Rivista Italiana di Paleontologia e Stratigrafia
volume 114
no. 2
6 pls.
pp. 233-262
July 2008
OSTRACODS ACROSS THE PERMIAN-TRIASSIC BOUNDARY IN WESTERN TETHYS:
THE BULLA PARASTRATOTYPE (SOUTHERN ALPS, ITALY)
SYLVIE CRASQUIN 1, M. CRISTINA PERRI 2, ALDA NICORA 3 & PATRICK DE WEVER 4
Received: February 27, 2008; accepted: May 26, 2008
Key words: Ostracods, Permian-Triassic boundary, Bulla
sec-
tion, Italy.
Abstract. Investigation of the ostracod fauna of the parastratotype of the Permian-Triassic boundary at Bulla in the Southern Alps
produced 62 species belonging to 31 genera. They are all discussed
and figured. This paper presents results of the first description of
ostracods from this important site. One genus, Bairdiacratia n. gen.,
and 13 species are new: Glyptopleurina pasinii n. sp., Knoxiella ventrospinosa n. sp., Knightina bullaensis n. sp., Bairdia ortiseiensis n. sp.,
B. cheni n. sp., B. (Rectobairdia) kershawi n. sp., Bairdiacratia qinglai
n. gen. n. sp., B. tergilata n. gen. n. sp., Microcheilinella lata n. sp.,
Parabythocythere chongpani n. sp., Cavellina bellerophonella n. sp.,
C. alpina n. sp. and C. triassica n. sp. The palaeocecological analysis
of each unit is produced. The unconformity-paraconformity U1 is
clearly reflected in the ostracod assemblages and is marked by a drop
in diversity and abundance of specimens. It was followed by a change
in the ostracod faunal composition. The Bulla Member displays maximum ostracod diversity and abundance linked with the trangressive
trend reported for this period. The unconformity-paraconformity U2,
at the boundary between the Bellerophon and Werfen formations
(Bulla and Lower Tesero members) is the main extinction level for
ostracods. The Lower Tesero, Lower Mazzin and Upper Tesero members have very poor faunas. The lower part of the Upper Mazzin Mb.
is characterized by an uneven burst of diversity before the great
period of taxonomic paucity observed during the late Griesbachian
all over the world.
Riassunto. Per mezzo dell'analisi delle faune ad ostracodi del
parastratotipo del limite Permiano-Triassico a Bulla (BZ) nelle Alpi
Meridionali, sono state identificate 62 specie appartenenti a 31 generi.
La loro discussione e documentazione iconografica eÁ argomento di questo lavoro. Vengono descritti un nuovo genere, Bairdiacratia n. gen., e 13
nuove specie: Glyptopleurina pasinii n. sp., Knoxiella ventrospinosa n.
sp., Knightina bullaensis n. sp., Bairdia ortiseiensis n. sp., B. cheni n. sp.,
B. (Rectobairdia) kershawi n. sp., Bairdiacratia qinglai n. gen. n. sp., B.
1
2
3
4
tergilata n. gen. n. sp., Microcheilinella lata n. sp., Parabythocythere
chongpani n. sp., Cavellina bellerophonella n. sp., C. alpina n. sp. and
C. triassica n. sp. Per ciascuna unitaÁ litostratigrafica viene riportata l'a-
nalisi paleoecologica. Le associazioni ad ostracodi riflettono in modo
evidente la superficie di discontinuitaÁ U1 (unconformity-paraconformity 1) che separa l'UnitaÁ ad ostracodi (Ostracod and peritidal dolomite
unit) e il Membro di Bulla (Bulla Mbr) della Formazione a Bellerophon,
con un calo drastico in diversitaÁ e abbondanza. La maggiore diversitaÁ e
abbondanza delle faune ad ostracodi eÁ associata alla fase trasgressiva del
Membro di Bulla. Il principale evento di estinzione ad ostracodi coincide con la superficie di discontinuitaÁ U2 (unconformity-paraconformity
2) posta in corrispondenza del limite formazionale fra la Formazione a
Bellerophon e la Formazione di Werfen, fra i membri di Bulla e del
Tesero Inferiore. I membri del Tesero Inferiore, Mazzin Inferiore e
Tesero Superiore presentano una fauna ad ostracodi veramente povera.
La parte inferiore del Membro del Mazzin Superiore eÁ caratterizzata da
un irregolare incremento di diversitaÁ che precede il periodo di scarsezza
tassonomica osservato a livello mondiale durante il tardo Griesbachiano.
Introduction
The Bulla section in the Southern Alps is the parastratotype section for the Permian-Triassic (P-T)
boundary. It was located at that time in western Tethys
whereas the P-T stratotype at Meishan in China was
located near the eastern border of Tethys. The Bulla
section, well-known from many investigations, has been
described in detail by Perri (1991), Farabegoli & Perri
(1998), Perri & Farabegoli (2003) and Farabegoli et al.
(2007 and references therein). Apart from a preliminary
note (Pasini 1981) on ostracods from the upper part of
the Bellerophon Formation at Digonera and Rio Bar-
CNRS, UMR 5143, Universite Pierre et Marie Curie, Laboratoire de MicropaleÂontologie, T.46-56, E.5, case 104, 75252 Paris cedex 05, France.
E-mail: sylvie.crasquin@upmc.fr
UniversitaÁ degli Studi di Bologna, Dip. di Scienze della Terra e Geologico-Ambientali, Via Zamboni 67, 40126 Bologna, Italy. E-mail:
mariacristina.perri@unibo.it
UniversitaÁ degli Studi di Milano, Dip. Scienze della Terra, Via Mangiagalli 34, 20133 Milano, Italy. E-mail: alda.nicora@unimi.it
MNHN, DeÂpartement Histoire de la Terre, GeÂologie, 43 Rue Buffon, 75005 Paris, France. E-mail: pdewever@mnhn.fr
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05BU2
05BU2A
*
*
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*
*
*
*
*
*
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*
Lower Tesero Mb
05BU7A 05BU7B
05BU9
05BU4A
05BU5
05BU6
Ű
*
*
*
*
*
*
*
*
*
*
*
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*
*
*
*
*
*
Ű
*
*
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*
*
*
*
*
*
*
*
*
*
Ű
Upper Mazzin Mb
L. Mazzin Mb
05BU3
05BU10 05BU11 05BU12 05BU13A 05BU14 05BU17 05BU18 05BU19 05BU19A 05BU21 05BU23 05BU24
Ű
Ű
05BUCS105BUSC2 05BU1A 05BU1B 05BU1C
*
*
*
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*
*
*
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*
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05BU1C
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"Ostracod Unit"
05BUCS105BUSC2 05BU1A 05BU1B
species
Ű
1 Cavellina alpina n.sp.
Ű
Ű
Ű
Ű
2 Cavellina bellerophonella n.sp.
Ű
Ű
*
*
3 Arqoviella sp.1
Ű
*
Ű
Ű
4 Knoxiella ventrospinosa n.sp.
Ű
5 Cavellina visnyoensis
Ű
Ű
6 Neoulrichia pulchra
Ű
Ű
7 Acratia zhongyingensis
Ű
Ű
8 Sargentina postacuta
9 Callicythere mazurensis
10 Fabalicypris? sp.A
11 Bairdiacypris sp.1
12 Sargentina n.sp.A
13 Bairdia sp. 7
14 Bairdia sp. 2
15 Bairdia sp. 1
16 Bairdia cf subcontracta
17 Paramacrocypris schallreuteri
18 Bairdia cheni n.sp.
19 Bairdiacratia tergilata n.gen. n.sp.
20 Parabythocythere chongpani
21 Bairdiacratia qinglaii n.gen. n.sp.
22 Sulcella suprapermiana
23 Bairdia sp. 6
24 Kirkbya cf buekkensis
25 Glyptopleurina pasinii n.sp.
26 Acratia sp.2
27 Bairdia sp. 3
28 Bairdia sp. 4
29 Cryptobairdia cf postilonga
30 Kirkbyidae sp.1
31 Kirkbyidae sp.2
32 Knightina bullaensis n.sp.
33 Paraparchites sp.1
34 Bairdiacratia n.gen. n.sp.2
35 Basslerella braccinii nov. nom.
36 Bairdia sp. 5
37 Bairdia galei
38 Bairdia (R.) kershawi n.sp.
39 Kirkbya sp.1
40 Bairdia permagna
41 cf Shleesha pinguis
42 Bairdia ortiseiensis n.sp.
43 Petasobairdia nantongensis
44 Acratia sp.1
45 Bairdiacypris wangi
46 Microcheilinella peraxilis
47 Microcheilinella lata n.sp.
48 Parahollinella visnyoensis
49 Bairdiacypris longirobusta
50 Bairdiacypris? caeca
51 Pseudoacanthoscapha striatula
52 Geffenina? sp.1
53 Fabalicypris cf vinsyoensis
54 Microcheilinella sp.2
55 Macrocypris sp.1
56 Ostracoda indet
57 Liuzhinia antalyaensis
58 Langdaia laolongdongensis
59 Kirkbyidae sp.3
60 Langdaia suboblonga
61 Cavellina triassica n.sp.
62 Kerocythere? sp. 1
63 Paracypris sp.1
64 Paracypris gaetanii
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05BU10 05BU11 05BU12 05BU13A 05BU14 05BU17
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05BU19 05BU19A 05BU21 05BU23 05BU24
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Number of species (triangles)
0
5
10
15
20
25
05BU28
05BU27
05BU26
Upper MazzinMb
05BU25
05BU24
05BU23
05BU21
05BU20
05BU19A
nb specimens
nb species
05BU22
05BU19
05BU18
05BU17
Upper Tesero Mb
05BU16
05BU14ter
05BU14bis
Lower Mazzin Mb
05BU14
05BU13A
Triassic
05BU12C
05BU12B
samples
Permian
05BU12
Lower Tesero Mb
05BU11
05BU10
05BU9
05BU8
05BU7B
U2
05BU7A
05BU6
05BU5
05BU4A
05BU3
Bulla Mb
05BU2A
05BU2
05BU1C
U1
05BU1B
950
05BU1A
05BUSC3
"ostracod unit"
05BUSC2
05BUCS1
0
50
100
150
200
250
300
350
400
450
500
Number of specimens (squares)
.$ *$!$ $!" 00" $% #0+" @F* A $%!$$%-
# " 0"% #$%" "%,$ %#"% /") ! #$%" /" &
! "#'+-" $% ++ . /" *(++ ' 4"0 $
% 0 + " %! (!
0 ++ ! 0"$" 0""% $% *(++ ' "
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+$) . "2 3$%- . . $+ . # /" G ! (%$9 $
% (00 "! /"" /" '(%!% %! !$,"" !
.(% 0$(++) $% #0+" @F* F 1 %! A "
! 2$/ /" "(% . % #+ #$%" 0"% %!$&
$ % %%""! 2$/ %-"$," 0/" /" 0"3 .
!$,"$) %! '(%!%" ("! $% #0+" @F* 1
!$ ! 0 $% /" ! ##(%$) &
("! '"2""% #0+" @F* A %! @F* A* 2$/
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. #$) . ='"- +$ " + 5@@A6 $%"0""!
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Cavellinidae
Beyrichiidae
Sansabellidae
Kirkbyidae
Paraparchitidae
Cytheracoidea
10%
20%
30%
40%
50%
Percentage of species
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%
.
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(0".#$+$" $% /" *(++ "$
% 5 0""%-"
60%
70%
80%
90%
100%
& 2 # +%
Ostracod indet
# #3 0
Kloedenellidae
0%
=$- F
3! , )
Bairdioidea
. 0"$"6
>F
Samples
05BU24
05BU23
05BU22
05BU21
05BU20
05BU19A
05BU19
05BU18
05BU17
05BU16
05BU14ter
05BU14bis
05BU14
05BU13A
05BU12C
05BU12B
05BU12
05BU11
05BU10
05BU9
05BU8
05BU7B
05BU7A
05BU6
05BU5
05BU4A
05BU3
05BU2A
05BU2
05BU1C
05BU1B
05BU1A
05BUSC3
05BUSC2
05BUCS1
1@
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