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Year 1997 1997 Probability of entering study area (pent ) survey (categorical) survey (linear) Probability of staying in study area (phi ) survey (categorical) age Probability of detection AICc (p ) region 492.03 region*survey (categorical) 492.04 Delta AICc 0.00 0.01 AICc Weights 0.41 0.41 Num. Par 14 19 1997 1997 survey (categorical) survey (categorical) age age survey (categorical) 494.41 region*survey (categorical) 497.08 2.38 5.05 0.13 0.03 16 22 1997 1997 survey (linear) survey (categorical) survey (categorical) region age*survey (categorical) region 498.87 511.15 6.84 19.12 0.01 0.00 11 24 1997 survey (linear)+region 511.78 19.74 0.00 17 1997 511.95 19.92 0.00 24 1997 survey (categorical)*region survey (linear)+region survey region (categorical)*region survey region (categorical)*region age*survey (categorical) region 515.67 23.64 0.00 22 1997 1997 survey (linear) survey (linear)+age constant region age*survey (categorical) region 516.88 517.99 24.85 25.95 0.00 0.00 7 23 1997 age age*survey (categorical) region 518.14 26.11 0.00 22 1997 1997 1997 1997 1997 1997 1997 1997 1997 1997 region survey (categorical) age survey (linear) region survey (linear) age survey (categorical) survey (categorical) survey (categorical)*region age*region age*survey (categorical) region constant region age age age age age age age*survey (categorical) region region region region region age region region age region 518.51 520.47 520.63 520.72 520.89 521.50 523.02 523.67 525.72 528.94 26.48 28.44 28.60 28.69 28.85 29.47 30.98 31.64 33.68 36.90 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 8 10 9 9 9 10 10 12 13 29 age region age*survey (categorical) region 529.48 537.58 37.45 45.55 0.00 0.00 13 34 1998 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc study area (phi ) (p ) age region*survey (categorical) 975.40 Delta AICc 0.00 AICc Num. Weights Par 0.97 19 1998 survey (categorical) age region*survey (categorical) 982.46 7.06 0.03 22 1998 1998 1998 1998 1998 1998 1998 survey (linear) survey (linear) survey (categorical) survey (categorical) survey (linear) survey (categorical) survey (linear)+region age age age age survey (categorical) age survey (categorical)*region region age region age region survey (categorical) region 14.32 14.51 15.32 15.72 16.31 16.34 16.92 0.00 0.00 0.00 0.00 0.00 0.00 0.00 9 10 12 13 11 16 17 1997 1997 Year 989.72 989.92 990.72 991.12 991.71 991.75 992.32 1998 1998 1998 1998 1998 1998 1998 1998 1998 1998 constant constant age age constant region survey (categorical) survey (categorical) region age*survey (categorical) 1998 survey (linear) survey (categorical) region age constant region survey (categorical) constant age survey (categorical)*region survey (categorical) 1998 19.41 20.49 22.09 24.16 25.72 26.85 26.89 27.20 28.91 32.38 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 7 10 9 10 6 8 14 10 9 29 age*survey (categorical) region 1009.55 34.15 0.00 24 survey (linear)+region age*survey (categorical) region 1011.00 35.60 0.00 22 1998 age age*survey (categorical) region 1017.02 41.61 0.00 22 1998 survey (categorical)*region survey (linear)+age survey region (categorical)*region age*survey (categorical) region 1017.75 42.35 0.00 24 1021.89 46.49 0.00 23 1999 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc Delta study area (phi ) (p ) AICc age region*survey (categorical) 1358.08 0.00 AICc Num. Weights Par 0.90 19 1999 1999 survey (linear) survey (categorical) constant age region 1363.72 5.64 region*survey (categorical) 1364.14 6.05 0.05 0.04 5 22 1999 1999 1999 1999 1999 survey (linear) survey (linear) survey (categorical) survey (categorical) survey (linear)+region region region region region region 1369.81 1371.90 1372.73 1374.30 1377.15 11.72 13.81 14.65 16.21 19.06 0.00 0.00 0.00 0.00 0.00 9 11 10 12 17 1999 1999 1999 1999 1999 1999 1999 1999 1999 1999 survey (linear) survey (categorical) survey (categorical) survey (categorical) region age constant constant region survey (linear)+region age survey (categorical) constant age survey (categorical)*region age age age survey (categorical) region region survey (categorical) constant age age*survey (categorical) age survey (categorical) age region region region region region region region 1377.19 1379.75 1380.84 1381.32 1381.73 1383.77 1386.64 1388.85 1393.85 1395.43 19.11 21.66 22.75 23.24 23.64 25.69 28.56 30.77 35.76 37.34 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 10 16 13 14 8 9 10 6 9 22 1999 1999 age survey (categorical)*region survey (linear)+age age region survey region (categorical)*region age*survey (categorical) region 1395.90 37.81 1397.74 39.65 0.00 0.00 10 24 1400.83 42.75 0.00 23 1998 Year 1999 region region region region region region region region region region 994.81 995.89 997.49 999.56 1001.12 1002.26 1002.29 1002.61 1004.32 1007.78 1999 survey (categorical) age*survey (categorical) region 1401.96 43.87 0.00 24 1999 1999 age*region age age region age*survey (categorical) region 1402.08 43.99 1411.75 53.66 0.00 0.00 13 22 1999 survey (categorical)*region age*survey (categorical) region 1432.88 74.80 0.00 29 Year AICc Weights 0.45 0.31 0.10 0.07 0.04 0.02 0.00 Num. Par 7 11 14 9 12 10 17 2000 survey (linear) Probability of staying in study area (phi ) constant survey (categorical) survey (categorical) age age constant survey (categorical)*region age AICc 2000 2000 2000 2000 2000 2000 2000 Probability of entering study area (pent ) survey (linear) survey (linear) survey (categorical) survey (linear) survey (categorical) survey (categorical) survey (linear)+region 0.00 19 2000 survey region (categorical)*region age age age*survey (categorical) region 1006.63 14.67 0.00 24 2000 2000 survey (categorical)*region survey (linear) survey (linear)+region 1006.86 14.91 1007.55 15.60 0.00 0.00 10 22 2000 2000 survey (categorical) survey (categorical) age age age 1009.15 17.20 region*survey (categorical) 1009.72 17.77 0.00 0.00 13 22 2000 survey (categorical) age*survey (categorical) region 1009.85 17.90 0.00 24 2000 survey (linear)+age age*survey (categorical) region 1010.84 18.89 0.00 23 2000 2000 age survey (categorical) age*survey (categorical) region 1012.44 20.49 1014.36 22.40 0.00 0.00 16 29 2000 survey (categorical) survey (categorical)*region age*survey (categorical) age*survey (categorical) region 1028.04 36.09 0.00 34 2000 2000 2000 2000 2000 2000 region region age age age*region age region age region age age age*survey (categorical) 1031.34 1033.04 1033.40 1035.12 1041.37 1042.11 39.39 41.09 41.45 43.16 49.42 50.16 0.00 0.00 0.00 0.00 0.00 0.00 8 9 9 10 13 22 Year 2001 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc study area (phi ) (p ) age region*survey (categorical) 859.26 Delta AICc 0.00 AICc Num. Weights Par 0.97 19 2001 survey (categorical) age 7.08 0.03 Probability of detection (p ) region region region region region region region Delta AICc 991.95 0.00 992.74 0.78 994.93 2.97 995.60 3.65 997.06 5.10 997.76 5.81 1002.08 10.13 region*survey (categorical) 1004.67 12.72 region region region region region region region*survey (categorical) 866.34 22 2001 survey (linear)+region 2001 2001 2001 survey (categorical)*region survey (linear) survey (categorical) survey (linear) survey (categorical) survey (linear) survey (categorical) survey (categorical)*region region survey (linear)+region region 870.02 10.76 0.00 17 region 884.86 25.60 0.00 24 region region region region region region region 894.60 900.32 904.02 904.96 907.73 908.14 910.13 35.34 41.06 44.76 45.70 48.47 48.88 50.87 0.00 0.00 0.00 0.00 0.00 0.00 0.00 11 14 7 10 9 12 29 region region age*survey (categorical) region 912.00 913.88 52.74 54.62 0.00 0.00 8 22 2001 2001 age survey (linear)+age region region age*survey (categorical) region 914.07 920.44 54.81 61.18 0.00 0.00 9 23 2001 survey (categorical) age*survey (categorical) region 928.89 69.63 0.00 24 2001 age*survey (categorical) age*survey (categorical) region 932.82 73.56 0.00 34 2001 2001 2001 2001 survey (categorical) survey (categorical) survey (linear) age age age age age*survey (categorical) age survey (categorical) age region 932.98 938.12 941.31 955.27 73.72 78.86 82.05 96.01 0.00 0.00 0.00 0.00 13 16 10 22 2001 2001 2001 2001 region age age*region survey (categorical) age age age constant region region region region 957.12 959.20 965.47 997.76 97.86 99.94 106.21 138.50 0.00 0.00 0.00 0.00 9 10 13 10 Year Probability of entering study area (pent ) survey (linear) survey (categorical) survey (linear) survey (categorical) survey (categorical) survey (categorical) Probability of staying in study area (phi ) constant constant tsa survey (categorical) survey (categorical) constant Probability of detection (p ) region region . region region region AICc Delta AICc 0.00 5.24 5.99 8.38 10.50 314.71 AICc Weights 0.87 0.06 0.04 0.01 0.00 0.00 Num. Par 7 10 11 13 14 10 Probability of entering study area (pent ) survey (linear) survey (categorical) survey (linear) survey (categorical) survey (categorical) survey (categorical) region Probability of staying in study area (phi ) age constant survey (categorical) age age survey (categorical) age Probability of detection (p ) age region region region age region region Delta AICc 0.00 0.45 2.79 4.79 6.49 9.36 20.45 AICc Weights 0.46 0.37 0.11 0.04 0.02 0.00 0.00 Num. Par 10 10 11 12 13 14 9 2001 2001 2001 2001 2001 2001 2001 2002 2002 2002 2002 2002 2002 Year 2003 2003 2003 2003 2003 2003 2003 survey (categorical)*region survey (categorical)*region survey (categorical) survey (categorical) constant constant age age age*survey (categorical) 683.06 688.29 689.04 691.44 693.55 997.76 AICc 410.37 410.82 413.15 415.16 416.86 419.73 430.82 2003 survey (categorical) age 21.68 0.00 22 2003 survey (linear)+region age*survey (categorical) region 437.92 27.55 0.00 22 2003 survey (categorical) age*survey (categorical) region 438.43 28.06 0.00 24 2003 2003 age*region survey (linear)+age age region age*survey (categorical) region 439.50 440.20 29.13 29.83 0.00 0.00 13 23 2003 survey region (categorical)*region age*survey (categorical) region 442.06 31.69 0.00 24 450.72 40.35 0.00 29 2003 survey (categorical)*region survey (categorical)*region age*survey (categorical) age*survey (categorical) region 463.48 53.11 0.00 34 2003 age age*survey (categorical) region 758.94 348.58 0.00 22 2003 survey (categorical) constant 997.76 587.39 0.00 10 Year 2004 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc study area (phi ) (p ) age region*survey (categorical) 662.07 Delta AICc 0.00 AICc Num. Weights Par 0.88 19 2004 survey (categorical) age region*survey (categorical) 666.59 4.52 0.09 22 2004 2004 2004 2004 survey (categorical) survey (categorical) survey (categorical) survey (categorical)*region survey (linear) survey (linear) survey (linear) survey (linear)+region constant age survey (categorical) survey (categorical)*region constant age age survey (categorical)*region age survey (categorical) age age*survey (categorical) region region region region 671.16 671.53 673.90 675.10 9.08 9.46 11.83 13.02 0.01 0.01 0.00 0.00 10 12 14 24 region region age region 675.16 675.38 675.96 676.65 13.09 13.30 13.89 14.57 0.00 0.00 0.00 0.00 7 9 10 17 age region survey (categorical) region 677.50 678.16 683.09 685.16 15.43 16.08 21.02 23.08 0.00 0.00 0.00 0.00 13 11 16 29 age*survey (categorical) region 690.40 28.33 0.00 24 2003 2004 2004 2004 2004 2004 2004 2004 2004 region*survey (categorical) 432.05 region 2004 survey (categorical) survey (linear) survey (categorical) survey (categorical)*region survey (categorical) 2004 survey (linear)+region age*survey (categorical) region 697.06 34.99 0.00 22 2004 2004 region survey (linear)+age region region age*survey (categorical) region 697.66 698.95 35.59 36.88 0.00 0.00 8 23 2004 2004 2004 2004 age constant constant region region survey (categorical) constant age 699.51 703.07 705.17 707.98 37.44 41.00 43.10 45.91 0.00 0.00 0.00 0.00 9 10 6 9 region region region region 2004 2004 2004 age age*region age age region age region age*survey (categorical) region 710.42 715.80 724.07 48.35 53.73 61.99 0.00 0.00 0.00 Year 2005 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc study area (phi ) (p ) age region*survey (categorical) 891.04 Delta AICc 0.00 AICc Num. Weights Par 1.00 19 2005 2005 2005 2005 2005 2005 survey (linear) survey (linear) survey (categorical) survey (categorical) survey (linear) survey (categorical) age constant age constant age age region region region region age region*survey (categorical) 945.97 947.00 953.56 953.82 953.85 955.64 54.93 55.96 62.52 62.78 62.81 64.60 0.00 0.00 0.00 0.00 0.00 0.00 9 7 12 10 10 22 2005 2005 2005 2005 2005 survey (categorical) survey (categorical) survey (categorical) survey (linear) survey (linear)+region region age survey (categorical) region region 956.54 957.38 957.71 961.07 972.06 65.50 66.33 66.67 70.03 81.02 0.00 0.00 0.00 0.00 0.00 14 13 16 11 17 2005 survey (linear)+region survey (categorical) age age survey (categorical) survey (categorical)*region age*survey (categorical) region 981.06 90.02 0.00 22 2005 survey (categorical) age*survey (categorical) region 981.20 90.16 0.00 24 2005 survey (linear)+age age*survey (categorical) region 983.15 92.11 0.00 23 2005 survey (categorical)*region survey (categorical)*region region region age age constant constant age*region age survey region (categorical)*region age*survey (categorical) region 987.14 96.10 0.00 24 990.01 98.97 0.00 29 age region age region constant survey (categorical) age age*survey (categorical) 1001.29 1001.51 1003.37 1003.58 1004.03 1008.83 1009.66 1046.90 110.25 110.47 112.33 112.54 112.99 117.79 118.62 155.86 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 9 8 10 9 6 10 13 22 2006 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc study area (phi ) (p ) age region*survey (categorical) 676.91 Delta AICc 0.00 AICc Num. Weights Par 1.00 19 2006 2006 2006 2006 2006 survey (categorical) survey (linear) survey (categorical) survey (categorical) survey (linear) constant constant age age age 57.74 59.51 60.86 61.61 61.67 0.00 0.00 0.00 0.00 0.00 2005 2005 2005 2005 2005 2005 2005 2005 2005 Year region region region region region region region region region region region age age 734.65 736.43 737.78 738.52 738.58 10 13 22 10 7 12 13 10 2006 2006 survey (linear) survey (categorical) age age region 739.80 region*survey (categorical) 741.70 62.89 64.78 0.00 0.00 9 22 2006 2006 2006 2006 survey (categorical) survey (categorical) survey (linear) survey (linear)+region region survey (categorical) region region 742.62 745.73 751.03 765.24 65.70 68.82 74.12 88.33 0.00 0.00 0.00 0.00 14 16 11 17 2006 survey (categorical) survey (categorical) age survey (categorical) survey (categorical)*region age*survey (categorical) region 767.67 90.76 0.00 24 2006 2006 constant survey (categorical)*region age age*survey (categorical) region region 769.72 773.09 92.80 96.18 0.00 0.00 6 24 2006 2006 constant survey (categorical)*region region survey (linear)+region region region 773.49 773.49 96.58 96.58 0.00 0.00 9 22 2006 2006 region survey (linear)+age region region age*survey (categorical) region 773.74 774.02 96.83 97.10 0.00 0.00 8 23 2006 2006 2006 2006 2006 age age constant age*region survey (categorical)*region age age region survey (categorical) age age*survey (categorical) region region region region region 775.60 775.84 776.08 781.98 784.64 98.68 98.92 99.16 105.06 107.72 0.00 0.00 0.00 0.00 0.00 10 9 10 13 29 age*survey (categorical) region 801.00 124.08 0.00 22 2007 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc study area (phi ) (p ) age region*survey (categorical) 669.99 Delta AICc 0.00 AICc Num. Weights Par 0.95 19 2007 survey (categorical) age region*survey (categorical) 675.74 5.75 0.05 22 2007 2007 2007 2007 survey (linear) survey (linear) survey (categorical) survey (linear)+region region region region region 693.68 694.45 700.06 700.48 23.69 24.46 30.07 30.49 0.00 0.00 0.00 0.00 7 9 10 17 2007 survey (categorical)*region survey (categorical) survey (linear) survey (categorical) survey (linear) survey (categorical) survey (categorical) survey (linear)+region constant age constant survey (categorical)*region survey (categorical)*region age age survey (categorical) survey (categorical) age age age*survey (categorical) region 700.90 30.91 0.00 24 region age region region survey (categorical) age region 701.58 707.86 709.65 713.83 716.52 720.81 723.35 31.58 37.87 39.66 43.84 46.53 50.82 53.36 0.00 0.00 0.00 0.00 0.00 0.00 0.00 12 10 14 11 16 13 22 738.83 68.84 0.00 29 2006 Year 2007 2007 2007 2007 2007 2007 2007 2007 survey (categorical)*region age*survey (categorical) region 2007 survey (categorical) age*survey (categorical) region 741.66 71.67 0.00 24 2007 2007 region survey (linear)+age region region age*survey (categorical) region 746.06 746.39 76.07 76.40 0.00 0.00 8 23 2007 2007 2007 2007 2007 2007 2007 age region constant age constant age*region age region age constant age survey (categorical) age age*survey (categorical) region region region region region region region 748.15 789.44 790.05 791.54 796.95 797.90 824.47 78.16 119.45 120.06 121.55 126.96 127.91 154.47 0.00 0.00 0.00 0.00 0.00 0.00 0.00 9 9 6 10 10 13 22 Year Probability of staying in study area (phi ) constant constant age age Probability of detection (p ) region region region region*survey (categorical) AICc 2008 2008 2008 2008 Probability of entering study area (pent ) survey (linear) survey (categorical) survey (linear) survey (linear) 893.03 895.14 896.50 896.77 Delta AICc 0.00 2.11 3.48 3.75 AICc Weights 0.52 0.18 0.09 0.08 Num. Par 7 10 9 19 2008 2008 2008 2008 survey (categorical) survey (categorical) survey (linear) survey (categorical) survey (categorical) age age age region region age region*survey (categorical) 897.91 898.43 899.30 899.97 4.88 5.40 6.27 6.94 0.05 0.03 0.02 0.02 14 12 10 22 2008 2008 2008 2008 survey (categorical) survey (linear) survey (categorical) survey (linear)+region age region survey (categorical) region 901.45 901.96 904.40 918.30 8.42 8.93 11.38 25.28 0.01 0.01 0.00 0.00 13 11 16 17 2008 survey (categorical) age survey (categorical) age survey (categorical)*region age*survey (categorical) region 921.06 28.03 0.00 24 2008 survey (linear)+region age*survey (categorical) region 921.36 28.33 0.00 22 2008 survey (categorical)*region survey (linear)+age survey region (categorical)*region age*survey (categorical) region 921.96 28.94 0.00 24 925.43 32.40 0.00 23 survey (categorical)*region region age constant constant region age age*region age age*survey (categorical) region 931.32 38.29 0.00 29 region region constant survey (categorical) age age age age*survey (categorical) 993.04 995.12 997.02 997.20 1000.53 1002.61 1008.90 1019.98 100.02 102.09 104.00 104.18 107.50 109.58 115.88 126.95 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 8 9 6 10 9 10 13 22 2008 2008 2008 2008 2008 2008 2008 2008 2008 2008 region region region region region region region region Year 2009 Probability of entering study area (pent ) survey (linear)+age Probability of staying in Probability of detection study area (phi ) (p ) age*survey (categorical) region 2009 2009 survey (linear) survey (linear)+region 2009 2009 2009 2009 survey (linear) survey (categorical) survey (linear) survey (linear) age survey (categorical)*region constant age survey (categorical) age 2009 2009 survey (categorical) survey (linear)+region constant region age*survey (categorical) region 2009 2009 survey (categorical) survey (categorical) survey (categorical) age 2009 survey (categorical)*region survey (linear) survey (categorical) survey region (categorical)*region age age age*survey (categorical) region 963.84 Delta AICc 0.00 AICc Num. Weights Par 0.61 16 region region 965.37 968.71 1.53 4.88 0.28 0.05 9 17 region region region region*survey (categorical) 970.00 970.80 973.14 974.64 6.16 6.96 9.30 10.80 0.03 0.02 0.01 0.00 7 12 11 19 975.38 978.07 11.54 14.23 0.00 0.00 10 22 region 978.84 region*survey (categorical) 979.10 15.00 15.26 0.00 0.00 14 22 980.11 16.27 0.00 24 980.91 982.30 17.07 18.46 0.00 0.00 8 24 2009 2009 2009 survey (categorical)*region survey (categorical) survey (categorical) age*survey (categorical) age*survey (categorical) region 989.62 25.78 0.00 29 age age age survey (categorical) age*survey (categorical) region 990.68 995.14 998.91 26.85 31.30 35.07 0.00 0.00 0.00 13 16 34 2009 2009 2009 2009 2009 2009 2009 2009 region age region age constant constant age*region age region region age age constant survey (categorical) age age*survey (categorical) 1031.50 1033.56 1043.48 1045.55 1047.25 1047.79 1051.82 1053.81 67.66 69.72 79.64 81.72 83.42 83.95 87.98 89.98 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 8 9 9 10 6 10 13 22 Year 2010 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc study area (phi ) (p ) age region*survey (categorical) 984.96 Delta AICc 0.00 AICc Num. Weights Par 0.42 19 2010 2010 survey (linear) survey (categorical) age age region 986.36 region*survey (categorical) 987.10 1.40 2.14 0.21 0.14 9 22 2010 2010 2010 2010 2010 survey (linear) survey (categorical) survey (linear) survey (categorical) survey (linear) age age constant constant survey (categorical) age region region region region 2.53 3.22 7.42 7.83 13.77 0.12 0.08 0.01 0.01 0.00 10 12 7 10 11 2009 2009 2009 region region region region region region region region AICc 987.49 988.18 992.38 992.79 998.74 2010 2010 2010 survey (categorical) survey (categorical) survey (linear)+region 2010 survey (categorical) age survey (categorical) survey (categorical)*region age*survey (categorical) survey (categorical) region region 1000.14 15.18 1001.38 16.42 1009.54 24.58 0.00 0.00 0.00 16 14 17 region 1014.48 29.52 0.00 24 2010 survey (linear)+region age*survey (categorical) region 1018.61 33.65 0.00 22 2010 survey (categorical)*region survey (linear)+age survey region (categorical)*region age*survey (categorical) region 1020.28 35.31 0.00 24 1021.12 36.16 0.00 23 age*survey (categorical) region 1038.01 53.05 0.00 29 2010 2010 2010 2010 2010 2010 survey (categorical)*region region age constant age*region constant age age age constant age survey (categorical) age*survey (categorical) region region region region region region 1061.53 1063.60 1069.58 1069.87 1077.17 1085.01 0.00 0.00 0.00 0.00 0.00 0.00 9 10 6 13 10 22 2010 survey (categorical) age region*survey (categorical) 1366.07 381.11 0.00 22 Year 2011 Probability of entering study area (pent ) survey (linear) Probability of staying in Probability of detection AICc Delta study area (phi ) (p ) AICc age region*survey (categorical) 1256.57 0.00 AICc Num. Weights Par 0.77 19 2011 survey (categorical) age region*survey (categorical) 1259.08 2.51 0.22 22 2011 2011 2011 2011 2011 survey (linear) survey (categorical) survey (categorical) survey (linear) survey (linear)+region region region region region region 1266.13 1266.18 1270.96 1271.06 1273.56 9.56 9.61 14.40 14.50 16.99 0.01 0.01 0.00 0.00 0.00 9 12 10 7 17 2011 2011 2011 2011 2011 survey (linear) survey (categorical) survey (linear) survey (categorical) survey (linear)+region age age constant constant survey (categorical)*region age age survey (categorical) survey (categorical) age*survey (categorical) age age region region region 1274.58 1276.34 1276.58 1279.79 1281.87 18.01 19.77 20.01 23.23 25.30 0.00 0.00 0.00 0.00 0.00 10 13 11 14 22 2011 survey region (categorical)*region age*survey (categorical) region 1283.40 26.83 0.00 24 2011 survey (categorical)*region survey (categorical) 1286.47 29.91 0.00 24 2011 survey (linear)+age age*survey (categorical) region 1288.16 31.59 0.00 23 2011 survey (categorical)*region survey (categorical) age*survey (categorical) region 1293.24 36.67 0.00 29 age 1295.38 38.81 0.00 16 2010 2010 2011 survey (categorical) 76.57 78.64 84.62 84.91 92.21 100.05 2011 2011 2011 2011 2011 2011 2011 2011 region age region age constant age*region constant age region region age age constant age survey (categorical) age*survey (categorical) region region region region region region region region Year 2012 2012 Probability of entering study area (pent ) survey (categorical) survey (linear) Probability of staying in study area (phi ) age age Probability of detection AICc (p ) region 2157.44 region*survey (categorical) 2161.66 2012 2012 2012 survey (linear) survey (categorical) survey (categorical) age age age 2012 2012 2012 2012 survey (linear) survey (categorical) survey (linear) survey (linear)+region 2012 survey (linear)+region 2012 2012 2012 survey (categorical) survey (linear) survey (categorical) 2012 2012 2012 survey (categorical) survey (categorical)*region survey (categorical)*region survey (linear)+age 2012 2012 2012 2012 2012 2012 2012 2012 Year 2012 2013 2013 1434.90 1436.95 1456.19 1458.25 1462.56 1464.44 1465.42 1471.58 178.33 180.38 199.63 201.68 206.00 207.87 208.86 215.02 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 8 9 9 10 6 13 10 22 Delta AICc 0.00 4.22 AICc Weights 0.84 0.10 Num. Par 12 19 age 2163.56 6.12 age 2166.23 8.79 region*survey (categorical) 2167.18 9.74 0.04 0.01 0.01 10 13 22 age constant constant age*survey (categorical) region region region region 2168.51 2171.49 2173.04 2174.90 11.07 14.04 15.60 17.46 0.00 0.00 0.00 0.00 9 10 7 22 survey (categorical)*region survey (categorical) survey (categorical) age*survey (categorical) region 2176.72 19.27 0.00 17 region region region 2178.75 21.31 2179.65 22.20 2180.72 23.27 0.00 0.00 0.00 14 11 24 age survey (categorical) survey region (categorical)*region age*survey (categorical) region 2184.72 27.28 2194.06 36.62 0.00 0.00 16 24 2198.36 40.92 0.00 29 age*survey (categorical) region 2204.52 47.07 0.00 23 region age region age constant constant age*region age age age region region survey (categorical) constant age age*survey (categorical) region region region region region region region region 2314.50 2316.53 2317.39 2319.42 2322.18 2322.46 2322.66 2331.06 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 9 10 8 9 10 6 13 22 Probability of entering study area (pent ) survey (categorical) survey (linear) Probability of staying in study area (phi ) age age Probability of detection (p ) age age AICc AICc Weights 1.00 0.00 Num. Par 13 10 157.05 159.09 159.94 161.97 164.73 165.02 165.21 173.61 Delta AICc 2720.51 0.00 2737.65 17.14 2013 2013 survey (categorical) survey (linear) age age region 2738.61 18.10 region*survey (categorical) 2746.57 26.05 0.00 0.00 12 19 2013 survey (categorical) age region*survey (categorical) 2754.53 34.01 0.00 22 2013 2013 2013 survey (categorical) survey (categorical) survey (categorical) constant region age survey (categorical) age*survey (categorical) region 2756.65 36.14 2761.00 40.49 2767.01 46.49 0.00 0.00 0.00 10 16 24 2013 2013 2013 2013 survey (categorical) survey (linear) survey (linear) survey (categorical)*region survey (linear)+region survey (categorical) age constant age*survey (categorical) 2767.65 2769.36 2769.59 2777.82 47.14 48.85 49.08 57.31 0.00 0.00 0.00 0.00 14 9 7 29 age*survey (categorical) region 2779.38 58.87 0.00 22 2782.55 62.03 2783.08 62.56 0.00 0.00 11 24 region 2792.59 72.08 0.00 17 2013 survey (linear)+age survey (categorical) survey (categorical)*region survey (categorical)*region age*survey (categorical) region region 2013 survey (linear) survey (categorical)*region survey (linear)+region region 2794.53 74.01 0.00 23 2013 2013 2013 2013 region age age*region age age age age age*survey (categorical) region region region region 2987.39 2989.42 2995.51 3002.99 266.88 268.90 275.00 282.48 0.00 0.00 0.00 0.00 9 10 13 22 2013 2013 2013 2013 region constant constant age region survey (categorical) constant region region region region region 3006.71 3007.22 3008.58 3008.73 286.20 286.70 288.06 288.22 0.00 0.00 0.00 0.00 8 10 6 9 2013 2013 2013 region region region region