \0 \0 \0 Vi Vi Vi Vi Vi            e
                                                                                    ;.;:::
                                                N
                                                                                     0
                                                '°/\                                £
-NM...;t'Vi\Ol'-000,0
                                                       190
Table 9-3. Observed Lean Growth Rates in Growing-finishing Pigs (25 to 105 kg body weight) in Relation to
Growth Rates and Carcass Lean Yield Content (75-85 kg carcass weight)*.
* relates to carcass weights between 75 and 86.99 kg in Saskatchewan; 1995 Canadian carcass grading system;thc high,
medium, and unimproved lean growth rates correspond to lean growth rates of approximately 380, 340, and 300 g/d.
potential are essential for developing cost-effective          more likely to consume quantities of feed that are
feeding strategies in individual production units.             equivalent to 90% of the daily DE intake in growing-
                                                               finishing pigs as suggested by NRC ( 1987). Figure
                                                               9-4 represents feed intake curves for feeder pigs
  Clearly, the objectives in raising pigs are to               consuming a diet with a DE content of 3150 or 3400
optimize lean tissue gain and to minimize the accu-            kcal per kg and at 90% of voluntary feed intake
mulation of body fat. The overall goaJ is to meet              according to NRC ( 1987). This graph may serve as a
these objectives profitably.                                   benchmark to compare feed intakes on individual
                                                               pork production units.
                                                           191
dietary energy density (with an increase in daily feed     and not be sensitive to mechanical failure. No sharp
intake) in such a manner that the daily energy intake      objects or protruding objects should be present. If
remains constant. Grower (and starter) pigs are            well designed single space feeders arc used, one
generally unable to adjust feed intake with changing       feeder is sufficient for up to 12 pigs. When the water
energy densities. For these pigs physical feed intake      nipple is placed inside the feeder (wet feeders), feed
capacity, or "gut fill", determine feed intake, An         intake is generally higher ( l - 7%) as compared to
increase in dietary energy density will result in          that of standard dry feeders. Problems with wet
increases in the daily feed intake. It should be           pens, water wastage and the developments of molds
stressed that these principles do not apply to extreme     are more likely to occur when wet feeders are used.
dietary energy densities and will be affected by
environmental conditions. The effect of diet energy
density on daily energy intake can have important            Given the above considerations, it is important to
consequences for the optimum energy density in the         monitor feed intake as well as the main factors that
diet (see "diet formulation" later in the chapter).        affect feed intake.
Other nutrients have limited effects on feed intake
when present at levels in the feed which are within
                                                                                                                 --
                                                                  3.5
practical range. If feed is the suspected problem with
intake, then the various aspects of feed formulation       ,::,                                                            - L---
                                                                                                                            - ..
                                                                                                              .. .. - - . -
                                                                   3
and preparation should be quickly evaluated. These
include: ingredient composition and quality (inclu-
                                                           °'or
                                                           .Y.
                                                                  2.5
                                                                                              �- ..    --   i.--
                                                     192
energy adapting to their environment (e.g., shivering
                                                                                                      ,,
                                                                           "average" pig             "good" pig
to keep warm). If the environmental temperature is
suitable, pigs should not require feed to maintain a
constant body temperature, or combat diseases.                                                                                       Fat
                                                            Gain
                                                                                                     '
                                       Fat
                                                                                                             ''
                                                                                                                      ..        ·•
                                                                                                                                     Lean
                                                                                                                  "   -    \-
                                                                                                     /
                                                                                                     ..
                                                                                                         '�';'.       '
     Gain
                                       Lean
        0
                                                                       1       2      3          1            2                  3
                                      Maintenance
                                                                                   Feed Intake (kg/d)
                                                          193
                                                                                                           dietary lysine levels and lean growth in two types of
                                                                                                           pigs is demonstrated in Figure 9-8. At low levels of
                                                                                                           lysine in the diet there is no difference in
  ' -,                                                                                                     performance between the two types of pigs. Only
                                                                                                           when dietary lysine levels are increased can pigs with
         ' ' -,                                                                                            the higher lean tissue growth potential demonstrate
                                                                            ··gain
                       . ....: <,
                      <,
                                                                            - feed: gain
                                                                                                           their potential. Pigs with unimproved lean tissue
                                     ......                                                                growth potential will not respond to the increase in
                                              ......
                                                       ......               - lean gain : fat gain
Figure 9- 7a.
                            Feed Intake
                                                                    -                                      them, and to ensure that an animal with high
                                                                                                           performance potential receive sufficient quantities, it
                                                                                                           is important to monitor lean growth rates (see earlier
                                                                                                           section on lean tissue growth) and to establish the
                                                                                                           animal's lean growth potential.
   ' -,
          ' -,   <,
                                                                             ··gain
                                                                             - feed: gain
                       . ...:
                      <,
                                                                             -lean gain : fat gain
                                ' ...... .............
                                                                --- ---
                                                                                                                  500
                                                                                                     194
Table 9-4. The Effect of Health Status on Performance       Because animals require nutrients on a daily basis,
in Growing-Finishing Pigs (Williams 1994, Iowa State      estimates offeed intake per day are required to
University).
                                                          determine the optimum nutrient level in the diet.
                                                          This estimation is especially important for amino
Exposure to Disease           Low          High           acids in finishing pig diets. For example, based on
                                                          the animal's lean growth potential, a finishing pig
Feed intake, kg/d             2.47          2.36
                                                          may require 20 g of lysine per day. As there is
Growth rate, kg/d             0.89          0.78          generally no effect of energy intake on lean growth
Feed: Gain                    2.76          3.04          rates in finishing pigs, daily lysine requirements are
                                                          not affected by feed intake. ff the feed intake in the
Muscle in the carcass, %      57.0          52.5          finishing pigs is 2.50 kg per day then the optimum
                                                          lysine level in the diet should be 8 g/kg, When feed
                                                          intake is 3 kg/d, the lysine level should be 6.66 g/kg.
  The animal's lean growth potential may be               This reduction in required dietary lysine level will
estimated from observed lean growth rates in the          substantially reduce the cost of the finishing diet.
feeder barn (sec Table 9-3). However, take care in
interpreting lean growth rates. Other factors, such as      In growing pigs, up to approximately 60 kg body
feed intake or unbalanced diets, rather than the          weight, energy intake generally limits lean growth. A
animal's potential, may affect observed lean tissue       reduction in daily energy (feed) intake will reduce
growth rates.                                             lean tissue growth and the daily requirements for
                                                          amino acids. Iffeed intake happens to be reduced in
  A pig's daily nutrient requirements increase as it      grower pigs, there is no need to increase the dietary
grows heavier, but so does its feed intake. In fact, a    amino acid levels in the remaining food intake. For
pig's feed intake increases more rapidly than its         this reason, it is more important to balance the amino
increase in daily nutrient requirements. As a result,     acid levels in the grower pig diets based on amino
nutrient levels in the feed can be gradually decreased    acid to energy ratios than any other ratio, and to
as the animal grows (Figure 9-9). This concept            formulate finisher pig diets based on daily amino
forms the basis of phase feeding. Phase feeding uses      acid intakes.
different diet formulations to meet the pig's nutrient
requirements at each stage of its production, which
reduces over-feeding excessive quantities of                   Question: When should I determine the optimum
expensive nutrients.                                           amino levels in the pig's diet based on daily
                                                               intakes and when based on amino acid to energy
                                                               ratios?
-
>
�
 e                                                             Why: ln finishing pigs, energy intake docs not
-
 Cl)
·;:                                                            generally limit lean growth. As a result, there is no
-
 :::,                                                          relationship between energy (or feed) intake and body
 c
                                                               protein deposition; daily amino acid requirements arc
 Cl)
                                                         195
   The environment also affects a pig's nutrient               Due to the many factors that affect nutrient
requirements, especially nutrients needed for body           requirements, there is a wide range in recommended
maintenance functions. In modem and well                     nutrient levels for growing-finishing pigs. For
managed, confinement growing-finishing pig                   example, recommended lysine levels may range from
 facilities, environmental conditions can be optimized       as high as 1.3% in early grower diets for pigs with
and the amount of feed required for maintenance              extremely high lean growth potential to as low as
minimized. If the environment is too cold,                   0.5% in late finisher diets for pigs with poor lean
maintenance energy requirements quickly increase.            growth potential that consume large quantities of
Under cold stress conditions, growing pigs (25 to 60         feed. This variance again stresses the importance of
kg body weight) require approximately 25 g of extra          recognizing the main factors that affect nutrient
feed to maintain a constant body temperature per °C          requirements in various groups of pigs for
drop in environmental temperature. For finishing             developing unique management and feeding
pigs, this value is approximately 40 g of feed. This         strategies for individual production units. It also
demand means that total feed usage would increase            supports the need for a factorial, or modelling,
by approximately 15% for growing-finishing pigs              approach to estimating nutrient requirements in
kept in an environment that is effectively l O °C too        which the main factors that affect nutrient
cold for them. This usage is equivalent to a drop in         requirements are considered. However, when such a
feed conversion ratio from approximately 2.9 to 3.3.         modelling approach is used, different agencies or
The cold environment is one of the major reasons for         researchers will still provide different estimates of
the poor feed efficiency observed during the winter          nutrient requirements. For example, the estimated
months in low-cost, outdoor feeder pig housing               lysine requirements for a 50 kg pig with a given rate
systems. Remember that the effective environmental           of lean growth {body protein deposition), estimates
temperature to which pigs are exposed is determined          may vary by as much as 30% (Table 9-5). Obviously
not only by the environmental temperature but also           our knowledge of pig nutrition is not yet complete.
by other factors such air speed, air humidity, floor         As we continue to improve our understanding of
type, dryness of the floor, and the pig's body               nutrient utilization in pigs, these discrepancies will
condition.                                                   disappear.
   In developing feeding programs for feeder pigs it is      Table 9-5. Estimated Available (Apparent lleal
also important to consider the production objectives         Digestible) Lysine Requirements for a Growing Pig at
                                                             50 kg Body Weigbt1•
and the economic conditions. Different feeding
strategies and diet compositions are required when
the objective is to maximize income per pig rather               Moughan et al., 1987                        .54%
than income per pig place per year, or when
economic conditions change.                                      Stranks et al.. 1988                        .59%
                                                                 Fuller et al., 1989                         .43%
                                                                 TMV2, 1991                                  .62%
Nutrient Allowances and Diet
Formulation                                                      Moughan, 1992                              .65%
  The basic concepts of diet formulation are covered             Whittemore, 1993                           .58%
in chapter 5. In that chapter, the importance of
fonnu la ting diets on the basis of available rather than    I
                                                              Average lean tissue growth rate (340 g/d according to the
total nutrients is emphasized, as well as the                Canadian definition of lean , this is equivalent to 130 g/d of
importance of monitoring feed ingredient quality,            body protein deposition). The pig consumes 2.2 kg of a
feed preparation, and feed handling. As we continue          diet with a DE content of 3150 kcal/kg (feed intake is
to meet the animal nutrient requirements more                equivalent to 90% ofNRC [1987)). Pigs are assumed to
                                                             be in a thermo-neutral and relatively disease free
closely through phase feeding, split-sex feeding, and        environment.
feeding for different genetics, these basic concepts of      2
                                                                 Technisch Model Varkensvocding.
diet formulation and quality control become
increasingly important.
                                                       196
  The optimum energy density in feeder pig diets is               In growing pigs, physical feed intake capacity, or
determined largely by the cost per unit of energy in           gut fill, generally determines feed intake. An in-
the available feed ingredients. As mentioned earlier,          crease in diet DE content will thus result in propor-
finishing pigs (over 60 kg body weight), are                   tional increase in the daily DE intake. As illustrated
generally able to adjust feed intake with changes in           by the data in Table 9-7, an increase in energy density
dietary energy density in such a way that the total            in the diet for growing pigs will result in improve-
daily energy intake remains constant. at least when            ments in both feed efficiency and growth rate. For
the diet's DE content ranges between 3000 and 3400             growing pigs, the optimum energy density is not only
kcal/kg. In other words, if the diet's DE content is           determined by the cost per unit of energy in the diet,
reduced by I 0% from 3300 to 3000 kcal/kg, the                 but also by the effect of energy density on growth
finishing pig will simply consume 10% more feed,               rate, and thus throughput in the grower barn. As the
growth rate will not be affected, and the feed                 animal's Jean tissue growth potential continue to
conversion ratio (feed/gain) will simply increase by           increase, energy intake will become a limiting factor
10%. In this case, the optimum energy density in the           up to higher live body weights. This situation may
diets should be based on purely the cost per unit of           also mean that finishing pigs with extremely high
energy in the diet as outlined in Table 9-6.                   lean tissue growth potential may be unable to com-
                                                               pletely maintain a constant daily energy intake as the
                                                               energy density in the finishing diet is reduced.
Table 9-6. Estimation of the Lowest Cost per Unit of           Clearly, more information is required on the interac-
Energy in Growing Pig Diets (varying in energy
density)*.                                                     tive effects of pig genotype and body weight, as well
                                                               as tbermaJ environment and animal density, on the
                                                               optimum energy density in the diet.
                         Diet 1     Diet 2     Diet3
DE content, kcal/kg       3150       3250       3350
                                                                 In most practical swine diets, lysine is the first
Ingredient comp.(%):                                           limiting amino acid. It is thus appropriate to first
 Barley                   59.30      35.29       3.12          define the target (available) lysine level in the diet
 Wheat                    17.40     42.80       75.51          and to derive the required level of other amino acids
                                                               from that oflysine based on an optimum amino acid
 Soybean meal, 4 7%       15. JO     18.10      18.15
                                                               balance (outlined in chapter 3). The suggested
 Canela meal              5.20       0.70        0.00          allowances of available (apparent ileal digestible)
LysineHCI                                        0.03          lysine in relation to body weight and lean growth
 Premix                   3.00       3.11        3.19          rates are presented in Table 9-8. Allowances are
                                                               expressed either as lysine to energy ratios or as daily
Calculated nutrients:**
                                                               lysine intakes. These suggestions are based on a
 DE, kcal/kg              3150       3250       3250           factorial estimation of lysine requirements to support
 Avail. Lysine,%          0.70       0.72       0.74           the indicated lean growth rates. A safety margin of
 Avail. Threonine,%       0.46       0.47        0.48          about 5% for the grower phase and I 0% in the
Prices:
 $/tonne                 132.62     138.44      146.11         Table 9-7. Effect of Dietary Energy Density on the Voluntary
 $/Meal DE                42.10     42.60       43.61          Feed Intake and Performance of Entire Male Pigs between 22
                                                               and 50 kg Body Weight (Campbell and Taverner, 1986).
Estimated Feed usage:
 Kg per pig            160            155       150.5          DE content,
                                                               kcal/kg                  2820 3035 3250 3466 3600
 Feed: Gain           3.55           3.44        3.34
 Meal DE : kg gain        11.18      11.18      11.18          Voluntary Intake, kg/d    2.19   2.21    2.19    2.17    2.05
                                                               Voluntary DE, Mcal/d      6.14   6.62    7.10     7.48   7.39
* Prices of ingredients: barley $80/tonne, wheat $95/tonne,    Growth Rate, g/d
soybean meal $265/tonne, canola meal $190/tonne, lysine                                  695     776     847     898     913
HCL $3000/tonne, premix $600/tonne.                            Feed: Gain                3.16   2.89    2.61     2.39   2.25
** Levels of all nutrients are adjusted with diet's energy     Carcass P2               14.40   15.30   15.60   16.00   16.40
content to ensure a constant nutrient to energy ratio.
                                                              197
 finisher diets is already included in these values to                Once the requirements for available lysine are
allow for inaccuracies in feed preparation and varia-              established, the requirements for the other essential
tion in feed intake. In comparison to other factorial              amino acids can be determined based on the opti-
estimates of lysine requirements, the suggested                    mum amino acid balance, i.e., the concept of ideal
allowances can be considered high (Table 9-5).                     protein as outlined in chapter 3. It is estimated that
However, the suggested allowances are supported by                 per I 00 g of available lysine, growing-finishing pigs
empirical animal performance studies as well. The                  require 60 to 70 g available threonine, 56 to 64 g of
actual recommendations that are expressed as a                     available methionine plus cystine, and 18 to 20 g of
percentage in the diet in Table 9-8 relate to a diet               available tryptophan. At least 50% of the methionine
with an energy density of 3150 kcal/kg and for pigs                plus cystine requirements needs to be supplied by
consuming average quantities of feed (90% ofNRC                    methionine. The ratio of these amino acids to lysine
 l 987; Figure 9-4). If feed intake is different from the          increases with increasing body weight or with reduc-
indicated values or when the diet DE content is                    tions in lean growth potential.
different from 3150 kcal/kg, the available lysine
levels in the diet should be adjusted to maintain a
constant available lysine to energy ratio in the grower              Question: How do I adjust the suggestions in
diets, and a constant daily available lysine intake in               Figure 9-8 for the diets on my farm? My pigs
the finisher diets.                                                  have an average lean growth potential; I feed a
                                                                     grower and a finisher diet with DE contents of
                                                                     3300 and 3000 kcal/kg, respectively. I have
Table 9-8. Recommended Allowances of Available (ap-                  established feed intake curves on my farm:
parent ilea) digestible) Lysine in Relation to Lean Growth           consumption on my grower and finisher diets are
Potential and Body Weight in Growing-finishing Pigs.                 2.2 and 2. 7 kg/d, which is equivalent to 85% of
                                                                     estimated intake according to NRC (1987).
Body weight
(kg)          25            45      60      70         110
                                                                     Answer: The available lysine level in the grower diet
High*                                                                should be increased to 0.74 %. In the finisher diet it
                    2.70          2.35                               should be 0.65%.
                                                  20.00
                   (.86%)        (.74%)           (.67%)
Medium"                                                             Background: The lean growth potential are average so
                                                                    the recommendation derived for these pigs should be
                     2.25                 17.50                     the average lean growth rates - medium - in Figure
                    (.71%)               (.61%)                     9-8. In the grower diet the available lysine to energy
                                                                    ratio should be 2.25. At a DE content of3300 kcal/
Unimproved"                                                         kg, the available lysine level should be 7.40 g!kg or
                      1.95                15.50                     0. 74%. In the finishing phase, the daily available
                                                                    lysine requirements are 17.50 g/d. Given the daily
                    (.61%)               (.54%)                     feed intake of 2. 70 kg, the available lysine level in this
                                                                    finisher diet is 17.50 divided by 2.70, which is 6.50 g/
                                                                    kg or 0.65%.
* The average lean growth rates are 480, 440, and 400 g/d.
for pigs with high, medium, and unimproved lean growth
rates. respectively (see Figure 9-6 also).
                                                             198
  In Tables 9-10 and 9-11, the suggested additions of            Table 9-10. Recommendations for Vitamin Fortiflca-
vitamins and minerals to practical grower and fin-               tion of Grower and Finisher Diets=.
isher diets are summcrized. These suggested values
include a safety margin to account for inaccuracies in           Vitamin          Units/kg          Grower       Finisher
feed preparation, variation in requirements for                  Vitamin A           IU               7000         5500
different groups of pigs, loss of potency of vitamins
during storage, and the effect of stress on require-             Vitamin D           ru                700          550
ments. Given these safety margins and the cost of                Vitamin E           IU                351          251
these nutrients, it is not critical to adjust these              Vitamin B12        mcg                20            15
suggested levels of fortification for specific groups
of pigs. Exceptions may be made for breeding stock,              Vitamin C           mg                 -2           -2
animals with extremely high lean growth potentials               Vitamin K           mg                2.5           2
(such as entire males) or when diets with extremely                                                    _3.4
                                                                 Biotin             mcg                             _3,4
high energy densities are fed. In these cases the
                                                                 FolicAcid           mg                04            04
suggested additions may be increased by up to 20%.
                                                                Niacin               mg                25           20
                                                                199
Table 9-11. Recommendations for Mineral Levels in Grower and Finisher Diets.
Macromincrals - Total
Calcium                                                         %                             0.70                   0.60
 Phosphorus                                                     %                             0.60                   0.50
 Available Phosphorus                                           %                             0.25                   0.20
 Sodium                                                         %                             0.15                   0.15
 Chloride                                                       %                             0.18                   0.18
 Potassium                                                      %                             0.35                   0.35
Magnesium                                                       %                             0.05                   0.05
Microrninerals - Supplemented
 Iron                                                         mg/kg                            701                   501
 Copper                                                       mg/kg                            151                   JOI
 Zinc                                                         mg/kg                           JOO'                   901
 Iodine                                                       mg/kg                           0.30                  0.20
Selenium                                                      mg/kg                           0.30                  0.30
* The values in this table are minimum levels or supplemented quantities recommended for practical diets. Due to varying conditions and
the influence of diet type on mineral requirements. deviations from these numbers may be required in some circumstances. In pigs with
extremely high performance levels, when pigs may be used as future breeding stock, or when high nutrient dense diets are fed these levels
may be increased by 10 - 20%.
I
 Copper may be used as a growth promotant at 125 mg/kg in the diet. Due to interactions in the utilization of copper, zinc and iron, the
levels of iron and zinc should be raised by approximately 60 mg/kg when copper is included at growth promoting levels.
Photo 9-2.
                                                                                                i
                                      ..........,
                                                                         ...            �
                                                                                        ,,     '? �
                                                                               Y:   -
                                                                    _.....
                                                                        -<,,            '
                                                 ../
Modem feeder barns provide a climate controlled environment, designed for the con fort of pigs as well as farm workers. A
proper environment is required if feeder barn goals are to be achieved.
                                                   -=---��������������������������
                                                                  200
Typical Diets
  There are a wide variety of options available to                  Table 9-12 contains diets that might be used in
feed growing-finishing pigs. Tables 9-12 and 9-13              areas where barley and wheat are most available.
present some alternative formulae for the various              Table 9-13 offers typical diets for areas where corn is
types of pigs.                                                 the predominant grain.
2 3 4 5 6
 Ingredients, %
  Wheat                                                      50.00          50.00        60.00         55.35      60.00
  Barley                                       73.23         23.10          21.70         7.50                    16.72
  Peas                                                                                                 20.00
  Soybean meal - 47%                           11.70                        16.90        18.50         10.35       17.60
  Soybean meal - 44%                                           11.8
  Canola meal                                  10.00          10.00          7.40        9.00          9.30
  Fat/oil                                       1.00           l.00                      1.00          l.00        1.40
  Lysine HCl                                    0.07           0.10                                                0.18
  Threonine                                                                                                        0.10
  Premix                                        4.00          4.00           4.00        4.00          4.00        4.00
 Nutrients, minimum %
  D.E., kcal/kg                                3,100          3,100         3,250        3,350         3,350      3,350
  Crude protein                                 17.2           18.3          19.9         21.l          20.6       18.5
  Digestible lysine                             0.73           0.73          0.78         0.84          0.84       0.84
  Digestible methionine                         0.20           0.20          0.22         0.23          0.23       0.23
  Digestible TS.A.A.                            0.42           0.42          0.47         0.52          0.52       0.52
  Digestible threonine                          0.45           0.45          0.51         0.56          0.56       0.56
  Digestible Tryptophan                         0.13           0.13          0.15         0.16          0.16       0.16
  Calcium                                       0.75           0.75          0.75         0.75          0.75       0.75
  Phosphorus                                    0.65           0.65          0.65         0.65          0.65       0.65
  Sodium                                        0.15           0.15          0.15         0.15          0.15       0.15
  Chloride                                      0.15           0.15          0.15         0.15          0.15       0.15
These are sample diets only and are for illustrative purposes only. While every attempt has been made to present exam-
pies that reflect successful commercial formulations, these examples are not intended for actual use without assistance
from a qualified nutritionist.
All amino acid concentrations arc expressed as apparent ilea! digestible amino acids.
                                                              201
Table 9-13. Examples of Growout Diets Based on Corn.
                                                                        2                3         4             5
  Ingredients, %
   Corn                                               69.25          70.18          64.30       40.70          47.10
   Wheat                                                                                        30.00
   Peas                                                                                                        25.00
   Soybean meal - 47%                                 26.75                         27.80       23.50          13.85
   Soybean meal - 44%                                                24.65
   Canola meal                                                                                                 7.50
   Lysine HCI                                                         0.10
   Fat/oil                                                            1.05          3.90         l.80          2.55
   Threonine                                                          0.02
   Premix                                               4.0            4.0              4.0       4.0           4.0
  Nutrients, minimum %
   D.E., kcal/kg                                      3,440          3,450         3,650        3,500          3,500
   Crude protein                                      18.60          17.00         18.70        18.67          19.30
   Dig. lysine                                         0.84           0.79          0.86         0.79           0.79
   Dig. methionine                                     0.23           0.22          0.24         0.22           0.22
   Dig. T.S.A.A.                                       0.52           0.46          0.50         0.47           0.47
   Dig. threonine                                      0.56           0.49         0.53          0.51           0.51
   Dig. Tryptophan                                     0.16           0.14         0.15          0.15           0.15
   Calcium                                             0.75           0.75          0.75         0.75           0.75
   Phosphorus                                          0.65           0.65          0.65         0.65           0.65
   Sodium                                              0.15           0.15         0.15          0.15           0.15
   Chloride                                            0.15          0.15          0.15          0.15           0.15
These are sample diets only and are for illustrative purposes only. While every attempt bas been made to present exam-
pies that reflect successful commercial formulations, these examples are not intended for actual use without assistance
from a qualified nutritionist.
All amino acid concentrations are expressed as apparent ilea! digestible amino acids.
     All diets will support a reasonable level of per-            Formulations employing commercial supplements
formance. These formulations are examples only and              are not included in the tables because their nutrient
many other combinations are possible. Due to the many           composition varies among suppliers. Combinations
factors that influence pig performance, results with these      of grains with supplements are possible and inter-
diets may vary from fa11TI to farm.                             ested persons are encouraged to contact their feed
  These formulations are only samples. Exact                    supplier for assistance to develop a balanced diet.
rations will depend on the 'actual' composition of the            No one diet formulation will be the best for all
ingredients on hand, which can be determined only               farms under all economic conditions. Flexibility is
by having the feeds tested. Rations used on indi-               the key to success in attaining the most value from
vidual farms may vary from these examples and                   your feed dollar.
should be formulated by a qualified nutritionist.
                                                          202
Feed Additives                                                    A situation where feed additives may be particu-
                                                               larly effective is when feeder pigs are just moved into
   A whole range of feed additives are available for           the grower-finisher units, especially when pigs arc
inclusion in feeder pig diets. They include enzymes,           commingled from different sources with varying
acidifiers, probiotics, feed flavours, medications,            health status. In these situation, pigs should be fed
toxic binders, etc .. Many of these are discussed in           palatable, highly digestible diets that arc hightly
chapters three and eleven. Most of these additives             fortified with vitamins and minerals and contain
are more effective in diets for weaner and starter pigs        appropriate levels of feed medication. The use of
and for sows than in diets for feeder pigs. This can           feed additives, such as acidifiers, may be considered
be attributed to the relative maturity of the feeder           as well. A practical means to ease the adjustment to
pig's digestive and immune system, and the lack of             the new environment is to feed a good quality pig
sudden changes in levels of production and feed                starter diet for the first week after the pigs have
intake (such as those around the time of farrowing).           arrived or until feed intake has increased to accept-
The effectiveness of feed additives also differs               able levels.
between individual production units, and in particular
with variation in the herd's health status and environ-          It should be stressed that feed additives should not
mental management. The average effects of the                  be used to compensate for poor management and that
various feed additives that are listed in Table 9 -14          feed medication, at the allowed levels, will be inef-
may differ substantially from their effects on indi-           fective in treating clinical levels of disease.
vidual pig production units. It is thus difficult to
make general statements about the cost-benefit
relationship of each of these feed additives. Produc-          Phase Feeding
ers that are considering the use of various feed
                                                                  Various options are available for feeding growing-
additives are encouraged to demand "proof', or the
results of on-farm studies, from the suppliers of these        finishing pigs. One option is phase feeding, in which
products. It should be noted that the use of many              more than one diet is fed to pigs between their arrival
feed additives, and of medications in particular, are          in the feeder barn and their attainment of market
under govenment control and subject to the Feeds               weight. Phase feeding allows producers to meet the
Act (see also chapter 10).                                     pig's nutrient requirements more closely at its various
Table 9-14. Pig Response to Antimicrobials in Feed of Growing-finishing Pigs (16 kg to market weight) (adapted from
Zimmerman, 1986).
                                                              203
live body weights. For example, a diet with rela-            Table 9-15. Estimated Value of a One versus Two
tively high DE and amino acid levels can be fed to           versus Three Phase Feeding Program in Pigs with
the grower pigs, and energy density and amino acid           Average Lean Grol, th Potential*.
levels can be reduced in the finishing diet. At the
same time, the supply of excessive nutrients can be                                                     Phase
reduced when phase feeding is applied (see Figure 9-
                                                                                             One         Two     Three
9). It should be noted that a large proportion of feed
used in the feeder barn is consumed by the finishing         Animal 12erfQrmance
pigs. Based on a typical feed intake and growth              Growth rate, g/d                0.772      0.784    0.789
curve and a three phase feeding program, feed usage          Feed: Gain                      3.28        3.24    3.22
between 25 and 45 kg body weight (grower I), 45
                                                             Carcass index                  106.20      106.30   106.30
and 70 kg body weight (grower II) and 70 and I 05
kg body weight (finisher) would be 18%, 27% and              Financial 12erformance
55% of total feed usage, respectively. This implies          Feed cost, S/pig                38.44      37.03    35.89
that close attention should be paid to the formulation       Gross margin, $/pig             48.85      50.30    51.55
and cost of the finisher diet and that producers can
afford to invest in high, nutrient dense grower I diets.     Gross margin, $/pig place/yr 156.23        163.14   168.16
                                                             ex
                                           Barrows                        Gilts             Difference
Initial weight, kg                           23.9                         24.4
Final weight, kg                             105.1                        104.1
Feed intake, kg/d                            2.42                         2.16                +11%
Gain, kg/d                                    .84                          .78                 +8%
Feed: Gain                                   2.87                         2.78                 +3%
Dressing percentage, %                       80.9                         80.3                +.7%
Carcass lean yield, %                        48.1                          50.5                -5%
"Calculated as performance of barrows - gilts divided by the average performance of the two sexes.
** 1993 carcass grading system.                              Source: Prairie Swine Centre Inc., I 993
                                                       204
barrows and gilts has been generated. As indicated                                               the daily feed allowance for barrows may be reduced
by the results in Table 9-16, barrows consume more                                               in the finishing phase during which barrows deposit
feed, grow faster, have poorer feed efficiency, and                                              large quantities of body fat. This reduction wi II
lower carcass lean yields than gilts. Generally, the                                             improve feed efficiency and carcass value slightly. If
differences in performance between these two sexes                                               you choose split-sex feeding as a feed option in your
increases as pigs grow heavier. Up to approximately                                              operation, keep in mind that difference in perform-
25 kg body weight, identifying any differences in                                                ance between the two sexes, and thus the optimum
performance between gilts and barrows is difficult,                                              feeding strategy, may vary somewhat with the differ-
but differences in feed intake and growth rate may be                                            ent pig breeds and genotypes. Monitoring feed
as high as 15% during the finishing phase of                                                     intake and performance in the two sexes is important
production. This is illustrated by the feed intake                                               whenever split-sex feeding is applied.
curves for Prairie Swine Centre barrows and gilts that
are presented in Figure 9-10.
                                                                                                 Table 9-17. Effect of Sex and Dietary Lysine Level on
                                                                                                 Performance in Finishing Pigs (50.7 to 104.6 kg Body
                                                                                                 Weight; NCR-42 Committee on Swine Nutrition 1993)
I �i : : : _. -- . -· .: - ·. -�                     : --· · ·: : : : :
       2.8 •• • •••• • ••• • • ••••••• • • •• • •••••••••••• • •••••       • • ••• • ••• • •
!
'\:S                                                            --··----                  ...
                                                                                                205
                                                                Photo 9-4.
  *   Feeding diets with a higher energy density to             There is no doubt that in the future barrows and gilts will be
  gilts than to barrows, especially in the finishing            fed separatley, some producers are already doing so.
  phase. In gilts, energy intake will limit lean growth
  up to higher body weights than in barrows.
                                                                meat derived from entire male pigs. Lean growth
                                                                potential are higher in entire males than in gilts.
                                                                Entire males require more nutrient-dense diets but
   * Slaughtering castrates at a lighter weight than            can utilize these diets more efficiently than gilts.
  gilts. Because gilts are leaner, carcass grades can
  be maintained at heavier weights. Barrows tend to
  deposit fat sooner and faster, therefore should be            Feeding to Appetite Versus Restricted
  marketed at a lighter weight.
                                                                Feeding
                                                                   Most pigs in Canada are fed to appetite, which
  *  Feeding a different daily feed allowance to each           means they are given continuous access to feed. The
  sex. Barrows' feed intake can be restricted to reduce         objective of this feeding method is to maximize feed
  the deposit of excessive fat in the carcass. Barrows          intake, thus growth rate. There are several benefits to
  grow faster than gilts but they also lay down fat             feeding to appetite. Carcass merit will suffer
  more quickly.                                                 somewhat when pigs are fed to appetite, but the
                                                                reduced value of a slightly fatter carcass is thought to
                                                                be more than off-set by a faster growth rate. Ad
                                                                libitum feeding is more easily automated than
  An additional advantage of managing the two                   restricted feeding so that labour costs are lower.
sexes separately is that barn space will be used more           Restricted feeding also has benefits, the major one
efficiently. Because barrows grow faster than gilts.            being that feed efficiency and carcass value may
growth rates within pens will be more uniform when              improve and growth rates may be reduced when feed
the two sexes are kept separately. The barrow pens              intake is reduced (see Figure 9- 7).
will empty more quickly so can be turned over faster
than the gilt pens. As a result, more pigs can be
produced per year using the same amount of space                    The results in Table 9-18 indicate that feed intake
when split-sex feeding is utilized.                             should be maximized when the production objective
                                                                is to maximize income per pig place per year. Yet, a
                                                                 l 0% reduction in feed intake is expected to increase
   The benefits of split-sex feeding will be increased          the income per pig. Efforts to improve carcass
when entire males, rather than barrows, are used in             quality in Canada have been directed largely al
commercial pork production. In North America,                   genetic selection and diet formulation. As the lean
entire males cannot (yet) be used for pork                      growth potential of pigs continue to increase due to
production. This situation may change as more                   genetic selection and improvements in health status,
information becomes available about the factors that            the potential benefits of restricted feeding will
cause the strong, adverse smell (boar taint) in some            decline.
                                                          206
Table 9-18. Estimated Effect of Level of Feed Intake on     feeder pigs were allowed continuous access to two
Animal and Financial Performance in Pigs with a             feeders with different feeds. One diet was formu-
Slightly Better than Average Lean Growth Potential*.        lated to contain high levels of available nutrients to
                                                            meet the highest possible nutrient requirements of
Level of feed intake      Average* Average - 10%            any pig in the pen. The second diet was formulated
                                                            with low available nutrient levels to meet the require-
                                                            ments of the pig with the lowest possible nutrient
Growth rate, g/d           823.00         729.00            requirements. The theory tested was that each pig
Feed: Gain                  2.97            2.88            would adjust its consumption of the two diets to meet
                                                            its nutrient requirements. Gilts would consume more
Carcass dressing %          79.70          79.20
                                                            of the high nutrient diet than barrows, and all the pigs
Carcass index               107.80         110.30           would consume relatively more of the low nutrient
Gross margin per pig($) 30.22              31.42            diet as they grew heavier and required fewer nutrients
                                                            per kg of feed. Unfortunately, the observations in
Gross margin                                                many studies were different from the expectations.
per pig place per year($) 98.50            92.50            The pigs tended to over-consume on the more expen-
                                                            sive ' hizh
                                                                     0
                                                                        nutrient dense diet. Plus, the choice of
                                                            diets was affected by the presence of small quantities
*Upper limit to lean growth is approximately 350 g/d.
Average feed intake is considered 90% of voluntary fe�d
                                                            of unpalatable ingredients in both diets. Clearly,
intake according to NRC 1987. 1993 Saskatchewan prices.     more research is needed before choice feeding can be
                                                            applied to commercial pork production units.
                                                           207
months because they are consuming more feed.                       Table 9-19. Performance Data Comparing Shelter and
Based on the observations at the University of                     Conventionally Raised Feeder Pigs During the
Manitoba, the dietary levels of amino acids, vitamins              Summer {May through August, Trial 1), Fall (August
and minerals may be reduced by approximately 8                     through November, Trial 2) and Winter (November
                                                                   through February, Trial 3). (University of Manitoba,
and 15% during the fall and winter months, respec-
                                                                   Courtesy of Dr. Connor)
tively, as compared with diets for pigs housed in
conventional indoor facilities. Despite the reduction
in the dietary levels, the daily allowance of these                                        Trial 1
nutrients should still be maintained somewhat higher                                       Shelter       Conventional
to account for the larger variation in feed intake and
                                                                   # of pigs                 175               150
feed wastage in pigs that are housed outside in large
groups.                                                            Initial weight, kg        23.9            28.20
                                                                   FinaJ weight, kg        100.80            I 01.00
                                                                   Feed intake, kg/d         3.05             2.98
Feed Separation                                                    Weight gain, kg/d         0.90             0.92
   Feed separation was discussed in detail in chapter              Feed/gain                 3.39             3.24
5. Table 9-20 summarizes the results of a Swedish
study reported by Dr. Ove Olsson. The study looked                 Carcass index           103.70            103.60
at three feed handling systems: I) mixed feed added
directly to 50 kg bags, 2) bulk feed with careful                                          Trial 2
handling, and 3) bulk feed with ordinary handling.                                         Shelter       Conventional
Study results show that separation decreased uniform
                                                                   # of pigs                 177               150
growth and feed efficiency. Separation is a problem
that costs money and should receive much more                      Initial weight, kg       32.50            34.10
attention than it does.                                            Final weight, kg        101.00            100.80
                                                                   Feed intake, kg/d         3.38             2.92
                                                                   Weight gain, kg/d        0.92              0.89
Photo 9-5.
                                                                   Feed/gain                3.67              3.28
                                                                   Carcass index           103.20            102.80
                                                                                           Trial 3
                                                                                           Shelter       Conventional
                                                                   # of pigs                 178               150
                                                                   Initial weight, kg       31.7              32.3
                                                                   Final weight, kg         100.5             101.1
                                                                   Feed intake, kg/d        3.62              3.19
                                                                   Weight gain, kg/d        0.90              0.96
                                                                   Feed/gain                4.02              3.32
                                                                   Carcass index           103.40             103.0
                                                             208
Table 9-20. Effect of Feed Handling on Pig Performance.
No. of pigs                                                  24                      24                    24
Initial weight, kg                                         20.500                 20.500                 20.500
Final weight, kg                                           97.000                 96.000                 96.300
Average daily gain, kg                                     0.580                   0.560                  0.550
Variation in weight gain    1
                                                           0.030                   0.038                  0.042
Feed efficiency                                            3.310                   3.440                  3.550
   Feed wastage is particularly high when pigs are             Feed form and feed processing are also related to
floor fed. Results from British studies (Table 9-21)         feed wastage. As illustrated by the results in Table 9-
clearly demonstrate that, even when feed intake in           22, feed efficiency is approximately 8% better in
                                                            209
pigs fed pelleted feeds as compared to pigs fed the          minimal labour, provides extremely useful
same diet but in a meal form. As there is little effect      information.
of pelleting on the digestibility of nutrients by
                                                                The importance of good financial and production
grower-finisher pigs (as is the case in starter pigs),
                                                             records is well established. The computer has helped
the difference in performance can largely be attrib-
                                                             many of us monitor cash flow, animal flow, and
uted to feed wastage. This implies that the effect of
                                                             production in a very sophisticated manner. One note
feed pelleting on feeder pig performance will depend
                                                             of caution: records from a computer are only as
on feeder design and feeder management; it will be
                                                             correct as the information entered into it and are only
smaller when feeders are well managed and when
                                                             as accurate as the program being used. Anyone using
feed wastage is already minimal. For further discus-
                                                             a program should learn how the calculations are
sions on feed processing see chapter I 0.
                                                             made so he or she knows how the final numbers are
                                                             produced. Very often the summaries make important
Table 9-22. The Effect of Feed Form on Performance           assumptions that can influence how the data is to be
of ad libitum Fed Pigs (35 to 87 kg body weight;             interpreted. If unaware of these assumptions, a
combined results of various studies; approximately           producer may misunderstand the information and
1000 pigs per treatment).                                    make unsound management decisions. Companies
                                        Feed form            offering record keeping systems are generally more
                                                             than willing to provide assistance in this regard.
                              Pellets          Meal
                                                               Compare your computer summaries with 'real'
Feed Disappearance (kg/d)* 2.29                 2.17
                                                             data. For example, if the computer says you are
Grun (kg/d)                    0.75             0.78         weaning 21 pigs per sow per year and you have a
Feed:Gain                                                    herd of I 00 sows, are you really weaning 2100 pigs
                               3.06             2.82
                                                             per year? Checks like this one will give you greater
*includes feed wastage                                       confidence in your computer results.
Derived from Walker. N. 1990; Pig News and Information
Vol. ll(l)pp.31-33.                                            A computer is not necessary to keep records,
                                                             though. If the number of pigs in a barn is reasonably
                                                             constant, you can estimate the days to market by
Monitoring Performance                                       measuring the rate of inventory turnover (if the
                                                             number of pigs in the barn fluctuates too much, the
   Once the 'correct' feeding regime has been se-            results will be misleading). To measure inventory
lected, it must be maintained. Performance must be           turnover, you must know the average number of pigs
monitored to see if goals are being achieved. If goals       on the farm throughout the year and the number of
are consistently not being achieved, solutions to            pigs sold per year, including gilts raised as replace-
existing problems must be found. If goals are being          ment breeding stock (see Table 9-23). Month-end
met, they should be re-evaluated and new ones set. If        inventories including nursing pigs, and weanling and
the goal is to maximize profit per pig, changes in           market hogs can be used for the calculation. Do not
feed prices relative to the price of pork may require        include the breeding herd in your calculation.
adjustments in the feeding program. If the objective
                                                                Surveys have shown that the average number of
in developing feeding programs is to meet the ani-
                                                             'days to market' is close to 200 days. The experi-
mals' nutrient requirements as closely as possible,
                                                             ence of some commercial producers, however,
nutrient levels in the feed should be adjusted with
                                                             indicates that 165 days for pigs to reach I 05 kg is a
changes in observed levels of feed intake.
                                                             realistic management objective. Better performance
  Monitoring feeder barn productivity is a very              ( 140 - 150 days) is possible with careful manage-
worthwhile activity. Full record keeping systems are         ment, all-in all-out housing, healthy stock, excellent
more comp I icatcd than those in the breeding and            diets. and maybe just a bit of luck! The data summa-
farrowing areas and may be daunting at first. Some           rized in Table 9-23 demonstrates that the average
type of monitoring, however, is important - even             number of days to market below 150 is possible
spot-checking of days-to-market or recording feed            under commercial conditions.
conversion on the odd random pen, which requires
                                                       210
Table 9-23. Calculation of Days to Market Based on Inventory Turnover.
Item l 2 3 4 5
Erem RecQrds
Avg. Inventory              325                    3215                 1256                   1230                 698
# of Pigs Sold/Year         728                    5883                 2135                   3038                 1459
Calculated
Inventory Turnover          2.24                   1.83                  1.70                  2.47                 2.09
Avg. Days to Market         163                     199                  215                    148                 175
                                                                  211
       Total weight gain= 204,650 - (2025 weanling x          ciency is required to estimate feed cost per pig. The
                20 kg/weanling)= 164,150 kg                   marginal feed efficiency is required to determine the
                                                              optimum shipping weight, i.e., the increase in feed cost
                                                              to raise pigs to heavier weights. For the development
           Feed conversion= 545,700 kg feed/year -
                                                              of a multi-phase feeding program, the estimated levels
           164, 150 kg weight gain in the feeder barn         of feed intakes can be used to determine the optimum
                  - 3.32 kg feed/kg gain                      dietary nutrient levels in diets at each individual phase.
                                                              Prairie Swine Centre Inc. has created a computerized
                                                              performance monitoring system that allows for the de-
   The major disadvantages of inventory-based                 velopment of a feed intake and growth curve based on
performance monitoring systemsare that they do not            a limited number of detailed observations on a selected
provide information on uniformity of growth and               number of pens. The program can also be used to esti-
flow of pigs through the barn and on performance at           mate feed cost and gross margins per pig or per pig
the various stages of growth. Since highly variable           place per year.
growth rates among pigs leads to reduced barn
                                                                                                                  Feed Intake Curuc
utilization, many producers use individual animal
                                                              1.0,--------------------.
tattoos to monitor average and ranges of days to                     • Obserued !Jat.a                                                                                                   e
                                                                     • • fl tted Intake Curue                                                                                            e
market. If there is too much variation in starting and               - Standard lntalce Curuo <NRCl                                                                                      d
                                                                                                                                                                                         y
keeping systems is to accurately monitor perform-             8.8'--------'------'---'--...;.__...___
                                                                 18 ze se i8 se 60 ?e
                                                                                                                                                                                         )
                                                                                      89 '8    109  110
ance in a limited nwnber of representative pens in the                                               Obserued Lluo Bocly Weight Ckgl
growing-finishing barn. Based on feed usage and
body weight gain in the monitor pens, performance
                                                                                                               An 1... 1 Grouth Cw-uc
in the rest of the barn can be estimated. If feed
disappearance and body weight gain is monitored at
                                                              110.9,----------------��
                                                                    • Obscrucd Do.ta .> •                                                              r
                                                                      •• Fitted - Curuoc                                                                                                 B
regular intervals, complete feed intake and growth                                                                                                                                       Q
                                                                                                                                                                                         d
                                                              85.8
curves can also be developed.                                                                                                                                                            y
                                                                                                                                                                    :
    The feed intake and growth curves, presented in fig-      60.0
                                                                             .-·-- -·,-
                                                                                      : ···'·                            -f           i                             fl
                                                                                                                          l I !
                                                                                                          ,r":.,,..,,.    ··-- !---•·•-•••,•••••·•···•-:·•
                                                                                                                                 00
                                                                                                                                                                                         g
ures 9-11 a and 9-1 lb, are derived from observations                                   l          .l--                                                             i                    h
                                                                                             »:"
                                                                                                                                                                                         t
on six different pens over a 14 day period. Observa-
tions were recorded for feed intake (based on feed dis-
                                                              35.8
                                                                             ,        /,·;          s,       ·l-··-··-i···- "j·1 ------!--··-·'t', - :! .                  .             (
                                                                      . ..              i                                                      : :                                       k
                                                                                                                              i
                                                              10.e�---'----------------�
and the number of days the pigs are in the barn. These               9           12     21         36         '18        60       72        111       'J6         108              120
                                                        212
 Table 9-24. Estimated Performance of Feeder Pigs Based on Feed Intake and Growth Curves Presented in
 Figure 9-11.
 Wt. Range         Marg.     Cumm.        Feed Int.    MargGain Cum. Gain           Marg.FCE Cum. FCE
  (kg)             Days       Days        (kg/day)      (g/day)  (g/day)              (g/g)    (gig)
 25 - 30             8.0        8.0         1.475          628.4        628.4          2.35          2.35
 30- 35              7.3       15.3         1.671          682.4        654.3          2.45          2.40
 35 - 40             6.9       22.2         1.868          727.9        677.1          2.57          2.45
 40-45               6.5       28.7         2.034          766.0        697.3          2.66          2.50
 45 - 50             6.3       35.0         2.184          797.5        715.3          2.74          2.55
 50 - 55             6.1       41.0         2.322          823.1        731.3          2.82          2.60
 55 - 60             5.9       47.0         2.447          843.5        745.4          2.90          2.64
 60- 65              5.8       52.8         2.562          859.0        758.0          2.98          2.68
 65 - 70             5.7       58.5         2.666          870.2        769.0          3.06          2.73
 70- 75              5.7       64.2         2.762          877.1        778.6          3.15          2.77
 75 - 80             5.7       69.9         2.848          880.3        786.9          3.24          2.81
 80- 85              5.7       75.6         2.927          879.9        793.8          3.33          2.85
 85 - 90             5.7       81.3         3.000          876.1        799.6          3.42          2.90
 90-95               5.8       87.0         3.066          869.2        804.2          3.53          2.94
 95 - I 00           5.8       92.9         3.126          859.2        807.7          3.64          2.99
 100 - 102           2.4       95.2         3.164          850.5        808.7          3.72          3.01
                                                             213
feeder design, and location, will also affect space re-      Table 9-25. Recommeded Pen Floor Space Allowances
quirements and performance of growing-finishing pigs.        for Growing Pigs*
If 15 finishing pigs are grouped in one pen, a pen size
of 4.3 m x 2.3 mis suggested. This size is based on the
                                                             Body weight        Fully slatted      Partial slats
amount of space required according to the Canadian
code of practice to accommodate 15 pigs at 95 kg body        kg      (lb)       m2      (sq ft)      m2 (sq ft)
weight. Given variations in animal performance, the
first pig is already shipped from the pen when the av-       25      (55)       .30      (3.2)      .33   (3.6)
erage body weight of all pigs in the pen reaches 95 kg.
                                                             50     ( 110)      .48      (5.2)      .53   (5.7)
   Controlling the barn temperature is also an impor-        75     (165)       .62      (6.7)      .70   (7.5)
tant practice. The effective barn temperature, which is
the combined effect of many factors, including air tem-      100    (220)       .76      (8.2)      .85   (9.1)
perature, air speed, air humidity, floor type, dryness of
floor, and the pig's body condition, all have a direct       *Recommended Code of Practice for the Care and Handling
effect on feed intake and animaJ performance. If the         ofFann Animals
effective environmental temperature is too cold, extra
feed is required to maintain a constant body tempera-
ture and feed intake will increase. If, on the other hand,   Additional Reading and References
the effective environmental temperature is too hot, feed     Agriculture and Agri-Food Canada Publicaiton 1898/
intake will quickly reduce and growth rates will de-         E. 1993. Recommended code of practice for the care
cline. The optimum temperature range, i.e., the tem-         and handling of farm animals - pigs. Communication
perature range in which pigs are neither too hot nor too     Branch, Agriculture Canada, Ottawa, ON, Canada.
cold, is relatively narrow, and will reduce with increases
                                                             Veterinary Infectious Disease Organization. 1987.
in live body weight. It is recommended that barn tem-
                                                             Feeder Barn Design and Management. Saskatoon, Sas-
peratures do not fall below 18 - 20°C for growing pigs.
                                                             katchewan. 48 pp.
Lower temperatures (l2°C) may be acceptable in all-
in, all-out barns when the pigs in a given room reach        Forbes, J.M., M.A. Varley and T.L.J. Lawrence (Ed.).
75 kg of body weight. As pigs approach market weight,        1989. The voluntary food intake of pigs. Occasional
a barn temperature of 12°C is quite acceptable if the        publication of the British Society of Animal Produc-
pigs are dry and free of drafts.                             tion. P.O. Box 3, Penicuik, Midlothian Eh26 ORZ,
                                                             Scotland.
   Other housing recommendations include the provi-
sion of one waterer for every 15 pigs. The scientific        Midwest Plan Service. 1983. Swine Housing and
basis for this recommendation is limited, so it should       Equipment Handbook. Iowa State University, Ames,
be interpreted as a useful guideline only. Water quality     IA. 112 pp.
and water flow rates should be monitored regularly. It
                                                             Miller, E.R., D.E. Ullrey and A.J. Lewis (Ed.). 1991.
is recommended that the flow rate for growing-finish-
                                                             Swine Nutrition. Butterworth-Heinemann, 80
ing pigs be between 1.0 and 1.5 litres per minute (0.22-
                                                             Montvale Avenue, Stoneham. MA 02180, USA. pp.
0.33 imperial gallon per minute). Consult the manu-
                                                             1-673.
facturer's recommendations for optimum waterer place-
ment.                                                        Moughan, P.J., M. W.A. Verstegen and M.I. Visser-
                                                             Reyneveld (Ed.). 1995. Modelling growth in the pig.
                                                             EAPP publication No. 78. Wageningcn Pers, P.O. Box
                                                             42, 6700 AA Wageningen, The Netherlands.
                                                             English, P.R., V.R. Fowler, S.Baxter and B. Smith.
                                                             1988. The Growing and Finishing Pig - Improving
                                                             Efficiency. Farming Press, Ipswich. 555 pp.
                                                             Zhang, Y. 1994. Swine building ventilation - a guide
                                                             for confinement swine housing in cold climates. Prai-
                                                             rie Swine Centre Inc. Saskatoon.
                                                       214
10. DIET PROCESSING AND DELIVERY
  Processing and delivery of feed to pigs is obvi-            3. To isolate specific parts of an ingredient.
ously a critical part of pork production. Whether            For example, oat hulls can be separated from oats
feeds are prepared at a commercial feed mill or              to produce oat groats for use in starter diets. The
mixed on the farm, proper processing and delivery            remaining oat hulls can be used in gestating sow
are absolutely essential to achieving high quality,          or cattle diets.
consistent feed mixtures for pig consumption.                 4. To improve handling. The objective of feed
Poorly processed feeds result in impaired feed               processing is to create a feed mixture that flows
utilization, poor performance and in some instances,         freely and does not separate. In some cases
serious health problems. Understanding the nature            producers may wish to increase bulk density to
of feed processing, the reasoning behind each                reduce storage and hauling costs. For example,
procedure and the needs of the pig are important to          pelleting has been found to increase the bulk
all pork producers, whether they buy prepared feeds          density of wheat shorts by 80%. Bulkiness of the
or manufacture their own diets on the farm.                  feedstuff can also reduce feed intake. By increas-
                                                             ing the physical density of the diet, daily nutrient
                                                             intake can be improved. For example, gut
There are several reasons for processing diets
                                                             capacity often limits the quantity of feed con-
before feeding them to swine:
                                                             sumed by the young pig. Therefore, physical
                                                             density of the feed will influence the young pig's
  1. To alter the physical form or particle size.            nutrient intake.
 For example, grains are ground to reduce particle
 size and thus improve their compatibility with               5. To improve palatability. Mixing unpalat-
 other ingredients in the diet.                              able, but necessary ingredients with those that are
                                                             more appealing to the pig, increases the intake of
 2. To improve nutrient availability. All grains
                                                             required nutrients over the amount that would be
 must be ground before being fed to swine if
                                                             eaten if ingredients were offered individually.
 nutrient availability is to be maximized. Studies
 at the University of Alberta found that lysine is            6. To preserve. Drying or treating high mois-
 12% more available to the pig from ground                   ture grains with organic acids improves the length
 wheat as compared to rolled wheat. The average              of time they can be stored.
 improvement in availability of all essential amino           7. To detoxify. Cleaning removes undesirable
 acids is more than 6%. Pelleting and extruding              weed seeds that may prove to be toxic to the pig.
 are also processes used to improve nutrient                 Researchers are currently looking at chemical
 availability.                                               detoxicants as a way to improve the feeding
                                                             value of moldy grains.
Photo 10-1.
                                                              8. To create a uniform final feed mixture.
                                                             This is particularly critical in feeds for young
                                                             pigs, who eat relatively small quantities offeed
                                                             per day. However, a proper feed mixture is
                                                             necessary for all classes of swine to ensure that
                                                             they achieve maximum performance. Research
                                                             has shown that pigs do not possess "nutritional
                                                             wisdom" to the extent that they will consume a
                                                             properly balanced diet. They will seek out
                                                             certain nutrients such as salt if required, but they
                                                             are incapable of balancing a complete diet. For
                                                             this reason, a nutritionally balanced mixed feed
                                                             must be offered to pigs. Jf the feed is not uni-
A modern commercial feed plant.                              form, daily nutrient intake could fall below
Photo courtesy of Federated Co-operatives Ltd.               requirement and thus impair performance.
                                                       215
  A large number of processing methods are avail-                Table 10-1. Effect of Fineness of Grind on the
able to the swine industry. They include mixing,                 Performance of Weanling Pigs Fed Barley-based Diets.
grinding, rolling, cracking, popping, extruding,
micronizing (pulverizing), roasting, dry pelleting,              Screen size, mm       3.2      4.8       Difference
steam pelleting, steam rolling and reconstituting. In            Particle size.urn    634       767
practice, only a few, including mixing, grinding,                Daily gain, g        386       367            +5.2%
extruding and steam pelleting, are commonly used.                Daily feed, g        653       653              nil
The selection of one or more processes will depend               Feed:gain            l.70      1.79           +5.0%
on the ingredients employed, the age of the pig
being fed and the cost/benefit relationship.                     Source: Adapted from Goodband et al., 1993.
� 30
                                                               Dietary particle size also affects sow performance
-"'
a> 20
                                                             (Table 10-3). While the smaller particle size im-
                                                             proved productivity, the incidence of ulcers was
s
Cl
                                                             also increased.
"O
� 10
....Q.
0
                                                             Photo 10-2.
E
-        0
             Dry Matter     Energy          Protein
                            •   34kg
                            •   70kg
                                                           216
Table 10-3. Effect of Particle Size on the Perform-         grinding if they are not replaced. Based on a report
ance of Lactating Sows Fed a Corn-based Diet.               by Pouteaux, 1988, screen wear is much more
                                                            serious than hammer wear (Table 10-5).
                               Particle size, um
                        400    600     900     1200           Other factors may also play a role in the quality of
Litter size, d2 l       8.9    9.4     9.0     9.0          grinding. These include the speed of the mill, the
Sow wt. loss, kg        8.0    6.8     8.5     3.7          design of the hammers and screens, the ingredient
Sow backfat                                                 transporting system and the physical characteristics
loss, mm                3.3    3.8     4.6     4.1          of the grains, most important of which is moisture.
Litter weight, kg       50.4   50.4    48.9    46.9         Broad recommendations suggesting a specific
Feed intake, kg         4.4    4.3     4.2     4.2          screen size must be interpreted with care since a
Diet digestibility, %   88.6   86.7    85.4    84.4         universal guide is not possible. This is graphically
                                                            shown in Table l 0-5. The same screen size and
Source: Adapted from Goodband et al., 1993.                 hammer speed were employed for both the hammer
                                                            and screen studies. Only the brand of hammer mill
                                                            differed. Nevertheless, mean particle diameter,
  In addition to the potential for increased ulcers,        particle surface area and particles per gram varied
there are other down sides to seeking a more finely         significantly, even with new equipment. The best
ground diet: mill output is reduced, the power              alternative is to determine mean particle size on
demands per tonne of diet increase (Table I 0-4 ),          each diet and adjust mills accordingly. Mean
palatability may suffer and dustiness will increase.        particle size for market hogs and sows should be
Another consideration is shrink, a factor often             650 to 750µ.m.
ignored when calculating the true cost of manufac-
turing feeds. It refers to material lost as dust and
moisture and is believed to increase in parallel with
                                                            Photo 10-3.
fineness. Although shrink is difficult to quantify,
most estimates range from one to three percent.
                                                          217
Table 10-5. Effect of Screen and Hammer Wear on Grinding Effectiveness
  Barley
  New Screen                           692                          229                     118,200
  Worn Screen                          388                          419                     311,400
  (Difference)                       (l .78x)                    ( l .83x)                  (2.46x)
  New Hammers                          633                         274                      180,700
  Worn Hammers                         627                         250                      124,100
  (Difference)                       (l.Olx)                     (1.1 Ox)                   (l .46x)
  Wheat (HRS)
  New Screen                           772                         186                       73,290
  Worn Screen                          358                         427                      938,200
  (Difference)                       (2.16x)                     (2.30x)                    (12.8x)
  Wheat (Winter)
  New Hammers                          443                         390                     2.319,500
  Worn Hammers                         433                         392                     1,841,900
  (Difference)                       (l.02x)                       (--)                      (l.26x)
x: The number of times the smaller value exceeds the larger value in a column.
All studies were conducted with a 7/64" (2.78mm) screen. Both the screen and hammer study employed a 100 HP
tear-drop circular hammer mill operated at 3600 RPM. The mill used for the screen study though, was a different
brand than that used for the hammer study.
Adapted from Pouteaux, 1988. Proc. Alberta Pork Congress. Red Deer, pp.20.
  The most commonly recommended screen sizes              analyses of particle size, therefore visual inspection
for swine diets based on barley range from 3.0 to         of the ground feed remains important. In terms of
3.5 mm. Often. producers should use these as              final particle size, screen size is Jess important for
guidelines only, considering them with the other          barley than for wheat, as shown in Table I 0-5.
variables involved in manufacturing swine diets
such as moisture content, nature of the grain and the       As introduced previously. the incidence of gastric
feed mill itself. Producers do not have access to         ulcers tends to increase with the use of finely
                                                          ground feeds (Table I 0-6 and I 0- 7).
                                                        218
Table 10-6. Effect of Fineness of Grind of Corn on Incidence of Ulcers and Other Stomach Lesions in Swine
Adapted from Wu and Allee. 1984. Kansas State University Swine Day Progress Report pp. 83-88.
Table 10-7. Effect of Fineness of Grind of Barley on            It should also be noted that as farms move to
the Incidence of Ulcers and Other Stomach Lesions            phase feeding, where nutrient supply is closely
in Swine.
                                                             linked to nutrient requirements, the importance of
                                                             diet uniformity will be much greater; this is due to
                    Hammermill Screen Size, (mm)             the fact that single phase diets, for example, tend to
                         3.2              6.4                provide nutrients in excess of requirement and thus
Fineness of Grind                                            provide some degree of protection from poor diet
  Mean Particle Size                                         mixing.
  (microns)                 71 l               1159
  Surface Area (cm2/gm)      79                50              In all cases, the variation of the nutrient within the
Stomach Lesions(%)                                           batch is expressed as the coefficient of variation
  Normal Stomachs            17                67            (C.V.). In general, the objective of most mixing
  Comifications              33                17            systems is to reduce nutrient variability within a mix
  Erosions                    0                17            to a C.V. of less than 10% and certainly no greater
  Slight Ulcers               0                0             than 15%. Excessive variability within a mix
  Serious Ulcers              0                0             reduces growth rate and feed efficiency (Tables I 0-
                                                             8 and 10-9).
Adapted from Goodband 1986. Master's Thesis. Kansas
State University, Manhattan, KS.
                                                        219
  While proper mix time is required to achieve              Table 10-9. Impact of Mixing Time on Diet Uniform-
adequate mixing, worn or poorly adjusted parts may          ity and Performance of Finishing Swine.
also contribute to a poor mix. Operators must be
careful not to exceed the recommended capacity of                                         Mixing time, min
the mixer; otherwise poor mixing efficiency is                                       0      0.5     2.0    4.0
almost a certainty.                                         Coefficient
                                                            of variation           53.8      14.8     12.5    9.6
   There are two general types of mixers: horizontal        Daily gain, g           776      808      794     785
and vertical. The following comments are generali-          Daily feed, g          2945     2904     2886    2881
ties only; differences will exist among manufactur-         Feed conversion        3.80     3.60     3.63    3.67
ers due to variation in design. Typically, horizontal
mixers are more expensive to purchase than vertical         Source: Adapted from Traylor et al., 1994.
mixers but have a lower maintenance cost, in part
        '
because they have a slower ribbon speed of rotation
(30 to 40 rpm vs. 200 to 300 rpm). Also, horizon-           Pelleting
tal mixers can generally handle higher quantities of          Swine diets are pelleted for handling reasons and
added liquids than vertical mixers.                         to improve performance. The increased feed
                                                            efficiency is believed to be due to reduced wastage
    Horizontal mixers may employ either paddles or          and perhaps improved digestibility. A survey of
ribbons, or a combination of the two, circulating           117 experiments showed an average increase in
within the batch to achieve a uniform mix. Typical          growth rate of 6.6% and an improvement in feed
mixing times for horizontal mixers range from 3 to          efficiency of7.9% due to pelleting. The benefit
6 minutes, while for vertical mixers, a somewhat            depends on the nature of the ingredients used.
longer period- 8 to 15 minutes - may be required to
bring the coefficient of variation below the desired          Generally, improvement in performance is greater
 I 0%. Portable grinder mixers are a version of the         with more fibrous feeds. Therefore greater benefits
vertical mixer described above. Studies in which            from pelleting would be expected from barley than
mixing efficiency has been properly evaluated               from wheat or com. One study found that the
suggest that for a mixer in good repair, mixing for         advantage to pelleting corn in terms of feed effi-
 I 2-17 minutes after the addition of the last ingredi-     ciency was approximately 11 % while improvement
ent should suffice.                                         in growth rate ranged from 4-9%, depending on the
                                                            age of the pig; this was confirmed by a recent report
  The proper order of addition of ingredients is also       from Kansas State University (Table 10-10).
important. For best results, one-half to two-thirds
of the major ingredient (eg. barley) should be added          A survey of l O experiments using barley-based
first. Intermediate ingredients, such as supplement,        diets suggested that on average, pelleting improved
soybean meal or canola meal can then be added,              growth rate and feed efficiency by 15%. On bal-
followed by the minor ingredients such as premix or         ance this appears to be high, but improvement in the
salt. Finally, the remainder of the major ingredient        range of7-9% would not be unexpected. For
can be added.                                               example, Bell and Keith reported an average 4.7%
                                                            improvement in growing pigs and an average 12.6%
                                                            improvement in finishing pigs fed barley- or barley
                                                            and wheat-based diets (Table I 0-11 ).
                                                          220
Table 10-10. Impact of Pelleting Com-based Diets on Finishing Pig Performance, Nutrient Digestibility and
the Incidence of Ulcers.
'Lesions and degree ofkeratinization in easophageal region of the stomach scored on a scale ofO to 3, with 3 being
the most severe.
Source: Adapted from Wondra et al.. 1992a.
                                                         221
Table 10-1 J. Effect of Pelleting Diets Based on Canela (seed and/or meal) and Cereal Grains (barley or barley
plus wheat) on Pig Performance
Source: Adapted from Bell and Keith, 1991. Annual Report, Prairie Swine Centre, Saskatoon, SK. pp. 21-24.
  Pellet quality is a subject that attracts a great deal      Although many factors are believed to be in-
of attention in feed manufacturing and probably             volved. gastric ulcers tend to increase when pigs are
receives more attention than is necessary if a pro-         fed pelleted diets. Part of the problem may be
ducer's major concern is pig performance. Pellet            related to the fineness of grind used to prepare
durability is heavily influenced by diet composition.       grains for pelleting. A finer grind results in better
Wheat in particular is a good binding agent, while          quality pellets; consequently, pellet-mill operators
pellets made of com do not hold together as well.           tend to use finely ground grains in an attempt to
When pigs have access to pellets and fines, they            minimize customer complaints. With a coarser
appear to prefer the pellets and let the fines collect      grind. pellet quality will suffer but the incidence of
in the corner of the feeder. This prompts concern           ulcers will be reduced. The heating process associ-
on the part of the producer. Recent studies at              ated with pelleting results in gelatinization of the
Kansas State University reveal that the presence of         grain starches and has also been implicated as a
fines has little effect on growth rate, but reduces         cause of ulcers. In the example presented in Table
feed efficiency (Tables 10-12 and 10-13).                   10-10, the grains used in both the meal and the
                                                            pelleted diets were ground to the same particle size.
Table 10-12. Effect of Pellet Fines on Performance of Finishing Pigs Fed a Corned-based Diet (Initial wt.= 54kg).
                                                         222
Table 10-13. Effect of Pelleting and Fines on Performance of Newly-weaned Pigs Fed Corn-based Diets (Initial
wt.= 5. 7 kg).
d7 to d35
Ave. gain, kg/d                             0.47                     0.49                     0.49
Ave. feed intake, kg/d                      0.78                     0.73                     0.76
Feed conversion1•3                          1.67                     1.50                     l.54
I
  Effect of pelleting significant, P<0.01
2
  Effect of fines significant, P<0.05
3
  Effect of fines significant, P<0.07
Source: Stark et al., 1993.
Liquid Feeding
  Liquid feeding was popular many years ago but              True liquid feeding systems are also appearing on
fell into disfavour for a variety of reasons. Most of     the Canadian market. The new systems offer more
the reasons were related to problems with the             precision in mixing and delivery and if they are
mechanical equipment. New approaches have                 linked to computer systems, they provide a much
rekindled interest.                                       greater degree of control than was previously
                                                          possible. It is too early to offer a definite evalua-
  For example, one "version" of liquid feeding is         tion of these newer units, but they do offer consid-
the use of "wet-dry" feeders, a compromise                erable advantages over previous liquid feeding
between liquid and dry feeding. Feed is delivered         systems. Small improvements in pig performance
to the self-feeder in dry form, but a nipple waterer      can be expected with such feeding systems. The
located below the feeding platform allows the pig to      main drawback is cost and the difficulty of adding
drink water at the same time as it is eating. The pig     antibiotics to certain diets without contaminating
can even mix the dry feed with water in the dish          others.
located below both the feeding platform and the
waterer.                                                  Photo 10-5.
                                                        223
   Liquid or paste feeding has shown some benefits         Photo 10-6.
for the young weanling pig. Liquid feeding refers
to a feed with a high water content (2: I water:feed
ratio), while paste feeding is drier but still moist
(J .3-1.5: 1 water:feed ratio). Young pigs tend to eat
wet feed more readily. The problem is maintaining
feed freshness. Starter diets are high in milk prod-
ucts and tend to develop off-flavours very quickly.
However, if suitable antioxidants arc included in the
diet and the feed delivery system is managed
properly, these concerns can be largely overcome.
The key to liquid or paste feeding systems is to
provide fresh feed on a regular basis.
Drying Grains
  Depending on cropping practices and weather
conditions, grains may be dried to reduce moisture         Grain dryers are used to reduce the moisture content
content and thus prevent spoilage during storage.          of wet grains thus preventing spoilage.
An alternative to drying is to store the product as a
high moisture grain. This method is discussed later
in the chapter. Research has shown that corn can be           Sealed, oxygen-limiting silos are the most
dried to 12-15% moisture using temperatures of             convenient systems for storing high-moisture grains,
up to 110°C with no adverse affects on pig                 but they are also the most expensive. Vertical
performance. Temperatures in excess of 150°C               concrete silos or horizontal silos can be used
have been shown to affect the palatability of corn         successfully, although the level of management
to the pig.                                                required is greater than with the oxygen-limiting
                                                           systems. Mechanization of feeding is also more
                                                           difficult. especially in the case of the horizontal
High Moisture Grains                                       silos.
  High moisture storage and feeding of grain is
popular in areas where grains are harvested wet and          The grain should be ground and well packed in
must then be dried before conventional storage.            the silo to eliminate oxygen. All doors should be
High moisture storage involves the use of either           sealed; unsealed systems result in losses of 2-5%
oxygen limiting structures or the addition of              compared to oxygen limiting systems. Once a
acidifying compounds. Both systems inhibit                 vertical silo is opened, a minimum of three inches
undesirable microbial deterioration, allowing the          must be removed per day to prevent spoilage. This
wet grain to be stored until fed. The systems add to       may have to be increased during warm weather.
the cost of storage but save on the cost of drying.
                                                             On a dry matter basis, there is very little
  The use of high moisture grain can improve               difference in the feeding value of com or barley of
harvesting management, since a dryer tends to slow         equal quality stored in an oxygen limiting unit
down the grain handling process. High moisture             compared to being conventionally dried. High
grains stored in silos should normally fall within the     moisture storage appears to increase the availability
range of 22-28% moisture. This range limits the            of phosphorous in the grain but it may also reduce
period of time for harvesting but provides ideal           the amounts of vitamins A and E. The main point
moisture to generate the desired level of                  to keep in mind is that rations must be adjusted to
fermentation.                                              compensate for the extra moisture content of the
                                                           grain.
                                                         224
  Organic acids have been used extensively as                [fa proportioner-type mill is being used, the
preservatives for high moisture grains. The                bushel weight should also be measured. Bushel
different types include propionic, acetic, isobutyric,     weight should be checked on a regular basis
formic and benzoic acids or their various                  because changes in bushel weight will alter mixing
combinations. These acids preserve the grain by            accuracy. Appendix II outlines the steps required to
preventing mold growth and by killing the grain            calibrate a typical proportioner-type mill, one of the
germ. Organic acids can be corrosive to any metal          common mixing systems.
they come in contact with such as transportation and
storage equipment. If metal granaries are to be               It has been suggested that the moisture content of
used, they must be lined with a protective covering        the ingredients will influence the rate of addition in
to avoid premature rusting.                                proportioner-type mills. This is because moisture
                                                           content affects bushel weight. However, moisture
  Acid preservation systems are more flexible than         tests from a feed testing laboratory may not be
the use of oxygen limiting silos. The advantage is         completely accurate. This is because the moisture
that continuous feeding is not necessary. The acid         content of a sample of grain can change a great deal
remains with the grain until feeding: therefore            from the time it is sampled at the farm until it is
spoilage outside the granary is eliminated.                finally measured in the lab. This problem is
                                                           relatively minor for dry grains, but increases as
  Pigs normally perform very well on high-moisture         actual moisture content increases. Researchers go
grain. Dustiness is reduced so the wet grain tends to      to great pains to measure moisture accurately under
be very palatable. It should not be used for the very      laboratory conditions that would not be practical for
young pig though, due to the high moisture content         farmers. Consequently, a good bushel weight is
of the grain and the limited gut capacity of the small     likely to be the best overall measurement on a grain
pig. Once high moisture grain is harvested and             sample, to determine if mill re-calibration is
stored, there tends to be very little resale market        required.
available. The owner is generally committed to
feeding it to livestock.                                     For cereal grains, calcium and phosphorus
                                                           analysis may not be required, as they are quite
                                                           consistent from year to year and because they
Mixing Feeds on the Farm                                   contribute relatively small proportions of the total
  In Canada, a high but declining proportion of
feeds are manufactured on the farm. There are              Photo 10-7.
many indications though, that many home-
maufactured diets are not well mixed. Toe results
of surveys in several provinces indicate that up to
75% of feeds submitted to feed testing laboratories
for analysis fail to meet the nutrient requirements of
the pigs being fed. The problem can occur at any
number of the steps involved in feed manufacturing:
diet formulation, feed mixing or feed delivery. A
proper quality control program must be in place to
ensure diet quality and consistency. A system for
quality control was discussed in detail in Chapter 5.
Ingredients grown on the farm should be analysed at
least for crude protein and moisture.
                                                         225
quantities in the diet. Purchased ingredients should       Table 10-14. Separation of Feeds Following Mixing on
be analysed for protein, calcium, phosphorus and           the Farm.
salt on a regular basis, perhaps every 2-3 months, to
ensure that the assumed nutrient content is indeed                               Portion of Storage Tank
correct.                                                   Nutrient          Start       Middle         End
                                                                                           (%)
  Mixed diets should be sampled monthly and                Protein           17.60        17.40        15.70
analysed as required. This step ensures that mixing        Calciwn            1.09         0.99         0.74
and formulation steps have been completed                  Phosphorus         0.95         0.85         0.65
correctly, and provides a final check on the quality       Fibre              3.80         4.20         9.00
of the feeds being offered to the pigs. Refer to           Salt               0.74         0.71         0.61
Chapter 5 for a recommended feed sampling and                                      (parts per million)
analysis schedule.                                         Iron               310          200          200
                                                           Zinc               260          140           140
Processing Problems                                        Manganese           50           40           40
  Achieving a proper mix on the farm is not easy,          Copper              23           20           20
based on the results of surveys showing generally
poor quality control. Surveys continually reveal           A pre-grower diet was mixed using a proportioner-type
                                                           mill and stored in a hopper-bottomed bin before feeding.
that diets lack nutrient balance and often uniformity.     Feed samples were collected at three stages of emptying:
                                                           beginning, middle and end of the batch.
  One major problem is separation. For example,
Table l 0-14 summarizes the results of a study
conducted on a commercial farm in Saskatchewan             proven beneficial in removing nuisance dust, but is
using a typical proportioner-type mill. Feed was           very poor at controlling respirable dust - that which
mixed in batches and stored in a hopper-bottomed           is small enough to enter the lungs and thus poten-
(centre flow) steel bin. Feed samples were                 tially contribute to respiratory problems. Including
collected for analysis at the beginning, middle and        whole seed canola or soybeans in the formula is one
end of the batch as it left the storage bin.               way of accomplishing this objective and has the
Considerable separation was taking place, such that        added benefit of increasing the energy content of
pigs were receiving diets of varied composition,           the diet. Sprinkling oil directly in the barn is much
depending on the location of the feed within the           more effective in reducing dust - by up to 80% -
storage tank. Since this diet was formulated to be a       and at less cost than adding the oil to the diet.
pre-grower, the very high crude fibre and low
protein level found in the latter part of the batch    Care must be taken to increase amino acid
was a serious cause for concern.                     (protein) content of the diet when any fat source is
                                                     used; otherwise, poor carcass grades may result.
  To improve uniformity, dispersion tubes should     For every l % vegetable fat added to the diet, the
be placed inside feed bins to prevent segregation of limiting amino acids should be increased by about
ground feeds. Coarse grinding may also help, but     4%. For example, for every l % of fat added to a
losses in performance must be avoided. Pelleting of finisher diet, lysine should rise by about 0.03
course would solve the problem, but is generally not percentage units (eg. lysine increased from 0.75%
available on most farms.                             to 0.78%). This can be accomplished most easily
                                                     by increasing the amount of protein by 4% (i.e.
  Dustiness is another problem. Recognition that     crude protein increased from 16.5% to 17.2%).
dust in the barn poses a threat to the health of     When whole seed canola or soybeans are used, they
humans as well as pigs has focused greater attention add protein as well as fat, so the increase in protein
on this subject. Adding 0.5-2% fat to the diet has   per l % fat would be 1.5-2.0%.
                                                         226
Feeds Act and Regulation                                        2. When livestock diets are fed to animals
   Everyone manufacturing feeds, whether they are a            resulting human food products, such as meat or
commercial feed company or a private farmer                    milk, do not pose any kind of health threat to
                                                               persons consuming them. For example, the Act
mixing feed for his own use, should be familiar
                                                               specifically forbids the use of in-feed drugs in
with three publications: the Feeds Act ( 1976). the
                                                               such kind or quantity that would result in residue
Feeds Regulations ( 1983, revised) and the Com-                in the meat intended for human consumption.
pendium of Medicating Ingredient Brochures
(CMIB). These are federal statutes and guides                   3. Animal feeds do not pose a health threat to
governing the manufacture and use of livestock                 the animals consuming them.
feeds in Canada. Copies of all three can be ob-                 4. Animal feeds do not pose a threat to the
tained from the Federal Government. They detail                environment.
the legal responsibilities associated with the produc-
tion of feeds intended for use in livestock opera-            The Food Production and Inspection Branch
tions; contrary to common thinking, these rules            administers the Feeds Act and Feeds Regulations.
apply equally to both commercial feed companies            As a consequence of changes in the nature of the
and farmers manufacturing their own feeds. How-            livestock feed industry, and in their view, to obtain
ever, the Feeds Act states that it does not apply to a     the best results from a limited budget, they are
feed:                                                      focusing less attention on random visits to commer-
                                                           cial feed mills, a major activity in the past, and
 "that is manufactured by a livestock producer if it       spending more time following up on suspected
 is not offered for sale and has not had incorpo-          abuses of in-feed drug usage at commercial mills or
 rated into it any drug or other substance that may        on livestock farms.
 adversely affect human health or the environment
 or that is sold by the individual grower thereof if
                                                             At the present time, all feeds imported,
 it is free from prescribed deleterious substance�."
                                                           manufactured or sold in Canada must be registered
                                                           with the Federal Government, unless they are
  Simply stated, if the individual pork producer is
mixing his own diets for his own use and is not            s�eci�ically exempted by the Feeds Act; fortunately,
                                                           since rt takes considerable time and effort (and
adding anything to the feed that will be harmful to
                                                           S95.00) to register a single feed, the majority of
human health or the environment, then he or she is
                                                           feeds are exempted. For example, a feed is
not bound by the legislation contained in the Act.
                                                           exempted from registration if it does not contain
However. if feed medications are used such as
                                                           any medicating ingredients and meets specified
antibiotic-type growth promotants, then the Feeds
                                                           minimum requirements with respect to nutrient
Act must be adhered to.
                                                           composition. These nutrient specifications are
                                                           outlined in Table 4, Schedule I of the Feeds
  The following discussion summarizes general
                                                           Regulations. Table 10- 7 summarizes the
points of interest to livestock producers. It is not
                                                           information as it relates to swine. It must be
intended to be comprehensive; anyone interested in
                                                           recognized that Table 4 of the Regulations is
answers to specific questions should consult the Act
                                                           continually being reviewed and adjusted; therefore,
or contact their nearest Agriculture Canada office.
                                                           the reader is cautioned that changes to the
                                                           Regulations may have occurred since this book was
  Essentially, the objectives of the Feeds Act are to
                                                           written and that Table 10-7 may not be completely
ensure that:
                                                           up to date.
  I. All animal feeds are efficacious in terms of
 the original purposes for which they were                   As indicated, registration requirements do not
 intended. For example. a sow lactation diet               apply if the feeds are manufactured for a livestock
 should support milk production in the sow.                producer's own use and do not contain medications;
                                                           similarly, if the feed contains medications added at
                                                           levels defined in the CMIB, the registration
                                                         227
requirements are again waived. The CMIB defines                     It is clear that the government wishes to be very
which drugs are permitted to be used in swine diets,             careful about permitting the use of medications in
what levels may legally be added to the diet and                 livestock feeds. Anyone mixing diets which include
identifies restrictions on their use, such as                    medications may be required to submit to an on-
withdrawal times. Levels of drugs or combinations                farm inspection. The Act discusses such inspec-
of drugs, not included in the CMIB cannot be used                tions of feed mixing facilities. They are intended to
in swine diets unless the producer or feed                       ensure that, if medications are going to be used,
manufacturer receives a veterinary prescription.                 they are mixed adequately and safely. In this
                                                                 respect, the on-farm feed mixing facility is treated
  Agriculture Canada is primarily concerned with                 exactly the same as a large commercial feed mill.
diets that contain medications or other substances               Such an inspection could include a listing of all
that pose a potential hazard to animal or human                  medicating ingredients used and determine if such a
health and if not used properly could result in                  premix is approved for use, a check of feed han-
carcass residue. They are also concerned about the               dling, delivery and mixing facilities to ensure they
use of other substances that may be harmful to the               are in good repair and adequate for the job, and an
environment such as high levels of copper, which                 evaluation of manufacturing practices such as those
accumulate in manure and become concentrated on                  related to preventing inadvertent contamination of
land where it is spread.                                         other, un-medicated feeds mixed at the same site.
Table 10-15. Nutrient Guarantees Required for Swine Diets that are Exempt from Registration Requirements.
Minimum and maximum nutrient composition specified in the Feed Regulations for unregistered diets fed to various
classes of swine:( I) 1- l O kg bodyweight; (2)1actation; (3 )breeding; (4 )all classes not specified; (5)up to 20 kg
bodyweight; (6)20 kg to market; (7)up to 60 kg; (NRS)No requirement specified. Diets offered for sale which fall
within these ranges need not be registered. The limitations specified in this table do not apply to individual livestock
producers who are exempt from these regulations (see above).
                                                           228
  Clearly, the use of medications in livestock diets      Additional Reading and References
is considered a serious matter which must be              Goodband, R.D., 1986. The effects of barley on
carefully controlled to ensure that the pork industry     starter and finishing pig performance. Master's
produces a safe, wholesome product. It is the             Thesis, Kansas State University, Manhattan, KS.
responsibility of each individual producer to ensure
that proper procedures are followed.                      Goodband, R.D., M.D. Tokach and J.L. Nelssen.
                                                          1993. Diet particle size influences pig
  Because they are in the business of selling feeds,      performance. Proc. Manitoba Swine Seminar,
the commercial feed manufacturer is expected to           Winnipeg, MB. pp. 91 - 104.
meet certain standards beyond those applying to the
individual farmer. All feeds must be carefully            Stark, C.R., K.C. Behnke, J.D. Hancock, and R.H.
labelled. Although the Regulations carefully define       Hines. 1993. Pellet quality affects growth
labelling requirements, the major objective is to         performance of nursery and finishing pigs. Swine
provide the buyer with sufficient information to use      Day Progress Report pp. 67-70.
the feed effectively and safely. Labels include
feeding instructions, withdrawal times if                 Traylor, S.L., J.D. Hancock, K.C. Behnke, C.R.
medications are involved and cautions with respect        Stark, and R.H. Hines. 1994. Mix time affects diet
to proper use. All diets manufactured for sale by         uniformity and growth performance of nursery and
feed companies that contain nutrients outside the         finishing pigs. Swine Day Progress Report pp. 171-
bounds defined in Table 4 of the Feeds Regulations        175.
must be registered with the Federal Government.
An exception is what the Regulations refer to as a        Wondra, K.J., J.D. Hancock, K.C. Behnke, C.H.
customer formula feed. If the pork producer               Fahrenholz, C.R. Stark, and R.H. Hines. 1992a.
specifically asks for a feed that may contain levels      Effects of mill type (hammer vs roller) and particle
above or below the standards outlined in Table 4          size uniformity on growth performance, nutrient
and provides a signed request form listing all the        digestibility, and stomach morphology in finishing
ingredients to be used, the feed manufacturer does        pigs. Swine Day Progress Report pp.126-129.
not have to register the feed with the Federal
government. Consequently, requests from farmers           Wondra, K.J., J.D. Hancock, K.C. Behnke, G.A.
to commercial feed manufacturers should keep such         Kennedy, and R.H. Hines. l 992b. Does diet form
restrictions in mind.                                     (pelleted vs meal) affect optimum particle size of
                                                          com for finishing pigs? Swine Day Progress Report
  Agriculture Canada provides the 'Medicated Feed         pp. 122-125.
Information Guide'. Like any legal document, the
Feeds Act is written in very precise legal language       Wondra, K.J., J.D. Hancock, K.C. Behnke, R.H.
which is sometimes difficult to understand. This          Hines, and C.R. Stark. 1993. Effects of
Guide is useful because it explains the Feeds Act         hammerrnills and roller mills on growth
and associated documents in simpler terms.                performance, nutrient digestibility, and stomach
Everyone mixing feeds on the farm would be well           morphology in finishing pigs. Swine Day Progress
advised to read it, so they are familiar with their       Report pp.135-138.
legal obligations. The Guide also provides useful
tips on feed mixing and handling to help avoid
potential problems.
                                                        229
230
11. TOXIC SUBSTANCES IN FEEDS
  While the main focus of animal nutrition is the
need for - and supply of - nutrients, the feedstuffs                Potential Toxins or Toxin Producers
which supply these nutrients may also carry with                              Which May be
them certain toxins or antinutritional factors. Tox-              Present in Common Feed Ingredients
ins are generally considered to be poisonous sub-
stances present in the feed, often produced by the                Endogenous                 Exogenous
host plant or by an organism living in the feed,
which could adversely affect the health and wellbe-               Phytates                   Fungi
ing of animals; conversely, antinutritional factors               Saponins                   Aspergillus
are substances which impair normal feed digestion                 Estrogens                  Fusarium
or nutrient utilization. A diet can be perfectly                  Trypsin Inhibitors         Penicillium
balanced but fail to support optimal growth if it                 Tannins                    Ergot
contains anti-nutritional factors that adversely affect           Glucosinolates
appetite, feed utilization or animal health.                      Lectins                    Bacteria
                                                                  Thiaminase                 Escheriscia coli
   Toxins may be substances that occur naturally in               Oxalates                   Salmonella
certain species of plants. Examples are tannins and               Gossypol                   Campy lobacter
phytates; their presence is readily predicted and                                            Listeria
appropriate adjustments can be made in the diet                                              Clostridia
formulation. Toxins can also be the result of
contamination by bacteria or fungi. The bacteria
themselves may be toxic to the pig, or if the bacte-
ria or fungi are not poisonous. they may produce            to formulate rations around them. Research is
substances which are. This is not a predictable             underway to negate the effects of many of these
occurrence and therefore. can be much more diffi-           toxic factors. The following is a discussion of some
cult to deal with. Listed adjacent are some of the          of the more common toxicants which may be
toxins or toxin producers which may be found in             encountered when formulating pig diets in Canada.
feeds in Canada. It does not include weed seeds
which may also contain noxious substances.                  Phytates
Mycotoxins are metabolites (products of metabo-             Phytic acid is present in many plant seeds, contain-
lism) of fungi, the most common in Canada being             ing 6 phosphorus molecules linked to an organic
ergot alkaloids, ochratoxin A, zearalenone and the          compound called inositol. In the plant kingdom, it
tricothecenes. These are discussed in detail later in       serves as an important storage form of phosphorus.
the chapter.                                                Phytic acid may chelate or bind with a variety of
                                                            minerals, including calcium, magnesium, iron and
                                                            zinc to form phytate. In pig diets, phytate typically
Toxicants Produced by Plants                                binds calcium and magnesium as well as phospho-
  Many common feed ingredients contain natural              rus; destroying phytate through enzymatic or
toxicants and/or toxins which may impair pig                processing activity will therefore increase the
performance. For example, raw soybeans contain              utilization of all three mineral elements, not just
trypsin inhibitors and the enzyme urease. They are          phosphorus as normally assumed.
part of the normal plant or seed and are generally
predictable in both quantity and impact on the pig.           In mature cereal grains, 60% to 80% of the total
Although their presence is undesirable, our knowl-          phosphorus is so bound; in soybean meal, about
edge of anti-nutritional factors allows nutritionists       half of the phosphorus exists as phytate phosphorus.
                                                          231
Table 11-1 provides information on the relative               Table 11-1. Phytate Phosphorus Content of Common
proportion of total phosphorus present in the form            Feedstuffs.
of phytate in common feedstuffs.
                                                                                   Total              Phytate
  Phytate is of nutritional significance because              Feeds tu ff      Phosphorus, %        Phosphorus,
phosphorus in phytic acid is poorly absorbed by the                                                  % of total
pig. In fact, the availability of total phosphorus            Alfalfa meal          0.30                   0
from grain sources ranges from 20% in com to 45%              Barley                0.34                  56
in wheat; barley and oats are intermediate at about           Com                   0.26                  66
30% availability. These low availabilities are the            Cottonseed meal       1.07                  70
result of poor utilization of phytate-bound phospho-          Grain sorghum         0.31                  68
rus - essentially zero bioavailability in com and             Oats                  0.34                  56
oats, 10% in barley and 40% in wheat. In contrast,            Sesame meal           l.27                  81
inorganic phosphorus supplements, such as                     Soybean meal          0.61                  61
dicalcium phosphate are about 80% available or                Wheat                 0.30                  67
better.                                                       Wheat bran            1.37                  70
                                                              Wheat middlings       0.47                  74
   A number of factors will influence the availability
of phytate phosphorus. Germination releases the               Source: Ward, 1994.
naturally-occurring phytase in cereal grains; within
one week of germination, the amount of phytate
phosphate will be reduced by 50%. Some grains                 Table 11-2. Naturally-occurring Phytase in Common
contain more natural phytase than others; wheat in            Feedstuffs.
particular, is known to be a relatively rich source of
                                                               Feedstuff               Relative Phytase Activity
phytase and thus has a higher phosphorus availabil-
ity (Table 11-2).                                              Wheat, rye                       High
                                                               Barley                         Medium
   Phytase supplements can also be purchased for               Oats, soybean meal               Low
inclusion in pig diets; these are normally derived             Com, sorghum                   Very low
from a genetically-modified Aspergillus strain.                Cottonseed                    Undetectable
Biological action will vary with the source, but as a
rule, approximately 500 units of pbytase per kg of            Source: BASF Technical Bulletin No. 9201.
mixed feed is equivalent to 0.1 % supplemental
phosphorus in the diet. Phytase supplements must              Saponins
be properly handled to obtain maximum benefit.              Saponins are a family of compounds found in
Because phytase is an enzyme, it is susceptible to        many plants of economic importance, including
the effects of heat and moisture. Therefore,              canola. alfalfa, soybeans, peas, sugarbeets, sunflow-
pelleting diets containing phytase must be carefully      ers, oats, chickpeas and beans. They are character-
controlled to avoid die temperatures in excess of         ized by a bitter taste, but are used commercially in
70°C as losses in enzyme activity may exceed 25%;         such diverse consumer products as shampoo, soft
alternatively, phytase can be added as a liquid post-     drinks, soap and fire extinguishers. The type of
pelleting or used in mash diets.                          saponin, the concentration present and the physi-
                                                          ological effect on swine varies a great deal within
  Certain micronutrients, such as iron sulphate,          and among plant species.
copper sulphate or choline chloride will contribute
to phytase breakdown. Therefore, phytase is best            If injected, saponins are highly toxic, attacking
not added to premixes containing trace minerals or        and altering the structure and function of cell
choline; because these ingredients are diluted in         membranes. However, when administered orally,
complete feed, their impact on phytase in mixed           their effect is greatly reduced. Saponins impair pig
diets is greatly reduced.                                 performance, due to their bitter taste and irritating
                                                        232
effect on the lining of the mouth and gut. There is        formance independent of their impact on protein
some suggestion that the absorption of certain             digestion, at least in some species. In some species,
nutrients, including vitamins and trace minerals is        but not the pig, the effect of trypsin inhibitors can
altered by saponins, presumably due to changes in          be reduced by supplementation with sulphur amino
the structure of the lining of the gut; for example,       acids; this occurs because such animals respond to
soybean saponins bind zinc and render it less              trypsin inhibitors by producing more trypsin, a
available, while ginseng saponins actually increase        protein which is particarly rich in sulphur amino
the absorption of iron. Interestingly, saponins also       acids. In the pig, inhibition of protein utilization
lower blood cholesterol levels.                            occurs because the quantity of trypsin inhibitor
                                                           present exceeds the available supply of trypsin, and
  In animal nutrition, concern about impaired              increased supply is either not possible or is inad-
performance due to saponins is most often associ-          equate.
ated with alfalfa. The saponin content of alfalfa
varies according to season, being highest in mid-            Trypsin inhibitors are readily destroyed by heat;
summer and tapering off thereafter; not surprisingly,      fifteen minutes at 100°C is sufficient to remove
the saponin content of second cut alfalfa tends to be      almost all negative effects. A simple, indirect test,
higher than that of first or third cuttings. Plant         called the urease test, is often employed to deter-
breeders are now developing low saponin alfalfa            mine the level of trypsin inhibitor left after
varieties.                                                 soybeans have been processed. The enzyme urease
                                                           is also present in soybeans; it converts urea to
Estrogens (Phytoestrogens)                                 ammonia and can be tested quite easily. It is as-
  The estrogen content of feeds can occur from the         sumed that the destruction of urease during soybean
plant directly or from contamination with molds            processing parallels that of trypsin inhibitor, so that
which produce estrogenic mycotoxins. Alfalfa and           if one is inactivated, the other will be similarly
soybeans can produce an estrogenic compound                destroyed. If the urease test is negative, this serves
called coumestrol. The use of alfalfa pellets in sow       as an indirect indication that trypsin inhibitor has
diets has declined, due in part to the fear that           also been destroyed.
possible estrogenic compounds may impair repro-
ductive performance.                                         Heating ingredients to destroy protease inhibitors
                                                           must be carried out with great care; this is because
Protease Inhibitors (Trypsin Inhibitors)                   excess heat may damage certain amino acids,
  Most legume seeds (eg. soybeans, faba beans),            especially lysine, rendering them unavailable to the
some grains (eg. com, rye, barley and triticale),          pig.
alfalfa and potatoes contain protease inhibitors.
These are protein molecules which bind to and              Tannins
inactivate trypsin or other digestive enzymes that           The term "tannin" was originally used to describe
help to break down protein molecules in the small          a group of plant extracts used in tanning leather.
intestine. Of the protease inhibitors, trypsin inhibi-     They are found in many crops, including rapeseed,
tors are best known, but inhibitors of chymotrypsin        soybeans, fababeans, sunflower seeds, alfalfa and
also exist in nature. In practical terms, while            sorghum. More technically speaking, tannins are a
protease inhibitors are widespread throughout the          group of compounds that bind to proteins, including
plant kingdom, the ones of greatest significance in        dietary proteins, digestive enzymes and proteins
pig nutrition are those found in soybeans, other           which assist in nutrient absorption in the gut. It is
beans and triticale.                                       not surprising, then, that tannins impair the ability
                                                           of the pig to use dietary protein, carbohydrates and
  ln swine nutrition, protease inhibitors are most         fats; however, the most significant impact is directly
often discussed in the context of impaired perform-        on protein utilization. Tannins also reduce palat-
ance due to reduced protein digestibility. However,        ability, due to a sour taste.
trypsin inhibitors have been shown to impair per-
                                                         233
  Tannins are divided into two broad categories:               Lectins are known to impair growth, cause diarrhea
condensed and hydrolyzable. The latter are named               and interfere with nutrient utilization.
due to their ability to be readily broken down while
the condensed tannins are much more stable and                Found in highest concentration in legumes,
complex in structure.                                      including soybeans, lectins were first associated
                                                           with castor beans. The ability of the lectins to bind
  The tannin content of sorghum differs among              to cells along the intestinal tract is highly specific,
varieties, with yellow sorghum having very low             such that certain Jectins will bind and others will
levels. Tannin content is directly related to bird         not. If binding occurs, the lectins may enter the cell
resistance in the field; consequently, plant breeders      and. if toxic, can initiate a range of adverse reac-
are faced with the decision to select varieties for        tions, from greatly enlarged intestinal tissue to
low tannin content, desirable in animal nutrition, or      impaired nutrient absorption and depleted body
high resistance to predation by birds, which is            muscle, fat and glycogen reserves.
desirable in regions where sorghum is a human food
staple, but is vulnerable to attack by wild birds.         Thiaminase
Sorghum tannins are of the condensed kind.                   Thiaminase is an enzyme which destroys the B
                                                           vitamin thiamine. It is not found in any feedstuff
  The most effective means of addressing the issue         common in pig diets, but is present in certain fish,
of tannins in swine nutrition is to select crops and       including carp. If such fish are fed in uncooked
varieties of crops with little or no tannin content.       form to swine, problems may occur. Since feeding
Alternatively, affected crops can be treated with          raw fish to swine is unlikely, the risk of problems
dilute alkali or polyethylene glycol to improve their      due to thiaminase is remote.
feeding value. Heating is somewhat less effective
because tannins are heat stable.                           Oxalates
                                                             Oxalic acid is a compound with the ability to bind
Glucosinolates                                             calcium, rendering it less available for absorption
  Glucosinolates are common in plants in the               by the pig. Oxalate poisoning, while common in
Brassica family, including rapeseed, mustard,              grazing species exposed to certain plants, is rare in
turnips, crambe and kale. They reduce palatability         swine. However. some feedstuffs that may be
and impair the function of the thyroid gland. Since        included in swine diets, such as alfalfa. contain
the thyroid gland is so important in regulating            oxalate, lowering the bioavailability of calcium.
animal growth, this can be a serious problem.
                                                           Gossypol
  Canadian plant breeders have developed low                 Gossypol is a yellow pigment found throughout
glucosinolate varieties of rapeseed; to distinguish        the cotton plant: in the seed. gossypol is found in
these new varieties with highly different nutrition        the pigment glands. During processing. the glands
characteristics, the term "canola" was coined.             are ruptured and gossypol released. A portion of
Continued research on lowering the glucosinolate           the gossypol will bind to protein, primarily the
level to essentially zero is on-going; however, it is      amino acid lysine, rendering it less available to the
not yet known if these further improvements will           pig. Thus. the higher the portion of bound as
affect animal performance. Meal from current               compared to free gossypol, the lower will be the
varieties of canola can be fed at high levels to swine     biological value of the protein. The remaining, or
without any apparent effect due to glucosinolatc           free. gossypol is reactive and affects animals in a
activity.                                                  number of ways: depressed appetite. lower weight
                                                           gains, impaired lung, heart and liver function,
Lectins                                                    anemia and male infertility. Free gossypol levels
  Lectins arc either protein or carbohydrate-protein       should not exceed I 00 ppm in pig diets.
compounds which share a common characteristic:
the ability to selectively bind certain carbohydrates.
                                                         234
  The effects of gossypol can be reduced by adding        feed trucks that arc regularly sanitized as a further
ferrous sulphate to the diet; the generally accepted      safeguard against disease transmission. The data in
recommendation is 1 unit of iron as iron sulphate         Table 11-3, based on the results of a survey reported
per unit of gossypol. Even with added iron, the           by the Veterinary Diagnostic Laboratory at the
maximum upper limit of free gossypol in swine             Pennsylvania State University, illustrates that
diets is 400 ppm. Increasing dietary protein is           removing ingredients of animal origin will not
helpful as well, but rarely economical.                   eliminate the risk of Salmonella contamination.
Exogenous Microbes and Toxin                              Table 11-3. The Results of Salmonella Assays
                                                          Reported by the Veterinary Diagnostic Laboratory at
Producers                                                 the Pennsylvania State University.
Bacteria
  Contamination of feed by pathogenic microbes                Ingredient          No. Samples        % Positive
such as certain strains of Salmonella may lead to
                                                              Animal Protein           62                63
infection of the herd and an outbreak of disease. In
                                                              Distillers Grains        51                 0
the case of Salmonella, there are more than l ,000
                                                              Canola Meal               11               18
different serotypes, yet only a handful tend to be of
                                                              Oats                     15                 7
significance in pork production. including S.
                                                              Soybean Meal             20                20
choleraesuis and S. typhisuis. Some other Salmo-
nella serotypes have a broad spectrum of hosts and        Adapted from John, R.E., 1990. Proc. Symp. Feed
could therefore infect swine if the conditions were       Quality Assurance, Centre for Veterinary Medicine,
right.                                                    USDA, Arlington, VA.
                                                        235
Fungi and Mycotoxins                                       aflatoxin, and Fusarium roseum which produces
  Fungi impair feed quality in at least two ways.          zearalenone and vomitoxin. Several other fusarium
They may alter the nutritional quality of feeds by         varieties also produce toxins while Penicil/ium and
destroying or making unavailable important nutri-          Aspergillus produce ochratoxin. These are ex-
ents such as vitamins or they may produce                  plained in detail later in the chapter.
myeotoxins that are harmful to the pig.
                                                               Ergot
  The direct effect on nutrient composition is               Ergot alkaloids are very potent toxins produced
difficult to predict and attempts have failed to           by the fungi ergot (Claviceps purpureay. Triticale
demonstrate impaired pig performance due solely to         and rye are the most susceptible grains, but wheat,
fungal growth. Documented cases of feeding                 barley, oats and corn can also become infected.
extremely moldy corn to swine without difficulty           Visual inspection of suspect grain is useful, since
demonstrate that mold by itself probably represents        the infected kernels are usually dark, swollen and
only minimal risk to the pig. It is not until the fungi    have a rough coat. Once the grain is ground,
produce mycotoxins that animal performance                 detection requires laboratory analysis.
suffers. Some toxins need only be present in very
small concentrations to affect pig health.                    Ergot alkaloids can severely affect pig health and
                                                           performance. They impair blood flow to the
  While it would be useful to have tables summariz-        extremities, affecting tissues such as the kidney and
ing maximum tolerances for the various toxins of           uterus. Reduced blood flow to the reproductive
importance in swine production, the reality is that        tissues can result in spontaneous abortion in poi-
such tables are very difficult to produce. The             soned sows. In fact, in ancient times, abortions in
maximum tolerance for any particular toxin will be         humans were induced by feeding ergot. Piglets
affected by many factors, including the response           born to infected sows have poor viability, due to
criteria (animal performance versus tissue damage),        impaired lactation of the dam.
time frame (short-term versus long-term effects)
and, interestingly, the nutrient specifications of the       Poor blood flow also explains other symptoms of
diet. For example, some toxins are involved in the         ergot toxicity such as blackening of the tips of ears
metabolism of highly toxic compounds called free           and sloughing of hooves. In these examples, poor
radicals; protection from free radicals is obtained by     blood flow actually results in the death of the
consuming increased quantities of antioxidants,            respective tissue. Ergot poisoning can also reduce
such as vitamin E. Consequently, the toxic level of        feed intake, induce convulsions and muscle incoor-
certain mycotoxins will vary, depending on the             dination, cause respiratory distress and precipitate
vitamin E level of the diet.                               an outbreak of diarrhea.
  Another problem in determining the toxicity of a            At the present time, only imprecise recommenda-
moldy grain sample is the occurance of multiple            tions on ergot are available. Suspect grains should
toxins, some of which may be present at low levels         never be fed to breeding animals and the quantity of
or which are difficult to identify. While the toxin(s)     contaminated kernels fed to growing pigs should
assayed may be below tolerance, associated toxins          not exceed 0.1 % of the total grain in the ration ( one
which may not have been measured may contribute            infected kernel for every 1000 clean kernels).
to the overall toxicity of the sample. This is a
particularly troubling aspect of mycotoxin technol-        Ochratoxin A
ogy, since there are literally dozens of toxins known        Ochratoxin affects the kidneys in swine. Post
to man and one cannot practically assay them all!          mortems of infected pigs reveal swollen kidneys
                                                           which are lighter in colour and firmer than healthy
 The molds of greatest interest in Canada and the          kidneys. Ochratoxin can cause birth defects when
United States areAspergillusjlavus which produces          sows are infected and impairs growth in market
                                                         236
hogs. Swollen kidneys will occur at levels much          Photo ll-1 a.
lower than those required to impair performance.
lmmunosuppression has also been associated with
Ochratoxin A.
Zearalenone
  Zearalenone, sometimes referred to as F2 toxin, is     Aborted and mummified piglets.
a mycotoxin produced by the fungi Fusarium. It is
often associated with corn but can infect many other
crops including wheat. It is an estrogenic com-          Photo 11-lb.
pound; as a result, its effects on the pig are similar
to those of the natural hormone, estrogen. Estrogen
particularly affects reproduction and consequently,
most research on zearalenone bas been carried out
on sows and gilts, with less information being
available on growing pigs. Pigs, as a species, are
particularly susceptible to zearalenone toxicity.
Levels between 1 and 5 ppm are sufficient to elicit a
number of estrogenic symptoms.
              SYMPTOMS OF
          ZEARALENONE TOXICITY
                                                       237
  Zearalenone can also be carried in the sow's or          Vornitoxin contaminated feed does not appear to
gilt's milk and thus affect the growth rate of new-      affect reproductive performance in gilts. Levels up
born piglets. Up to 30 ppm of pure zearalenone           to 8 ppm vomitoxin did not produce adverse results
appears to have a minimal effect on litter size in       with regards to reproduction. Since vomitoxin is
gilts while levels of 60 to 90 ppm result in the         often found in grains containing zearalenone, it is
complete loss of fetuses. The impact of                  difficult to determine which of these toxins is
zearalenone remains with gilts for many months           causing a given problem. From research to date, it
after the infected diet is removed.                      appears reproductive impairment is most likely due
                                                         to zearalenone and not to vomitoxin contamination.
   Zearalenone appears to also affect boars. lt
reduces libido (sex drive) and delays sexual matu-         Other trichothecenes may be even more toxic than
rity in growing boars. However, diets containing up      vomitoxin. They cause feed refusal, vomiting,
to 60 ppm zearalenone have no apparent effect on         irritation of the skin and the gut and may also result
semen quality or libido of mature boars.                 in the birth of deformed piglets. One to JO ppm
                                                         diacetoxyscirpenol will reduce growth rates. Diets
  The effect of zearalenone on growth is less clear.     containing 8 ppm T-2 toxin do not affect weight
Some experiments have demonstrated that as little        gains but increasing levels to 16 ppm will depress
as l O ppm of zearalenone depresses pig growth and       growth rate.
appetite. Other experiments suggest that there is no
effect on growth at levels of toxin as high as 50        Aflatoxin
ppm. There is even the suggestion that growth is            Aflatoxin is produced primarily by the fungus,
actually enhanced by zearalenone. Part of the            Aspergillus. Ideal conditions for Aspergillus growth
problem with the variable responses produced by          include moisture content above 14%, temperature
zearalenone may be due to contamination of the test      above 25°C and the presence of oxygen. It is
ingredients with other toxins such as vomitoxin.         known to affect many feed ingredients including
Vomitoxin is known to impair performance and is          corn, rye. oats, wheat, barley, soybean meal, sun-
often found in samples of grain contaminated with        flower meal, rapeseed and alfalfa. There are four
zearalenone.                                             types of aflatoxin: B1, B2, G1 and G�. Aflatoxin B1
                                                         is considered to be the most potent for swine.
Tricothecenes
  Tricothecenes are a group of very toxic com-             The impact of aflatoxin on swine is age and dose
pounds which include deoxynovalenol (also called         dependent, and ranges from depressed perfonnance
DON or vornitoxin), HT-2 toxin,                          through immune suppression. Acceptable upper
diacetoxyscirpenol and T-2 toxin. They are pro-          limits have been defined by various agencies; for
duced by Fusarium molds and are very toxic to            young pigs, levels in excess of 20 ppb should be
pigs. T-2 toxin is unique in that it forms during        avoided, for finishing swine, the limit is 200 ppb
storage at low temperatures (6 - l 8°C).                 and for breeding animals, the limit is l 00 ppb.
                                                         Recovery from aflatoxin toxicity is rapid in grow-
  One feature of tricothecenes that is particularly      ing pigs. Seven days on clean feed will usually
troubling is its suppression of the immune system.       result in a full recovery. In younger pigs, scouring
Even low levels in the diet may increase the pig's       can result from aflatoxin poisoning. In sows, milk
susceptibility to pathogenic organisms that normally     production is impaired.
would not create health problems. Studies with
contaminated wheat indicate that as little as l ppm
vomitoxin reduces feed intake and growth rate in
market hogs. Levels in excess of 20-40 ppm cause
vomiting and feed refusal. One of the most striking
features of DON contamination is feed refusal.
                                                       238
Prevention of Mycotoxin                                   prevent mold from growing. At 21 °C, moisture can
                                                          rise to 15% before there will be a problem. In
Contamination
                                                          soybean meal, moisture should be kept below 14%.
  Since effective treatment methods have not been
developed, prevention of mold contamination is
                                                            In summary, it is important to recognize that mold
very important. Cropping practices can play a
                                                          contamination of grains can occur at many stages:
useful role in prevention. Rye and triticale are
                                                          during growth, at the time of harvest, during storage
examples of crops that are particularly susceptible
                                                          as seed or after being processed into livestock feed.
to ergot. The selection of more resistant grain
                                                          Field contamination is worse when kernels are
varieties is a major consideration. Late maturing
                                                          damaged, when the weather conditions are moist or
com may provide greater yields but the seeds are at
                                                          if insects infest the crop. The satisfactory storage of
a susceptible stage of development during the wet
                                                          grains or mixed feeds requires low moisture levels
weather that often occurs in the late summer and
                                                          and temperatures as cool as possible.
fall. Crop rotation can help break the life cycle of
infecting spores. Com and wheat can be rotated
with less susceptible crops. Profitability in the short   Treatment
term may be reduced. but this must be compared to           If grain contamination is suspected, samples
the long-term benefits. Researchers have also             should be submitted to a laboratory for analysis.
observed that scabbiness caused by Fusarium               Increasing numbers of labs are setting up to conduct
molds, especially in wheat, can be reduced by             mycotoxin analyses. It is a difficult analysis due to
fertilizer selection. The use of urea fertilizer in       the sophisticated tests required and also due to
place of ammonium nitrate can be helpful.                 problems with sampling. A heavy infestation of
  Fusarium infestation is favoured by warm (15°C-         only a small portion of a field of com can result in
350C), windy, rainy weather. These conditions             swine diets contaminated above acceptable limits.
provide an excellent environment for infecting            If sampling failed to include the contaminated area
spores to spread. Therefore. this kind of weather         of the field, tests would be falsely negative.
during late growth and harvest should raise con-
cerns about potential contamination.                        If grains are suspected of being infected, a sample
                                                          can be fed to a group of young gilts. If swelling or
  Once grains are harvested, other practices can be       reddening of the vulva appears or iffeed intake is
beneficial. The cleaning of grains to remove              impaired, contamination should be suspected. The
damaged kernels will help to remove sources of            infected grain should not be fed to breeding animals
infestation. Broken kernels and related material
often contain the highest concentration of
mycotoxins. Their removal greatly improves the
feeding value of the grain. The addition of mold
inhibitors will not kill mycotoxins already present
but will prevent further formation.
                                                        239
and if it is fed to market hogs, it should be diluted         Additional Reading and References
as much as possible with "clean" grain.                   Anon. 1987. Recent developments in the study of
                                                          mycotoxins. Kaiser Chemical Co., Cleveland, OH.
  Chemical treatments of infected grains are being
investigated. It has been shown that sodium               Cheeke, P.R. and LR. Shull. 1985. Natural
bisulphite added to infected corn and then                toxicants in feeds and poisonous plants. AYINon
autoclaved can reduce DON (deoxynovalenol or              Nostrand Reinhold Co., New York. 492 pp.
vomitoxin) concentrations by a factor of I 0. To
date, this is not commercially practical, because         d'Mello, J.P.F., C.M. Duffus and J.H. Duffus.
autoclaving is slow and expensive. However, other         1991. Toxic substances in crop plants. The Royal
heating processes may prove to be as beneficial.          Society of Chemistry, Cambridge. 339 pp.
  Another possibility is the addition of hydrated         Marquardt, R.R. 1993. Effects of moulds and their
sodium calcium aluminosilicate (HSCAS) to diets           toxins on livestock performance: A western Cana-
containing vornitoxin and/or zearalenone. Prelimi-        dian perspective. Proc. Western Nutrition Conf.,
nary results suggest that there may be some value in      Winnipeg, MB. pp. 115 - 121.
the use of such products, although more research is
required.
                                                          Trenholm, H.L., D.B. Prelusky, J.C. Young and J.D.
                                                          Miller. 1988. Reducing mycotoxins in animal
  If mild contamination is suspected, the nutrient        feeds. Publication #1827E, Agriculture Canada,
density of the diet can be increased to help compen-      Ottawa, Ontario.
sate for reduced intake. The vitamin, mineral and
amino acid concentrations should be increased by
5-20% depending on the severity of feed refusal
expected.
Safety
  Molds and mycotoxins can also be harmful to
humans. Symptoms include allergies, skin and eye
irritation, headaches and vomiting. Fusarium
spores are a well-known toxin and must be avoided.
Aspergillus poisoning has also been recorded. If the
grain must be handled, ventilation is important and
the use of gloves and respirators is advised.
                                                        240
12. WATER
  Like other nutrients, such as energy and essential
amino acids, water is an absolute requirement in the
diet of the pig. Lndeed, the pig can survive much
longer without energy or protein, minerals or
vitamins than it can without water. This is espe-
cially true in hot weather.
                                                       241
the drinker line overlooks two important problems:           underestimated; boredom and hunger can increase
the amount of water wasted and the inaccuracy of             water intake many fold over basic requirements.
most meters at very low or intermittent flow rates.
Waste can represent a significant portion of total             The obvious question relates to the importance of
water "disappearance." With nipple drinkers, waste           over-consumption. ls it a problem? For the most
can reach 50% or more of intake and will be even             part, we do not believe that it is, in terms of pig
higher when boredom or other stressors are present.          health or productivity; any water consumed above
Much of the earlier information published on water           requirement will be eliminated as urine. However,
intake failed to measure waste and must be inter-            over-consumption does waste water and perhaps
preted with great care.                                      most critically, increases the cost of slurry storage
                                                             and spreading. Excess water intake, and more
  Research at the Prairie Swine Centre and else-             importantly, direct water waste is a growing con-
where has found that true, free choice water intake          cern; the cost of slurry storage is rising in response
in young growing pigs fed ad libitum is about 2.2 to         to environmental concerns and the expense of
2.8 times the intake of feed. Thus, a pig eating 2 kg        spreading slurry is also increasing due to higher
of feed will normally drink at least 4.5 litres of           labour and fuel costs.
water per day.
                                                               Is under-consumption a problem? There is an
  Additional water, to compensate for elevated               increasing body of data which strongly suggests that
environmental temperature or excess mineral or               even under ad libitum conditions, pigs do not
protein in the diet, or to help the pig deal with            always drink as much water as they require for good
certain health problems. must be added to the above          health and maximum performance. Clearly, more
intake levels. It is difficult to define water intake at     research is needed, but the implication is that
high barn temperatures, since much of the reported           providing water free choice may not always ensure
research measured water disappearance, which                 optimum intake.
includes waste, rather than actual intake. Actual
intake probably rises by only 15% to 75% in hot              Gestating Sows
weather, but waste increases dramatically, such that            Water use by gestating sows is affected by both
total water disappearance from nipple drinkers               physiological need and behavioural influences.
might increase as much as 3- to 4-fold. Following            Since dry sows are limit fed, they consume addi-
are some of the factors that increase or decrease            tional water to help achieve a feeling of satiety (full
water consumption.                                           stomach). This additional intake has little to do
                                                             with "requirements" as defined in the classical
                                                             sense, but may be important to the sow's achieving
   INCREASE                     DECREASE                     an overall sense of well-being. For example, in
                                                             human nutrition, it is not uncommon to recommend
   Hunger                       Cold stress                  that people on diets consume additional water, in
   Boredom                      Warm water                   part to help achieve a sense of satiety.
   Heal stress                   temperature
   Increased dietary minerals   High mineral levels
                                                               In addition, dry sows are often housed in indi-
   Moderate mineral levels        in water
    in water                                                 vidual gestation stalls, where boredom is common.
   Pelleted feed                                             Sows will consume additional water, and play with
                                                             drinkers more often, merely to offset this sense of
                                                             boredom. This additional water intake and playing
                                                             with drinkers may lead to excessively wet, dirty
  Pigs do not drink solely to satisfy their                  conditions in the barn, depending on the design of
physiological need for water. They will also drink           the stalls. If it is a problem, providing water in the
water to alleviate a feeling of hunger or boredom.           feeding trough, rather than nipple drinkers, has
The impact of "luxury" intake must not be
                                                           242
proven helpful. The provision of small amounts of         this is not practical in most situations, placing water
straw, to serve as an object of foraging, or the          in the feeder for the first three to four days after
feeding of a bulky diet have also proven helpful in       farrowing, for those few sows who do not appear to
minimizing excessive drinking.                            be drinking or eating well, may be advised.
  Providing water in a properly-designed trough             Beyond day three, when this initial adjustment
which minimizes waste and spill is an ideal method        period of early lactation has passed, nipple drinkers
for providing water to the dry sow. In the case of        with flow rates between I and 2 L/minute, will
nipple drinkers, firm recommendations are not yet         suffice. While some recommendations exist for
available, but flow rates of 0.5 L/minute will more       much higher flow rates, research at the Centre for
than suffice.                                             Food and Animal Research in Ottawa suggest no
                                                          such advantage exists; the only real effect is greatly
Lactating Sows                                            increased wastage with attending problems and
   All things being equal, lactating sows have the        costs. Indeed, the few studies which have shown
highest relative requirement for water, due to the        impaired lactation performance required flow rates
demands of milk synthesis. Intake studies under ad        well below 0.5 L/minute!
libitum conditions have suggested that nursing sows
will drink between 9 and 20 litres of water per day.        Some suggestions have been made that sows will
This is a very large variation and reflects the tre-      spend no more than five to six minutes drinking
mendous differences one sees among animals.               water per day; this may be true for a few sows
Well-controlled experiments, where wastage is             during the first two to three days after farrowing,
accurately measured, reveal that the majority of          but is clearly not the case for most sows and cer-
sows will drink about 15 litres per day.                  tainly not beyond the third day of lactation. Re-
                                                          search has shown that sows will spend 15 minutes
   However, provision of water ad libitum may not         or more per day at the drinker.
be enough to ensure optimum intake. Drs. Fraser
and Phillips at the former Centre for Food and              Excessive flow rates will not only waste water,
Animal Research in Ottawa have studied water              but will also result in messy crates and wet piglets.
intake in newly-farrowed sows and found that litters      For this reason, addressing the issue of lethargic
tended to gain poorly during the first three days         sows by adding water to the feeder for those few
after birth if the sow drank less than 10 litres of       animals in difficulty is a more practical and eco-
water during this period. The researchers concluded       nomical solution.
that in some sows, poor early lactation performance
is associated with very low water intake. This low        Piglets
water consumption may be a symptom of other                 Recent research, again by Ors. Fraser and Phillips,
health problems, or just a case of simple lethargy on     suggest that piglets may require water immediately
the part of the sow. Until more information be-           after birth. They reported that litters gaining poorly
comes available, it may be prudent to encourage           during the first one to three days after farrowing
sluggish sows to become active as soon after              drink more water than faster growing litters. It has
farrowing as possible to stimulate drinking and thus      been suggested that when sows are not milking
promote better early lactation performance. Placing       well, piglets may not consume enough water from
nipple drinkers at different heights in the farrowing     their limited milk intake and might benefit from
crate docs not appear to alleviate the problem.           supplemental drinking water. Nipple drinkers are
                                                          not adequate for newborn piglets, because the
  How should water be provided to the lactating           piglets take too long to find and use the drinker; a
sow? In early lactation, lethargy is a problem in         dish drinker, located in the creep area, is a better
some animals and the best way to maximize the rate        alternative. Balancing cost and convenience,
of consumption is to provide water troughs. While         portable dish drinkers may be the best choice,
                                                        243
employing them only in newborn litters where sow                Water Delivery
milk production appears to be a problem.                          Nipple drinkers are by far the most common
                                                                method of providing water to pigs. They are of
   Some people have asked if the provision of                   relatively low cost, easy to maintain and generally
drinking water to very young piglets may be harm-               free of fouling. However, nipple drinkers do have
ful, reducing their motivation to suckle. A number              their problems. They encourage waste and cannot
of studies have clearly indicated this is not the case.         be easily monitored; often, it will be many hours or
Consequently, providing water to newborn piglets,               even days before a plugged drinker is noticed, if
although not a common practice, is strongly recom-              daily checks are not observed. Nonetheless, nipple
mended, especially in warm conditions where                     drinkers are probably the method of choice for the
piglets lose moisture rapidly.                                  time being for all classes of swine except newborn
                                                                piglets, where dish drinkers are clearly superior.
  One common belief is that providing water to
nursing piglets encourages creep feed intake.                     A relatively recent variation is the wet/dry feeder
Although this may be true, studies have shown that              which incorporates a nipple drinker inside the
the effect is relatively small until the pigs are four to       feeder bowl; in some respects, they combine the
five weeks of age.                                              features of nipple and dish drinkers. Since in some
                                                                circumstances they increase feed intake, wet/dry
Weanling Pigs                                                   feeders are increasing in popularity. They also
   At the time of weaning, pigs are suddenly re-                reduce water wastage; a Manitoba study by Froese
quired to obtain aU of their water from the drinker.            and Hodgkinson found that water usage was re-
It is not altogether clear how well the newly-weaned            duced by 40% with the use of wet-dry feeders.
pig can regulate water metabolism, or select a daily
intake appropriate to its needs. For example, a               Several studies have demonstrated that restricted
number of experiments have shown that water                 nipple-drinker flow rates can impair water intake
intake follows a peculiar pattern at weaning, being         and, under severe restriction, growth rates as well.
initially quite high, falling to a minimum at about         However, the flow rates must be quite low - prob-
four days post-weaning, followed by a gradual               ably below 200 mL/min. A recent survey of Sas-
increase as feed intake rises. Surprisingly, water          katchewan farms revealed typical flow rates that
intake is at its lowest when scouring is typically at       greatly exceed this minimum (Table 12-1 ). These
its worst, raising questions as to the adequacy of          data suggest that excessive flow rates, leading to
water intake at this time.                                  wastage, may be more of a concern than inadequate
                                                            flow rates. However, individual nipple drinkers still
                                                            need to be checked regularly to ensure against
                                                            blockage or malfunction.
                                                          244
   The critical question for pork producers is what          such as the age or health status affect the response
are the minimum and maximum flow rates neces-                of individual animals.
sary to optimize health and productivity? Unfortu-
nately, research on the subject is extremely limited;
in the absence of definitive experimental data,              Table 12-3. Canadian Water Quality Guidelines for
estimates are the best that can be provided. For             Livestock.
weanlings and growers, flow rates between 500 and
 1500 mL/min would appear to be satisfactory. For            Item                     Maximum Recommended
                                                                                          Limit, ppm
nursing sows, flow rates between 1000 and 2000
mL/min should cause little problem. In all cases,            Major ions
the major impact of excessive flow rates will be              Calcium                          1,000
excessive wastage.                                            Nitrate+ nitrite                  100
                                                              Nitrite alone                      10
                                                              Sulphate                         1,000
Table 12-2. Recommended Nipple Drinker Flow
Rates for Various Classes of Swine.                           TDS                              3,000
                                                           245
Table 12-4. Assays to Include in the Chemical               pH
Evaluation of Water Used in Pig Production.                   pH is rarely a concern because the vast majority
                                                            of samples fall within the acceptable range of 6.5 to
  Primary Test                     Supplementary            8.5. If elevated, pH impairs the efficiency of
                                            Test            chlorination; low pH may cause precipitation of
  Total Dissolved Solids                 Sulphates          some medications delivered via the water system.
  pII                                      Sodium           This could lead to carcass residues of these same
  Iron                                 Magnesium            drugs, if the precipitate is later carried in the water
  Hardness                                Chloride          to pigs nearing market weight.
  Nitrates/nitrites                       Calcium
                                        Potassium           Hardness
                                       Manganese           Hardness is caused by multi-valent metal cations,
                                                         the most abundant in groundwater being calcium
                                                         and magnesium. Although it has no effect on
                                                         animal health, hardness does impair the cleansing
   All minerals elevate TDS. but only a few cause        ability of water and also results in the accumulation
health problems; for example, calcium and magne-         of scale in water delivery and treatment equipment.
sium contribute both to hardness and TDS, but            Thus, hardness can lead to problems associated with
unless present at extremely high levels or in associa-   inadequate water supply if nipples, filters, etc.
tion with sulphate, have no adverse effect on pig        become plugged. Water is considered soft if
health. Sulphates, a major contributor to TDS in         hardness is below 50 ppm; if above 300 ppm, it is
some regions, can result in severe diarrhea at even      deemed very hard.
moderate levels. One can conclude that ifTDS is
low, the water is of good quality. lfTDS is moder-       Chloride
ate to high, it may still be perfectly safe to drink.      Chloride is not commonly elevated in
depending on what minerals are responsible. For          groundwater on the Prairies. If chloride, as sodium
this reason, ifTDS is above 1000 ppm, the supple-        chloride (salt), is elevated. it can be countered by
mentary tests are required to determine if those         adjusting the salt content of feed; this must be
minerals responsible for the high reading represent a    carried out by a qualified nutritionist to avoid
significant health risk to the pig.                      problems.
                                                      246
Table 12-5. Effect of Aeration and Seven Days            magnesium and sulphates present. As mentioned
Settling on Water Chemistry (mgfL).                      above, magnesium, along with calcium, is also used
                                                         to calculate total hardness.
Item                            Dai
                       0                    7             Sodium
pH                   7.92                 8.06               Sodium sulphate, also know as Glauber's salts, is
ms                   2388                 2378            a laxative and cathartic agent. By themselves,
Hardness              761                  760            magnesium and sodium normally pose little risk to
Alkalinity            408                  405            the pig, but their association with sulphate is a
                                                          major concern.
Sulphates             1268                1248
Sodium                 446                432             Manganese
Calcium                183                 189              Like iron, manganese can also cause staining of
Magnesium               75                  72            laundry and bathroom fixtures. The threshold level
Chloride                40                 45             of manganese is 0.05 ppm. Like iron, manganese
Potassium               10                  11            exists in groundwater in the soluble form and is
Nitrates                L.0                1.0            precipitated only when pumped to the surface.
Nitrites               0.3                 0.3
Iron                   2.5                 0.6            Nitrates
                                                            Many people are concerned about nitrates and
Source: Tremblay et al., Prairie Swine Centre Annual      nitrites in their water. For human infants, their
Report, 1989.                                             concern is well-founded since nitrates can be a
                                                          serious problem. The primary concern relates to the
  Iron in the water can also support the growth of        reduction in the oxygen-carrying capacity of the
iron bacteria, if the well has been contaminated;         blood. Among livestock, cattle are more suscepti-
however, not all wells with iron in the water will be     ble to the problem than pigs, because the bacteria in
contaminated with iron bacteria. Iron bacteria can        the rumen convert nitrate to the much more danger-
cause foul odours and in extreme cases, can actually      ous nitrite.
plug water systems, including the well itself. Such
foul odours and blockage is caused by the accumu-           Nitrates are often a man-made problem. They are
lation of dead and decaying bacteria.                     formed by the decomposition of organic material,
                                                          and can percolate through soil and into a well at up
  Shock chlorination may solve the problem, but in        to 1 metre (3 ft) per day! They are also persistent.
many cases, the benefits are temporary and the            Examples of barns tom down three decades earlier
procedure may need to be repeated on a regular            still causing nitrate problems have been
basis. One method is to mix 25 L of household             documented.
bleach with 900 L of water; this mixture is poured
into the new well and left there overnight. If              Nitrates as well as nitrites in the water have been
necessary, the chlorinated water can be drawn             shown to impair the utilization of vitamin A by the
through the pipes by turning on a tap at the farthest     pig and reduce overall animal performance. llow-
location from the well. Such water cannot be              ever, the levels required to affect performance were
consumed, so all waterers must be turned off until        so much greater than observed in most water that
the system is thoroughly flushed the following day.       there is little cause for concern under practical
                                                          conditions. Although pork producers should be
Magnesium                                                 cognizant of the potential hazards of nitrates to
   Magnesium sulphate, also known as Epsom salts,         livestock, they pose a greater risk to humans who
is undesirable in water because of associated             consume contaminated water.
diarrhea. Typical water analysis will not define the
amount of Epsom salts present, only the amount of
                                                        247
Sulphates                                                    Table 12-6. Effect of Water Quality on the
   Sulphates arc a primary source of problems                Performance of Weanling Pigs Fed a Diet Containing
associated with well water quality. Sulphates are            Antibiotics.
not well-handled by the gut of the pig, resulting in
diarrhea and possibly reduced performance. Wean-                                                  TDS <m2m)
ling pigs are most susceptible, but in severe cases,         Variable                     217       2350    4390
even adult sows have been afflicted. Depending on                                            Water quality, ppm
the level of sulphate in the water, pigs can adapt and         Sulphates                  83         1280     2650
over a period of a few weeks, become accustomed                Calcium                    24          184      288
to the water. This may explain why pigs at weaning             Chloride                    8           34       70
appear to be most susceptible, since prior to wean-            Magnesium                  15          74        88
ing, they have likely consumed very little drinking            Sodium                     24         446       947
water.
                                                               Hardness                   124        767      1080
Impact of Water Quality                                        pH                         8.4        8.1       8.0
  What effect does poor quality drinking water have
on pigs? Diarrhea is certainly the major concern,            Performance
especially in the weanling pig (adult swine may be           Avg daily gain, kg/day      0.43        0.43     0.44
effected in severe cases); the impact of water               Avg daily feed, kg/day      0.55        0.56     0.57
quality on animal performance remains                        Feed:gain                   1.28        1.31     1.30
controversial.
                                                             Avg daily water, kg/day' I.60           1.84     1.81
  Clinical reports suggest a major impact of poor            Scour score1             1.07           l.30     1.46
quality water on animal performance on some
farms. Yet, this has not been supported by control-
led research, which has consistently concluded that          NB. Pigs were weaned at 28 days of age and immediately
pigs have considerable ability to handle water of            placed on test for 21 days; feed and water were offered
                                                             free choice. Source: McLeese et al. 1991.
widely varying quality with no effect on perform-            1
                                                               Effect of water source significant, P<0.05.
ance. The observation of diarrhea associated with
poor quality water has never been questioned; the
impact on pig performance has.                             In the second experiment (Table 12-7), the starter
                                                         diet did not contain any antibiotic, and in this
   How does one proceed? The results of experi-          instance, the pigs on the poor quality water grew
ments summarized on Tables 12-6 to 12-8 provide          slower and less efficiently than those on the good
useful information. Tn the first experiment (Table       water. Finally, when pigs were given either good or
12-6), pigs received either good quality water or        poor quality water and exposed to either a normal
one of two samples obtained from local farms; in         or chilled environment (Table 12-8), the cold stress
all cases, the pigs pcrformed very well, despite the     did not appear to make the pigs more susceptible to
fact that sulphate was as high as 2650 mg/L.             the effects of the water. In fact, pigs actually grew
                                                         better on the poor quality water, apparently due
                                                         primarily to increased feed intake!
                                                       248
Table 12-7. Effect of Water Quality on the Perform-              Response to Poor Quality Water
ance of Weanling Pigs Fed a Diet Free of Antibiotics.
                                                                   ln response to water problems, some changes to
                                                                 the diet might be warranted. These must always be
                                        T.D.S.
                                                                 carried out with great care. Following are some
Item                         217 ppm              4390 ppm
                                                                 changes that are commonly adopted in response to
Avg gain (kg/day)        0.418                     0.360         water problems.
Avg feed intake (kg/day) 0.530                     0.521
Feed:gain1                1.33                      1.47         Dietary Salt Reduction or Removal
                                                                   This is a common practice on farms using water
NB. Pigs were weaned at 28 days and immediately placed           containing a high mineral load. Partial removal of
on test for 21 days. Feed and water were available free          salt can almost always be done without fear of
choice. Source: Mcleese et al., 1991
1
  Effect of water significant, P< 0.05)                          difficulty because most diets contain a reasonable
                                                                 safety margin. However, complete removal of salt
                                                                 from the feed must be done with great care, because
Table 12-8. Effect of Pen Temperature and Water                  removing dietary salt removes both sodium and
Quality on the Performance of Newly-weaned Pigs.                 chloride. While most water that is high in sulphate
                                                                 often contains high levels of sodium, the chloride
                           Environment       Water Quality       content of water on the Prairies is usually very low.
                                                                 Consequently, removing too much salt from the
                         Normal Chilled Good Bad
                                                                 feed could result in a chloride deficiency. There-
Avg gain/day, g12    1574            1057         960   1671     fore, careful water analysis must attend any adjust-
Avg feed/day, g1     2106            2170        1848   2428     ment of dietary salt levels. Furthermore, if salt
Avg water intake, s' 9632            7628        7382   9878     adjustment is employed, water analysis must be
Gain: feed ratio2     708             366         434    639     repeated on a regular basis to ensure that the min-
                                                                 eral content of the water has not changed.
Pigs were weaned at 28 days and immediately placed on
test for 10 days. Feed and water were available free               If the dietary salt is altered inappropriately and a
choice. Source: Maenz et al.. 1993                               chloride deficiency results, depressed appetite will
I   Effect of water significant, P<0.05                          occur. If severe, it could cause a greater production
2
    Effect of environment significant, P <0.05                   loss than the water! Interestingly, the beneficial
                                                                 effects of salt removal may be related to a marginal
                                                                 chloride deficiency which reduces feed intake and
  From the above, and from research conducted                    thus is "seen" to reduce diarrhea!
elsewhere, it is clear that pigs can perform very
well, even in the presence of very high levels of                Lowering Nutrient Density in the Diet
sulphate. Interestingly, in every case, scouring was               Lowering diet nutrient density has proven suc-
clearly related to water quality, confirming clinical            cessful in the case of the weanling pig, where the
observations. What may be most significant to the                stress of weaning combined with water proves to be
industry is that the presence of scouring should not             too much. Adding bulky ingredients, such as beet
be interpreted as causing impaired growth and                    pulp or even oats or barley appears to reduce the
productivity. Therefore, pork producers,                         visible signs of scouring, but also greatly reduces
veterinarians and nutritionists are cautioned to                 the growth potential of the pig. As with the case of
separate the effect of drinking water quality on                 salt removal, the cure may be worse than the dis-
diarrhea from that on animal performance and                     ease, so such dietary changes must be carried out
determine if indeed, performance is compromised.                 with caution.
This is critical as it has a major impact on how one
responds to a water quality problem. It appears to
run counter to popular thought, but the results of so
many experiments cannot be ignored.
                                                               249
Improved Animal Management                                      Chlorination is performed to provide disinfection
   A more desirable approach would be to lessen all           and remove deleterious bacteria and other disease-
stresses on the pig, by improving the overall                 causing microorganisms. It must be recognized that
environment for the pig (drafts, humidity, crowding,          protozoa and enteroviruses are much more resistant
disease, etc). This enables the pig to withstand the          to chlorination than arc bacteria; it is not altogether
effects of the water with less impact on its health           clear if chlorination is unsatisfactory in this regard,
and productivity. Jmproving housing, for example,             but there is definitely cause for concern.
is generally easier than altering the make-up of the
water, and would, in most cases, improve                        The effectiveness of disinfection and the quantity
productivity.                                                 of chlorine required in the water will depend on the
                                                              quantity of nitrites, iron, hydrogen sulphide,
Water Delivery                                                ammonia and organic matter content, as well as pH
  It is entirely possible that many of the problems           and temperature. For example, the higher the pH,
associated with water quality are related to its              the more chlorine must be added to achieve the
delivery. Poor quality water often plugs screens in           same degree of disinfection. The presence of
drinkers, leaves scale in pipes and generally disrupts        organic matter in the water converts the free
the flow of water from the well to the animals.               chlorine to chloramines which have less disinfecting
Poor delivery systems which actually restrict the             action. The length of contact will also influence the
availability of water to the animals is a serious             demand for added chlorine.
problem on many farms.
                                                                Water softeners of many types are available. The
Water Treatment                                               most common is the ion-exchange unit, which
  If the problem is excess sulphates, treatment may           replaces calcium and magnesium with sodium. This
not be an option due to cost. Reverse osmosis will            reduces the hardness of the water, but clearly has no
remove sulphates, but both the capital and operating          effect on overall mineral load. Because of sodium's
costs for a livestock unit are very high and the cost         role in the process, it is important to recognize that
relative to benefit must be carefully balanced.               water softened in this manner will have elevated
                                                              levels of sodium.
lncreasin� nipple flow rates docs little to increase water intake in pigs but greatly increases water wastage.
                                                           250
Additional Reading and References
Brooks, P.H. and J. L. Carpenter. 1993. The water
requirement of growing-finishing pigs - theoretical
and practical considerations. In (D.J.A. Cole, W
Haresign and P.C. Garnsworthy, Eds.) Recent
Developments in Pig Nutrition 2. Butterworths,
London, pp. 179-200
                                                      251
252
Appendix I. Typical Nutrient Composition of Selected Feed Ingredients
                          Alfalfa Meal     Barley-     Barley-    Beet Pulp   Blood Meal Bone Meal
                                            Grain      Hulless                Spray Dried
Energy, kcal/kg
 -Digestible                  1850          3100           3250     3000         3050
 -Metabolizable               1675          2960           3100     2850         2810
Proximate analysis, %
  -Crude protein              17.0          10.6           13.7      9.9         84.0       28.0
  -Crude fibre                25.6           5.1           3.6      15.3          0.8
  -Acid detergent fibre       29.2           7.1            1.2     17.7
  -NDF                        39.6          17.8
 -Ash                          9.5          2.4                                  4.4
 -Ether extract                2.8           1.8                                 1.0
Macrominerals, %
 -Calcium                     1.33          0.07           0.24     0.70         .40       30.00
 -Phosphorus                  0.23          0.35           0.37     0.08         .30       12.50
 -Avail. phosphorus             0           0.11
 -Magnesium                   0.30          0.11           0.21     0.25          .15       0.30
 -Potassium                   2.27          0.55                    1.00          .15       0.10
 -Sodium                      0.09          0.03                    0.19         0.40       5.00
 -Chloride                    0.46          0.13                    0.25         0.25
Microminerals, ppm
 -lron                         330           90                     275                     45
 -Copper                        10            7             9        13                      IO
 -Zinc                          21           24                       I                      12
 -Mangancse                     37           20            44        35                       5
 -Selenium                     0.6           0.2                    0.04
                                                     253
                           Buckwheat Canola Meal Canola Seed    Com-         Com         Com Gluten
                                                                Grain   Distillers Grain   Meal
Energy, kcal/kg
 -Digcstiblc                  3010         3100          4750   3550        3450          4150
  -Mctaboliz-
able                          2810         2900          4475   3360        3260          3840
Proximate analysis, %
  -Crude protein              11.2         37.7          20.7     8.5        27.6         60.0
  -Crude fibre                13.3         11.8           7.0     2.2        12.0          3.1
  -Acid detergent fibre       20.0         17.2                   3.4
  -NDF                                                           12.0
  -Ash                         2.1         4.8                    1.4        2.2           2.8
  -Ether extract               2.5         3.5                    3.6        9.0           2.4
Macrominerals, %
 -Calcium                     0.08         0.63          0.39   0.02         0.12          0.06
  -Phosphorus                 0.32         1.01          0.64   0.25         0.65          0.38
  -Avail. phosphorus                                            0.03
  -Magnesium                  0.10         0.51          0.38   0.11         0.15         0.07
  -Potassium                  0.45         1.22          0.73   0.30         0.43         0.24
  -Sodium                     0.05                              0.01         0.40         0.07
  -Chloride                   0.05                              0.04         0.18         0.06
Micromincr-als, ppm
  -lron                        40          142           95      20          200           230
  -Copper                       9           6             4       3           50            15
  -Zinc                         9           69           37      15           55            30
  -Mangancse                   34           49           22       4           23             8
  -Sclenium                                 I. I         0.6    0.05         0.35          0.8
                                                   254
                                  Fababeans     Feather Meal     Fish Meal-    Fish Meal-      Fish Meal- Flaxseed Meal
                                                                  Herring'     Menhaden'         Whitc1
Energy, kcal/kg
 -Digestible                         3150           2250             3725          3700           3550           4550
 -Metabolizable                      2960           2070             2625          2930           2385           4250
Proximate analysis,%
  -Crude protein                     26.7           85.0             71.0          61.2           63.8            25.3
  -Crude fibre                        7.8            1.4              0.8          0.9             0.5             9.5
  -Acid detergent fibre              10.0
  -NDF
  -Ash                                6.4            3.7             10.7          19.5           22.7
  -Ether extract                      1.2            2.5              9.4          9.9             5.4
Macrominerals, %
 -Calcium                            0.12           0.34             2.75          5.11           6.97           0.20
 -Phosphorus                         0.49           0.53             1.75          2.92           3.94           0.70
 -Avail. phosphorus
 -Magnesium                          0.13           0.13             0.18          0.14           0.20           0.40
 -Potassium                          1.08           0.20             l.22          0.72            I.OJ          0.80
 -Sodium                             0.08           0.35             0.81          0.45           l.l3           0.05
 -Chloride                                          0.23             0.83          0.55           2.00           0.04
Microminer-als, ppm
    -lron                             65             76              110           443             120
    -Copper                            4              7               5             11              s              12
    -Zinc                             42             72              135           150             80
    -Manganese                        12              IS              7             35             18
    -Seleniurn                                      0.90             2.0           2.1             1.5            0.4
I
    Published Ileal digestibilities for fish meal do not differentiate the source of the product. Variation is likely.
                                                               255
                             Lentils      Meal Meal   Meat and Molasses-Beet Oats Grain   Oats Naked
                                                      Bone Meal
Energy, kcal/kg
 -Digestible                 3065           2850            2825    2475        2800        3600
 -Metabolizable              2865           2585            2570    2350        2650        3420
Proximate analysis, %
  -Crude protein              24.6          55.0            50.0     7.1        10.8         13.2
  -Crude fibre                 3.9           1.3             2.7     0.0        11.0          3.6
  -Acid detergent fibre                                      4.1     0.0        15.3
  -NDF
 -Ash                                       23.2            31.9     8.9         3.4         2.0
 -Ether extract                             10.1             8.4     0.0         4.2         8.7
Macrorninerals, %
 -Calcium                     0.08          8.10            9.50    0.10        0.10         0.12
 -Phosphorus                  0.33          3.63            4.70    0.02        0.35         0.40
 -Avail. phosphorus
 -Magnesiurn                  0.94          0.30            0.30    0.20        0.15         0.11
 -Potassium                                 0.62            0.65    4.60        0.40         0.45
 -Sodium                      0.04          1.20            0.80    1.20        0.05         0.05
 -Chloride                                  0.90            0.70    1.20        0.10         0.08
  Macrominerals, ppm
 -Iron                         88           450             500      70          75           75
 -Copper                        7            10              10      15           5            9
 -Zinc                         46            80             100      15          30           36
 -Manganese                                  10              10       5          40           44
  -Selenium                                 0.4             0.4                  0.3
                                                      256
                              Peas        Potatoes    Poultry By-   Rye Grain   Screenings   Screenings
                                           Dried     Product Meal                #1 Feed     #1 Wheat
Energy, kcal/kg
 -Digcstible                 3400          3350            3250       3300        3100         3250
  -Metabolizable             3175          3160            3020       3085        2930         3075
Proximate analysis, %
  -Crude protein              23.4          7.7            60.0       11.5         11.2         12.9
  -Crude fibre                 5.5          5.1            2.5         2.4          4.9          5.5
  -Acid detergent fibre        8.2                                                  5.9          4.4
  -NDF                        14.7
  -Ash                         3.3          3.3            17.0        1.6
  -Ether extract               1.3          0.2            13.0        1.6
Macrominerals, %
 -Calcium                     0.09         1.20            4.30       0.06        0.07         0.04
 -Phosphorus                  0.50         0.85            2.30       0.32        0.29         0.32
 -Avail. phosphorus           0.15
 -Magnesium                   0.13         0.13            0.39       0.12        0.11         0.13
 -Potassium                   1.01         1.70            0.40       0.45        0.64         0.30
 -Sodiurn                     0.04         1.85            0.82       0.02        0.03         0.02
 -Chloride                    0.05         l.50            0.54       0.03        0.15         0.05
Microminer-als, ppm
 -Iron                        60            14             450         75          57           30
 -Copper                       9            45              14          6           7            6
 -Zinc                        28             3             120         32          24           36
 -Manganese                   23             4              II         57          21           29
 -Seleniurn                   0.4          0.06                       0.03         0.2          0.1
                                                     257
                           Skim Milk      SBM-     SBM-        Soybean     Sugar     Sunflower
                            Powder         44%      47%       Seeds, Raw           Meal - Dehulled
Energy, kcal/kg
 -Digestible                 3850         3500         3675     4200       3800         3115
 -Mctabolizablc              3565         3190         3350     3875       3610         2920
Proximate analysis, %
  -Crude protein              33.4        44.3         47.5      37.2                   40.5
  -Crude fibre                 0.0         6.9          3.9      17.2                   14.9
  -Acid detergent fibre                    8.3          7.4       8.0
  -NDF                                    12.0          9.3
  -Ash                        8.0          6.4          6.5      4.0                     6.8
  -Ethcr extract              1.8          0.7          1.4      15.0                    5.0
Macrominerals, %
 -Calcium                     1.20        0.29         0.29      0.25                   0.42
 -Phosphorus                  1.00        0.61         0.60      0.59                   0.90
 -Avail. phosphorus
 -Magncsium                   0.12        0.26         0.30      021                    0.70
 -Potassium                   l.55        1.90         2.10      1.70                   1.10
 -Sodium                      0.55        0.04         0.01      0.02                   0.20
 -Chloridc                    0.95        0.03         0.03      0.02                   0.20
Microminerals, ppm
 -Iron                         15         140           125       80                     30
 -Copper                       11         25            20        16                      4
 -Zinc                         40          55            60       26                    100
 -Mangancse                     3          30            40       30                     20
 -Selenium                    0.10        0.10         0.10      0.10                   0.10
                                                 258
                            Triticale      Wheat Grain Wheat Bran Wheat Shorts Whey Powder
                                             HRS
Energy, kcaVkg
 -Digestible                  3375            3425          2475      3140       3350
 -Metabolizable               3150            3210          2310      2905       3150
Proximate analysis, %
  -Crude protein               13.6           13.5          15.5      17.5        14.0
  -Crude fibre                  2.5            2.7          10.5       7.5         0.0
  -Acid detergent fibre                        3.5          13.1                   0.0
  -NDF                                        10.8          35.0
  -Ash                                         1.9           6.2      8.2         9.1
  -Ether extract               1.5             1.8           3.6      4.2         0.7
Macrominerals, %
  -Calcium                    0.05            0.04          0.12      0.10        0.98
  -Phosphorus                 0.32            0.34          1.16      0.85        0.79
  -Avail. phosphorus
  -Magnesium                  0.12            0.14          0.53      0.27        0.13
  -Potassium                  0.47            0.32          1.23      0.90        1.62
  -Sodium                     0.03            0.02          0.05      0.03        1.57
  -Chloride                   0.03            0.05          0.05      0.04        1.50
  Microminerals, ppm
  -lron                        30              31            150        85         140
  -Copper                       9               6             11        12         45
  -Zinc                        45              38             95       105          5
  -Manganese                   25              30            110       110          5
  -Selenium                                   0.10          0.60      0.50        0.06
                                                      259
                             Canola Oil          Lard         Poultry Grease    Soybean Oil         Tallow
Energy, kcal/kg
 -Digestible                    8800             7850             8625              8800             8200
 -Mctabolizable                 7300             7550             8200              7275             7900
Disclaimer:Every attempt has been made to provide accurate information on each ingredient. Due to the nature of
these materials. variability among samples will occur.
                                                        260
Appendix II. Calibrating Proportioner-type Mills
l. Ensure that the dial settings for the mill are         3. Tum on the mill and collect the ingredients until
correctly set for the diet being tested. Record the       a reasonable amount is added to all containers. The
setting for each dial.                                    more material that can be collected properly, the
                                                          more accurate the test will be.
2. Remove the back housing cover from the mill to
expose the proportioning augers and attach the            4. Weight the ingredients in each container and
testing tray. Place collecting containers below the       subtract the weight of the empty container. Be sure
tray to ensure accurate collection of each ingredient     to use a small scale that can accurately weigh
from each auger.                                          ingredients such as premixes and medications.
                                                        261
Useful Conversion Factors
   Length                                             Miscellaneous
   To convert:              Multiply by:              To convert:                  Multiply by:
   Centimetres to inches      0.394                   Calories to joules            4.184
   Inches to centimetres      2.540                   Kilocalories to calories       1000
   Millimetres to inches      0.039                   Megacalories to kilocalories   1000
   Inches to millimetres      25.40                   Joules to calories            0.239
   Feet to metres             0.305                   Percent to grams per kilogram 10.00
   Metres to feet             3.281                   Grams per kilogram to percent 0.100
   Metres to inches           39.37                   Gallons of water to pounds    10.00
   Inches to metres           0.254                   Megajoules to kilocalories     239
   Kilometres to miles        0.621
   Miles to kilometres         1.609
                                                      Temperature
                                                       op     oc            op       oc
   Area                                                212     100.0
   To convert:                 Multiply by:            106      41.l         55      12.8
   Square foot to square metre  0.093                  104      40.0         50      10.0
   Square metre to square foot   10.75                 102      38.9         45       7.2
   Acres to hectares            0.405                  100      37.8         40      4.4
   Hectares to acres            2.469                   98      36.7         35       l.7
                                                        96      35.6         30      -1. l
                                                        94      34.4        25       -3.9
                                                       92       33.3         20      -6.7
   Volume                                              90       32.2         15      -9.4
   To convert:                   Multiply by:          88       31. l        10     -12.2
   Millilitres to fluid ounces    0.033                85       29.4          5     -15.0
   Fluid ounces to millilitres    30.00                80       26.7          0     -17.8
   Litres to quarts               0.880                75       23.9         -5     -20.6
   Quarts to litres                1.137               70       21. l       -10     -23.3
   Gallons to litres              4.546                65       18.3        -15     -26.1
   Litres to gallons              0.220                60       15.6
   Litres to millilitres           1000
                                                262
                                              Common Abreviations
Weights
To convert:                  Multiply by:         Length
Grams to ounces                0.035              Kilometer        km
Ounces to grams                28.35              Meter             m
Kilograms to pounds            2.205              Centimeter       cm
Pounds to kilograms            0.454              Millimeter       mm
Micrograms to milligrams       0.001              Mile             mi
Milligrams to grams            0.001              Yard             yd
Kilograms to tonnes            0.001              Foot              ft
Tonnes to kilograms              1000
Tonnes to tons                 0.984
Tons to kilograms                1102
Tons to tonnes                  1.016
Tons to pounds                  2000
Parts/million to grams/tonne    1.000             Volume
Grams/kilogram to grams/tonne 1000                Litre             L
                                                  Millilitre       mL
$/bushel to $/tonne                               Micro litre      mcL
  (barley)                      45.94             Gallon           gal
  (wheat)                       36.75             Quart             qt
  (oats)                        61.25
  (corn)                        39.37
$/tonne to $/bushel
  (barley)                     0.0218
  (wheat)                      0.0272             Mass
  (oats)                       0.0163             Kilogram         kg
  (com)                        0.0254             Gram              g
                                                  Milligram        mg
                                                  Pound             lb
                                                  Ounce             oz
                                                  Hundredweight    cwt
                                                  Tonne (metric)    T
                                            263
264
Glossary
Abscess - Collection of pus formed by breakdown        Anemia - A condition in which the blood is defi-
of tissues.                                            cient in the amount of needed hemoglobin or in the
                                                       number of red blood cells or in both.
Absorption - The uptake of nutrients through the
intestinal wall.                                       Anion - An ion carrying a negative electric charge.
                                                       Chloride is an anion.
Acclimatization - The process of becoming accus-
tomed to a new climate or other environmental          Anorexia - Lack of loss of appetite.
conditions.
                                                       Anti-nutritional factors - Factors that work against
Acute - Having a short and severe course of devel-     the nutritional value of a feedstuff
opment; opposite of chronic.
                                                       Anti-oxidant - A substance that prevents fats from
Adaptation - The adjustment of an animal to a new      becoming rancid through oxidation.
or changing environmental condition.
                                                       APF (animal protein factor) - original name for
ADF (acid detergent fibre) - Fibre extracted with      vitamin B,r
acidic detergent in a technique used in appraising
                                                       Apparent digestibility - The amount of a nutrient
the quality of forages.
                                                       absorbed from the gut.
ADG (average daily gain) - Rate of body weight
                                                       Arginine - An essential amino acid.
gain, expressed on a 'per day' basis.
                                                       Arthritis - Inflammation of a joint and its adjacent
Adipose - Fat.
                                                       tissues.
Ad lib feeding - See ad libitum.
                                                       As fed - Refers to the nutrient composition of
Ad libitum - Self-feeding, or allowing swine to        feedstuffs; including moisture normally present in
consume feed to appetite.                              the feed at the time it is fed; differs from dry matter
                                                       basis, which defines nutrient concentration after aJl
Agalactia - Absence of milk production by the sow.
                                                       the water is removed.
Albumin -A water soluble plant (and animal)
                                                       Assay - The determination of the chemical compo-
protein.
                                                       sition of a feed or ingredient.
Aleurone - The protein portion of the endosperm of     Ataxia _ Lack of muscle coordination.
a seed.
                                                       ATP - Adenosine triphosphate: a source of energy
Alfatoxin - Any carcinogenic mycotoxin produced
                                                       for the cell.
by molds in stored agricultural crops.
                                                       Atrophic rhinitis - Inflammation of the mucous
Alkyl resorcinol - A colorless crystalline com-
                                                       membranes and turbinate bones of the nose, often
pound used in making dyes and pharmaceutical.
                                                       resulting in distortion in shape or size.
Amino acids - The building blocks of proteins;
                                                       Atrophy - Wasting away of cell or tissue.
hundreds are known, but only about 20 are nor-
mally found in proteins.                               Availability - That proportion of a nutrient that is
                                                       available to the animal.
Amylase - An enzyme that breaks down starches.
                                                       Available amino acids - The proportion of the total
Anabolic - The process by which food is changed
                                                       dietary amino acids that can be absorbed from the
into living tissue. Building tissue; opposite of
                                                       gut of the pig, and thus are actually used for growth
catabolic.                                             and production.
                                                     265
Bacterin -A suspension of killed bacteria (vaccine)             Calorie (cal) - A unit of energy measurement
used to increase disease resistance.                            defined as the amount of heat required to raise the
                                                                temperature of one gram of water from 14.5 to
Bacteria - Single celled living organism that
                                                                 15.5°C. Equivalent to 4.184 joules.
multiplies by simple division; some arc beneficial
and others cause disease.                                       Calorimetry- Measurement of the amount of heat
                                                                produced during normal metabolism.
Barrow - A young castrated male pig.
                                                                Capillary - Small blood vessel delivering arterial
Basal (energy) feeds - A group of grain and grain
                                                                blood to the tissues and venous blood from the
by-products containing not more than 16% CP and
18% CF.                                                         tissues. Walls of the capillaries are in close contact
                                                                with individual cells of the tissues.
Basal metabolism (BM) - The chemical changes
                                                                Carbohydrates - A class of compounds in the feed,
that occur in the pig in the fasting and resting state,
                                                                including starches, sugar, cellulose, and gums.
when it uses just enough energy to maintain basic
metabolic activity.                                             Casein - The major protein of milk.
Beta carotene - A source of vitamin A found in                  Catabolism - A breakdown of tissues; opposite of
some plant and plant products.                                  anabolism.
Beta-glucan - A polysaccharide that interferes with             Catalyst - A substance that increases the rate of a
digestion.                                                      chemical reaction. Enzymes are catalysts. Cata-
                                                                lysts are absolutely essential for the normal body
Bile salts - Compounds released from the gall
                                                                processes.
bladder into the intestine which help emulsify and
digest fats.                                                    Cathartic - A compound that acts as a laxative.
Bioassay - Using animals to evaluate feed quality.              Cation - An ion carrying a positive electrical charge
                                                                (i.e. sodium, potassium, and calcium).
Biopsy - The colJection and analysis of tissue
collected from alive animal.                                    Cecum, ceca - See caecume.
Biotin -A vitamin found in high levels in liver, egg        Cellulose - The carbohydrate constituent of plant
yolk and yeast.                                             cell walls that are resistant to normal digestive
                                                            processes in the pig.
Boar - Uncastrated male pig.
                                                            Chelating agent - A compound that can bind
Bomb calorimeter - An apparatus for measuring
                                                            mineral ions to improve their utilization by the pig.
the gross energy content of feed.
                                                            Cholecalciferol - Vitamin D3•
Bone meal (steamed) - Ground animal bones that
are steamed under pressure. It can be used as a             Coenzyme - A substance, usually a vitamin or
source of calcium and phosphorus.                           mineral, which works with an enzyme to help it
                                                            catalyze (speed up) metabolic processes.
Bran - The seed coat of cereal grains.
                                                            Colitis - An inflammation of the colon.
Brewer's grains - A by-product of the brewing
industry.                                                   Colon - The lower portion of the large intestine.
Bushel - Eight-gallon measure.                              Colostrum - The first milk, produced by the sow
                                                            during the first few days of lactation; rich in fats
Caecum (cecum) -A section of the large intestine
                                                            and antibodies essential for piglet survival.
in which bacterial action breaks down some fibre
that escaped digestion i the small intestine.               Comfort zone - The temperature range in which
                                                            pigs will not expend energy to keep warm.
Calciferol - Commonly known as vitamin D2"
                                                          266
Complete feed - Provides all the nutrients required      Digestion - The processes which feed undergoes
except water. A balanced diet.                           within the gastrointestinal tract to prepare it for
                                                         absorption.
Condition - Refers to the amount of flesh (fat) on
the body; the general health of animals.                 Diuresis - Increased urination.
Congenital - Characteristics of the fetus acquired       DL-methionioe - Synthetic source of methionine.
before birth, i.d. during the gestation period.
                                                         DNA - Deoxyribonucleic acid; also called the
Connective tissue - Tough fibrous tissue that            blueprint of life because it directs cells in the body
supports and connects tissues of an animal body.         to build proteins in certain ways.
Creatine phosphate - A storage form of energy for        Dressing percent - The portion of the carcass
cell metabolism.                                         remaining after removal of most internal organs,
                                                         feet, and in some cases the head.
Critical temperature - Environmental temperature
below which extra energy is required to maintain         Duodenum - The first portion of the small intes-
the pig's body temperature, and less energy is thus      tine.
available for growth purposes.
                                                         Ear notches - Slits or perforations in an ear used
Crude fibre (CF) - That portion of feedstuffs            for identification.
composed of cellulose, hemicellulose, lignin, and
                                                         Eczema - Skin disease characterized by redness,
other complex carbohydrates.
                                                         itching, loss of hair, and the formation of scales.
Crude protein (CP) - An estimate of the protein in
                                                         Edema - Swelling due to accumulation of fluid.
a feed. In calculating the protein percentage. the
feed is first chemically analyzed for its nitrogen       EFA (essential fatty acid) -A fatty acid that
content. Since proteins average about 16% (1/6.25)       cannot be synthesized in the body in sufficient
nitrogen, the amount of nitrogen in the analysis is      quantities for the body's needs.
multiplied by 6.25 to give the CP percentage.
                                                         Electrolyte - Any charged article: an ion.
Crumbles - Pelleted feed reduced to granular form
with corrugated rollers.                                 Electrolyte solution - A solution of simple sugars
                                                         and minerals often used in the treatment of scours.
Cyanocobalamine - Vitamin B12•
                                                         Endogenous - Internally produced in the body,
Cyst - A sac or bag-like structure, especially one       such as hormones.
that contains a liquid or semi-solid material.
                                                         Endogenous proteins - Sloughed cnterocytcs
Cystine - Amino acid that can replace up to one          together with enzymes.
half of a pig's requirement for methionine.
                                                         Endosperm - Part of the seed which provides food
Dermatitis - An inflammation of the skin.                for the developing embryo.
Dextrose - Glucose.                                      Endotoxins - Toxic substances stored inside bacte-
                                                         rial cells.
Dicoumarol - A chemical compound found in
spoiled sweet clover, which acts as an anticoagulant     Enteritis - Inflammation of the intestinal tract.
causing internal hemorrhages when eaten.
                                                         Enterotoxin - A toxin produced by microorganisms
Digestibility - A measure of the extent to which a       that disturbs the gastrointestinal tract.
feed is digested and absorbed by the animal.
                                                         Enzyme - Biological catalysts that increase the rate
Digestible energy (DE) - Gross energy minus fecal        of chemical reactions.
energy; generally pertains to feeds.
                                                         Enzyme activator - A substance which the enzyme
                                                         requires in order to be active.
                                                       267
Epidemiology - The study of relationships of                 Feed conversion - Amount of feed used per unit of
various factors that interact to cause or prevent            gain.
disease or other health related problems.
                                                             Folacin - Compounds derived from folic acid.
Ergocalciferol - Vitamin 02•
                                                             Gastric - Referring to the stomach.
Ergosterol - A plant sterol that (upon activation by
ultraviolet rays) becomes vitamin 02•                        Gastrointestinal - Referring to the stomach and
                                                             intestines.
Ergot - Undesirable fungi found in rye.
                                                             GE - Gross energy.
Ergonovine - A toxic alkaloid found in ergot.
                                                             Gluconeogenesis - Formation of glucose.
Ergotamine - A toxic alkaloid found in ergot.
                                                             Glutelin - A cereal protein.
Ergotoxine - A toxic alkaloid found in ergot.
                                                             Glycolysis - Degradation of simple sugars.
Erythropoieses - The production of red blood cells.
Occurs in bone marrow.                                       Goiter - Enlargement of the thyroid gland caused
                                                             by iodine deficiency.
Esophagus - Structure extending from mouth to
stomach.                                                     Gossypol - A toxic yellow pigment found in
                                                             cottonseed.
Essential amino acid - An amino acid that cannot
be synthesized in the body in sufficient quantities          GRAS - "Generally recognized as safe". Used by
for the body's needs; therefore required in diet.            the USDA as related to feed or feed ingredients.
Estrogen - A female sex hormone; promotes estrus             Gross energy (GE) - The amount of heat released
and stimulates the development of female second-             when a substance is completely oxidized (burned).
ary sex characteristics.                                     Gut - The digestive tract.
Estrus - Heat.                                               Heat increment (HI) - Heat of nutrient metabo-
Estrous cycle - The recurring sexual cycle.                  lism. This heat may be used to keep the body
                                                             warm.
Ether extract - Substances in feeds which are
soluble in ether. Used in feed analysis to estimate          Hemagglutinin - An antibody which causes red
fat content of a feed.                                       blood cells to stick together.
Exogenous - Produced or supplied from outside the        Hemoglobin - A protein that imparts a red color in
body.                                                    red blood cells.
F 2 generation - The second generation resulting         Histidine - The mechanisms by which animals
from the mating of F1 individuals.                       maintain a constant internal environment.
Fagopyrin - Photosensitizing agent found in              Homeostasis - The mechanisms by which animals
buckwheat.                                               maintain a constant internal environment.
Fagopyrism - Buckwheat poisoning.                        Hormone - A substance, secreted by one gland,
Farrow - To give birth.                                  which has an effect on other tissues.
Fat soluble vitamins - Vitamins A, O, E and K. HRS Wheat - Hard red spring wheat.
                                                       268
Hybrid vigor - Increased stamina or vitality of             Megacalorie (Meal) - Unit of energy equal to 101'
crossbred animals.                                          joules.
Hydrocephalus - Accumulation of fluid on brain.             Menadione - Vitamin K.
Ileum - The lower portion of the small intestine.           Metabolic body size - The weight of the animal
                                                            raised to the 3/4 power (W01s).
Immunoglobulins - Proteins (also known as
antibodies) that produce a state of immunity.               Metabolism - The sum total of the chemical
                                                            changes in the body, including building up {anabo-
International units (IU) - An arbitrary scale. used
                                                            lism) and breaking down (catabolism).
to compare sources of vitamins.
                                                            Metabolizable energy (ME) - Gross energy minus
Intestine - The portion of the gastrointestinal tract
                                                            fecal energy and urinary energy.
from the stomach to the anus.
                                                            Metabolite - Substance produced by metabolism.
intrinsic factor - A substance secreted by the
stomach which allows absorption of vitamin B1�.             Microvilli - Projections from the villi.
In utero - Within the uterus.                               Middlings - A by-product of flour milling contain-
                                                            ing endosperm, bran, and germ.
.n vitro - Outside the animal in an artificial environ-
ment such as a test tube.                                   Mycotoxicosis - Poisoning due to fungal or bacte-
                                                            rial toxin.
in vivo - Within the living body.
                                                            Mycotoxins - Toxic substances produced by molds.
Ion - Charged molecule.
                                                            Myrosinase - An enzyme found in rapeseed meal
Kcal (kilocalorie) - A unit of energy equal to 1000
                                                            capable of breaking down glucosinolates.
calories.
                                                            National Research Council - See NRC.
Keratin - Protein that composes hair, horn, claws,
and feathers.                                               NDF (neutral detergent fibre) - Fibre in the plant
                                                            cell wall, which is undigested by swine. Developed
Kjeldahl -0 A method of determining the quantity
                                                            to evaluate forages for ruminants.
of crude protein based on the nitrogen content.
                                                            Necropsy - Autopsy: postmortem examination.
Lactose - A simple sugar found in milk.
                                                            Necrosis - Tissue death.
Lesion - Change in the structure, color or size of a
part of the body.                                           Neurotransmitter -A substance involved in the
                                                            transmission of signals by the nervous system.
Lignin - A complex carbohydrate that is almost
completely indigestible.                                    Niacin - A vitamin.
Lipase - An enzyme that breaks down fat.                    NFE (nitrogen-free extract) - An approximation
Lipid - Fat.                                                of the carbohydrate content in feeds.
Macro (or major) minerals - Minerals present or             NRC - National Research Council. A division of
required in large amounts e.g., calcium, phospho-           the National Academy of Sciences promoting
rus, sodium, potassium, magnesium, and chloride.            utilization of scientific and technical information.
Mas ti tis - An inflammation of the udder.                  Ochratoxin - A mycotoxin, produced by the mold
                                                            aspergillus, which attacks the kidneys, reduces pig
                                                            performance and may lead to birth defects.
                                                          269
Odd chain fatty acids - Those fatty acids with                Tetany - A condition where muscles become rigid
uneven chain lengths e.g. containing 7, 9 or I l              and have spasms.
carbon atoms.
                                                              Thiamine - Vitamin 81•
Osteogenesis - Formation of bone.
                                                              Tocopherols - One of the compounds that make up
Osteomalacia - Softening of the bones.                        Vitamin E.
Osteoporosis - Reduction in bone mass.                        Tocotrienols - One of the compounds that make up
                                                              Vitamin E.
Pantothenic acid - Vitamin B5•
                                                              Total sulfur amino acids (TSAA) - Methionine
Parakeratosis - Skin disease.
                                                              plus cystine.
Parasite - An organism that lives at the expense of
                                                              Tricothecene - A group of mycotoxins, including
living animals.
                                                              vomitoxin (deoxynivalenol), HT-2 toxin, T2 toxin
Pepsin - A stomach enzyme which breaks up                     and diacetoxyscirpenol, produced by thefusarium
proteins.                                                     molds. They are very toxic compounds, causing
                                                              vomiting, depressed growth and suppression of the
Pepsinogen - Precursor to pepsin;. Secreted by the
                                                              immune system. The pig is especially sensitive.
stomach.
                                                              Ulcer - Erosion or disintegration of tissues; often
pH - Measure of the acidity or alkalinity of a fluid.
                                                              refers to stomach.
pH= 7 is neutral.
                                                              USDA - United States Department of Agriculture.
Phytin - A naturally occurring compound in many
cereal grains containing phosphorus oflow avail-          USP - United States Phannacopeia.
ability to the pig.
                                                          VFA - Volatile fatty acids.
Pneumonia - Inflammation of the lung.
                                                          Villi - Finger-like projections on the surface of the
ppm - Parts per million.                                  gut.
Serotonin -A hormone and neurotransmitter.                Volatile fatty acids - The short chain fatty acids
                                                          such as those produced in the rumen of cattle and
Starch - A white, tasteless, odorless complex             the cecume and colon of swine.
carbohydrate found in large quantities in potatoes,
                                                          Zearalerone - Also called F2 toxin, is a mycotoxin
rice and wheat.
                                                          produced by thefusarium molds.
                                                        270
Index
A
absorption 5, 7, 9, 12, 27                                carcass grading 70, 189, 193, 204
ADF 19                                                    carmelization 29
Aflatoxin 238                                             carotene 42, 43, 68
aflatoxin 236, 238, 239                                   carotenoid 42
albwnen 25, I 00                                          cecum 12, 27
alfalfa 18, 28. 42. 45 63, 232, 233,                      cellulose 19
allergic reaction 5                                       chemical analysis 1, 33, 68, 113, 245
ammonia 13, 27, 252                                       chewing 7
amylase 7, IO                                             chloride 15, 34, 35, 248, 249
anemia 8                                                  choline 41, 46, 150, 163, 199, 232
animal by-products 235                                    chymotrypsin I 0, 86, 91
antioxidant 236                                           cobalt 15, 34, 36, 37, 45
amino acid 22, 28, 147, 159                               colon 12
   synthetic 115                                          colostrwn 82, 111, 167, 168, 169, 172
appetite 9, 22, 34, 40, 46, 47, 48                        comfort zone 143
arginine 15. 24, 26, 31, 148, 149, 160, 161, 176          computer formulation 123, 124
available phosphorus 33, 34. 200                          computers 3
                                                          conception 133, 154, 155, 162, 170
B                                                         condition scoring 143, 144, 146
bacteria 6, 12, 27, 45, 67, 113. 167, 184. 231. 235       copper 15. 32, 36, 113, 150, 163 184, 200. 228, 232
                                                          com 15, 18, 32, 52, 76. 202. 239
barley 18, 65, 126, 130. 132, 183. 197. 218,
      232, 235, 253                                       com gluten feed 78
   high moisture 67                                       com gluten meal 45, 78
   hulless 69                                             com oil I 08, 170
                                                          creep feed 90, 169, 170, 171, 172, 179, 180, 183
barn 18, 119, 129, 141, 157, 200, 213
                                                          criticaltemperature 143, 147, 163, 184
beet pulp 71, 152, 251, 255
bile 9, 10                                                crumbles 171. 180, 221
bile salts 10                                             cyanogenetic glycosides 82
                                                          cystine 22, 29, 31, 36, 80, 85, I 00, 164, 198
biotin 15, 45, 46
bleeding ulcer 8, 38                                      D
blood meal 181, 182
bomb calorimeter 17                                       deflourinated phosphate 112
bone meal 33, 54, 55, 59. 60, 235                         dermatitis 46
buckwheat 71, 72, 85                                      diarrhea 34, 39, 41, 47
bushel weight 2 l. 66, 77. 79. l 02. J 23, 225            dicalcium phosphate 112, 232
                                                          dicoumarol 4 7
c                                                         dietary requirement 46
calciferol 43, 44                                         digestible nutrients 12
calcium 15, 32, 55, 59, 61, 64. 112. 124, 134, 163,       digestive enzymes 7, 8, 9, I 0, 12
      231,234,248,253                                     digestive tract 5, 6, 113. I 14
                                                          DL-methionine 115, 182
calorie 19. l 07
canola meal 24, 28, 33, 45, 52, 73, 83, 126, 130, 197     dressing percentage 58, 70, 189
                                                          dried bakery product 80
canola oil 170
canola screenings 85                                      duodenum 12
eanola seed 74, 75, 110, 11 l                             durum 100
carbohydrate 5, I 0. 12, 16, 19, 20, 32, 39. 45, 47
                                                        271
s                                                       u
saliva 7                                                ulcers 8, 9, 138, 221
salivary enzymes 7
salt 34, 35, 36, 38, 41, 46, 59, 80
                                                        v
sample diet 153, 182, 20 I, 202                         valine 15, 24, 28, 31, 148, 149, 160, 161, 176
saponins 63. 64, 94                                     villi 9, 10. 11, 12
scours 39, 40, 170                                      vitamins 15. 40, 150. 161, 228
selenium 31, 39, 40, 44, 64, 82, I 09, 168                vitamin A 15, 40, 42, 60, 150, 151, 162, 199, 249
self-feed 142, 143, 177                                   vitamins B 15, 36, 45. 47. 48, 150, 162. 163, 199
small intestine 8, 11                                     vitamin C 199
soapstocks I 08                                           vitamin D 15, 34, 42, 43, 44, 47, 199
sodium 34, 35, 36, 40, 47. 248, 249, 251, 252             vitamin E 15, 39, 40. 44, 45. 199. 213
sorghum 47. 72, 232, 233, 234                             vitamin K 42. 47, 199
soybean meal 19, 20, 24, 26, 45, 51, 73. 83, 96, 178      fat soluble vitamins 41
soybeans 27, 45, 93, 109.                                 stability 41
starch 5. 7. IO. 19, 65. 76. 80, 82, 90, 222              supplement 41
stomach 5, I 0, 12, 22, 27, 35, 45, I 03, 116             water soluble vitamins 41
stress 8, 12. 244, 250, 251. 252                        volatile fatty acid 12
sucrose 10                                              vomitoxin 77, 104, 236, 238, 239, 240
sulphur 22, 36, 54, 69, 74, 83, 85, 86                  vulva biting 139
sunflower meal 97. 98. 238
sunflower seed 97, 98. 99, 233                          w
                                                         water 168, l 71, I 77, 241
T
                                                          delivery 244. 250
tallow 59, 107, 108, 180                                  quality 245
tannins 63, 72, 81, 86, I 00, 231, 233. 234              wheat 84, IOI. 201
taste 7                                                   bran 65. 71, 98. 104. 152
thiamine 42, 48                                           middlings 104
threonine 29. 30, 31, 85, 100, 105, 106, 115.             red dog 104
      153. 160, 161 182. 198                              screenings 84
thyroxine 38                                              shorts 104. 105
total sulpher amino acids                                whey 170. 171. 261
toxin 231, 236. 237. 238, 240
tricothccenes 231. 238                                   z
triglyceride 154. 166                                    zearalenone 231, 236. 237. 238. 240
triticale I 00. IO I                                     zein 26. 76
 trypsin 100
 tryptophan 23. 25. 29. 30, 47, 55, 115, 153. 182
                                                       274
Authors
John F. Patience, Ph.D.
Dr. Patience received his B.Sc.(Agr.), majoring in Animal Science and
M.Sc. in Animal Science from the University of Guelph. Following
employment in extension and in the feed industry, Dr. Patience re-
turned to school, earning a Ph.D. in Nutritional Biochemistry from
Cornell University in 1985. He was employed as a Visiting Fellow at
the Animal Research Centre in Ottawa and, in 1987,joined the Univer-
sity of Saskatchewan as a Research Scientist and Director of the Prairie
Swine Centre. When Prairie Swine Centre was incorporated in 1991,
Dr. Patience was appointed its first President and Chief Executive
Officer. His research interests include nutrition of the growing-
finishing pig, dietary influences on acid-base balance and water quality
and requirements.
I I