Form-genus <i>Desmiophyllum</i> Lesquereux, 1878 Layers A to E (Fig. 5 A) SPECIMENS. ... more Form-genus <i>Desmiophyllum</i> Lesquereux, 1878 Layers A to E (Fig. 5 A) SPECIMENS. — MPZ 2010/1040 to MPZ 2010/1052. DESCRIPTION Fragments of elongate strap-like leaves up to 12 cm long and 2 cm wide.Margins straight and reinforced.Base and apex not preserved.Venation consists of numerous gross veins, <i>c.</i> 25 veins per cm, running parallel to leaf margins. Another four thin veins extended between two gross veins making a total of <i>c.</i> 125 veins per cm. REMARKS Similar leaves were also found in the Aptian-Albian of western Europe (Coiffard <i>et al.</i> 2007), Canada (Bell 1956) and in Albian-Cenomanian deposits of Alaska (Spicer & Herman 2001) and Russia (Spicer <i>et al.</i> 2002).
FIG. 6. — Fertile and sterile structures of aquatic plants: A, floral receptacles of presumable N... more FIG. 6. — Fertile and sterile structures of aquatic plants: A, floral receptacles of presumable Nelumbonaceae floral receptacle showing internal moulds of achene cavities; B, Aquatifolia cf. fluitans, aquatic plant with foliar lamina, petiole and fusiform float organ at base; C, almost complete lamina of Aquatifolia cf. fluitans showing venation. Scale bars: 1 cm.
FIG. 10. — Photographs of some selected sporomorphs from the Estercuel section (Teruel province, ... more FIG. 10. — Photographs of some selected sporomorphs from the Estercuel section (Teruel province, northeastern Spain): A, Classopollis major; B, Perinopollenites halonatus, mid focus; C, Transitoripollis sp. cf. T. similis, distal face; D, Retimonocolpites textus, distal face; E, Clavatipollenites sp., mid focus; F, Afropollis jardinus, mid focus; G, H, Pennipollis sp. cf. P. reticulatus in distal (G) and proximal (H) faces; I, Rousea sp., polar view; J, Phimopollenites augathellaensis, equatorial view; K, Senectotetradites varireticulatus, equatorial view; L, M, Striatopollis sp., equatorial view; L, high focus; M, same specimen, low focus; N, O, Tricolporoidites sp.; N, polar view; O, equatorial view. Scale bar: 10 µm. Full taxa names are given in Table 1.
FIG. 4. — Distribution of Afropollis jardinus Doyle, Jardiné & Doerenkamp, 1982 (z) and Cycloneph... more FIG. 4. — Distribution of Afropollis jardinus Doyle, Jardiné & Doerenkamp, 1982 (z) and Cyclonephelium chabaca Below, 1981 (*). The palaeogeographic map was created using the ODSN Plate Tectonic Reconstruction Service (Hay et al. 1999) back to 100 My (late Albian) in an orthographic projection: Afropollis jardinus, Laurasia: 1, Mons Basin (Yans et al. 2007), Belgium; 2, Bohemia (Pacltová & Lashin 1998), Czech Republic; 3, Marches-Ombrie Basin (Fiet 1999; Fiet & Pons 1998), Italy; 4, Charentes, Île Madame (Dejax & Masure 2005; SNEA[P]; Doyle et al. 1982), France; 5, Ecommoy (Azéma et al. 1972), France; 6, Neau (Azéma et al. 1972), France; 7, La Bironnière (Azéma et al. 1972), France; 8, Anjou (Yans et al. 2007), France; 9, between Lisbon and Foz da Folcao and on both sides of the Serra de Sintra (Hasenboehler 1981), Portugal; 10, Iberian Chain (this study), Spain; North America: 11, Lake Texoma, Oklahoma (Wingate 1980), USA; Gondwana, South America: 12, Sergipe and Alagoas Basins (Mü...
FIG. 8. — A, Antulsporites varigranulatus in proximal face (VA2B4Q524); B, Stereisporites antiqua... more FIG. 8. — A, Antulsporites varigranulatus in proximal face (VA2B4Q524); B, Stereisporites antiquasporites in proximal view (VA2A3H44); C, Aequitriradites spinulosus in distal view (FV2A1K19); D, Triporoletes laevigatus in proximal view (FV4 3_3R4); E, Aequitriradites sp. 2 in distal view (PS8a2_5F424); F, Triporoletes cenomanianus in proximal view (PS8bA3T573); G, H, Anapiculatiporites sp. in proximal (G) and distal (H) views (PS126P414); I, P, Camarozonosporites sp. 2 in proximal (I) and distal (P) views (FV3 3_3M352); J, M, Asbeckiasporites sp. in proximal (J) and distal (M) views (PS121X291); K, N, Camarozonosporites sp. 1 in proximal (K) and distal (N) views (PS124S56); L, O, Camarozonosporites insignis in proximal (L) and distal (O) views (PS10c4M523); Q, Asbeckiasporites sp. in distal view (PS10); R, S, Cicatricosisporites sp. cf. Anemia exilioides in proximal (R) and distal (S) views (FV4B1F58). Scale bar: A-E, G-P, R, S, 20 μm; F, Q, 10 μm.
FIG. 9. — A, B, Cicatricosisporites hallei in proximal (A) and distal (B) views (FV3_3N382); C, D... more FIG. 9. — A, B, Cicatricosisporites hallei in proximal (A) and distal (B) views (FV3_3N382); C, D, Cicatricosisporites hughesii in proximal (C) and distal (D) views (FV1B2G433); E, F, Cicatricosisporites imbricatus in proximal (E) and distal (F) views (PS121R37); G, H, Cicatricosisporites sp. cf. C. imbricatus in proximal (G) and distal (H) views (PS132O343); I, J, Cicatricosisporites minutaestriatus in proximal (I) and distal (J) views (PS8bB2O532); K, L, Cicatricosisporites myrtellii in proximal (K) and distal (L) views (PS121L254); M, R, Cicatricosisporites perforatus in proximal (M) and distal (R) views (PS7c3O402); N, O, Cicatricosisporites venustus in proximal (N) and distal (O) views (PS10b3bS504); P, Q, Cicatricosisporites pseudotripartitus in proximal (I) and distal (N) views (PS10A2S383); S, T, Cicatricosisporites subrotundus in proximal (S) and distal (T) views (FV3 3_3N384); U, Cicatricosisporites pseudotripartitus in proximal view (PS10a), SEM. Scale bars: A-T, 20 μm; U...
FIG. 3. — The palynostratigraphic distribution, for the presence of the individual spores and pol... more FIG. 3. — The palynostratigraphic distribution, for the presence of the individual spores and pollen grains identified within the positive 21 samples, based on the previous available literature.
ABSTRACT Exceptionally well-preserved specimens from the middle Albian of Spain corresponding to ... more ABSTRACT Exceptionally well-preserved specimens from the middle Albian of Spain corresponding to a nearly complete fertile frond and fragments of pinnae containing soral clusters of the tree fern Weichselia reticulata have provided new data about the structure and arrangement of pinnae and their associated fertile reproductive structure. This new material has been compared with the previous studies, and recostructions of this fern and the new data indicate the segregation of vegetative and fertile fronds within the main stipe of this fern.
ABSTRACT In this paper we present new macrofossils of semi-aquatic fern Family Marsileaceae consi... more ABSTRACT In this paper we present new macrofossils of semi-aquatic fern Family Marsileaceae consisting of leaves and leaflets from the upper Hauterivian- lower Barremian Pinilla de los Moros Formation, Salas de los Infantes village, Burgos province, north of Spain. The leaves show two elliptic to flabellate-shaped leaflets that are joined to a petiole. Leaflets present entire margins and dichotomous anastomosed vein pattern. Comparison between the studied leaves and other marsileaceous leaves indicates close similarities with Regnellites nagashimae species. These leaves were deposited in a related fluvial environment near to the place where semi-aquatic plants grew. The studied macrofossils constitute at present the earliest record of leaves of Family Marsileaceae in the Mesozoic of Eurasia at the moment.
ABSTRACT A well-preserved macroflora and rich palynological assemblages corresponding to the Cama... more ABSTRACT A well-preserved macroflora and rich palynological assemblages corresponding to the Camarillas Formation (early-middle Barremian) in the San Cristo´bal and Galve Mine sites from the Galve sub-basin in northeastern Spain are presented here. These remains represent the first fossil plant evidence from these deposits. Within plant macroremains, the cheirolepidiacean Pseudofrenelopsis aff. varians (Fontaine) Watson has been found. The palynological assemblage yielded well-preserved spores and pollen grains, mainly dominated by the genus Classopollis. Spores are also abundantly represented by schizaeacean spores (Cicatricosisporites and Plicatella). This spore assemblage supports an early-middle Barremian age for these localities. It is noteworthy that small basal angiosperm pollen grains of the genera Crassipollis and Retimonocolpites, together with other indeterminate grains, have been reported here. This flora constitutes the primary food producer for dinosaur at that time, and thus the information of the flora is important for the understanding of the ecological background for the dinosaur evolution during the Early Cretaceous.
Form-genus <i>Desmiophyllum</i> Lesquereux, 1878 Layers A to E (Fig. 5 A) SPECIMENS. ... more Form-genus <i>Desmiophyllum</i> Lesquereux, 1878 Layers A to E (Fig. 5 A) SPECIMENS. — MPZ 2010/1040 to MPZ 2010/1052. DESCRIPTION Fragments of elongate strap-like leaves up to 12 cm long and 2 cm wide.Margins straight and reinforced.Base and apex not preserved.Venation consists of numerous gross veins, <i>c.</i> 25 veins per cm, running parallel to leaf margins. Another four thin veins extended between two gross veins making a total of <i>c.</i> 125 veins per cm. REMARKS Similar leaves were also found in the Aptian-Albian of western Europe (Coiffard <i>et al.</i> 2007), Canada (Bell 1956) and in Albian-Cenomanian deposits of Alaska (Spicer & Herman 2001) and Russia (Spicer <i>et al.</i> 2002).
FIG. 6. — Fertile and sterile structures of aquatic plants: A, floral receptacles of presumable N... more FIG. 6. — Fertile and sterile structures of aquatic plants: A, floral receptacles of presumable Nelumbonaceae floral receptacle showing internal moulds of achene cavities; B, Aquatifolia cf. fluitans, aquatic plant with foliar lamina, petiole and fusiform float organ at base; C, almost complete lamina of Aquatifolia cf. fluitans showing venation. Scale bars: 1 cm.
FIG. 10. — Photographs of some selected sporomorphs from the Estercuel section (Teruel province, ... more FIG. 10. — Photographs of some selected sporomorphs from the Estercuel section (Teruel province, northeastern Spain): A, Classopollis major; B, Perinopollenites halonatus, mid focus; C, Transitoripollis sp. cf. T. similis, distal face; D, Retimonocolpites textus, distal face; E, Clavatipollenites sp., mid focus; F, Afropollis jardinus, mid focus; G, H, Pennipollis sp. cf. P. reticulatus in distal (G) and proximal (H) faces; I, Rousea sp., polar view; J, Phimopollenites augathellaensis, equatorial view; K, Senectotetradites varireticulatus, equatorial view; L, M, Striatopollis sp., equatorial view; L, high focus; M, same specimen, low focus; N, O, Tricolporoidites sp.; N, polar view; O, equatorial view. Scale bar: 10 µm. Full taxa names are given in Table 1.
FIG. 4. — Distribution of Afropollis jardinus Doyle, Jardiné & Doerenkamp, 1982 (z) and Cycloneph... more FIG. 4. — Distribution of Afropollis jardinus Doyle, Jardiné & Doerenkamp, 1982 (z) and Cyclonephelium chabaca Below, 1981 (*). The palaeogeographic map was created using the ODSN Plate Tectonic Reconstruction Service (Hay et al. 1999) back to 100 My (late Albian) in an orthographic projection: Afropollis jardinus, Laurasia: 1, Mons Basin (Yans et al. 2007), Belgium; 2, Bohemia (Pacltová & Lashin 1998), Czech Republic; 3, Marches-Ombrie Basin (Fiet 1999; Fiet & Pons 1998), Italy; 4, Charentes, Île Madame (Dejax & Masure 2005; SNEA[P]; Doyle et al. 1982), France; 5, Ecommoy (Azéma et al. 1972), France; 6, Neau (Azéma et al. 1972), France; 7, La Bironnière (Azéma et al. 1972), France; 8, Anjou (Yans et al. 2007), France; 9, between Lisbon and Foz da Folcao and on both sides of the Serra de Sintra (Hasenboehler 1981), Portugal; 10, Iberian Chain (this study), Spain; North America: 11, Lake Texoma, Oklahoma (Wingate 1980), USA; Gondwana, South America: 12, Sergipe and Alagoas Basins (Mü...
FIG. 8. — A, Antulsporites varigranulatus in proximal face (VA2B4Q524); B, Stereisporites antiqua... more FIG. 8. — A, Antulsporites varigranulatus in proximal face (VA2B4Q524); B, Stereisporites antiquasporites in proximal view (VA2A3H44); C, Aequitriradites spinulosus in distal view (FV2A1K19); D, Triporoletes laevigatus in proximal view (FV4 3_3R4); E, Aequitriradites sp. 2 in distal view (PS8a2_5F424); F, Triporoletes cenomanianus in proximal view (PS8bA3T573); G, H, Anapiculatiporites sp. in proximal (G) and distal (H) views (PS126P414); I, P, Camarozonosporites sp. 2 in proximal (I) and distal (P) views (FV3 3_3M352); J, M, Asbeckiasporites sp. in proximal (J) and distal (M) views (PS121X291); K, N, Camarozonosporites sp. 1 in proximal (K) and distal (N) views (PS124S56); L, O, Camarozonosporites insignis in proximal (L) and distal (O) views (PS10c4M523); Q, Asbeckiasporites sp. in distal view (PS10); R, S, Cicatricosisporites sp. cf. Anemia exilioides in proximal (R) and distal (S) views (FV4B1F58). Scale bar: A-E, G-P, R, S, 20 μm; F, Q, 10 μm.
FIG. 9. — A, B, Cicatricosisporites hallei in proximal (A) and distal (B) views (FV3_3N382); C, D... more FIG. 9. — A, B, Cicatricosisporites hallei in proximal (A) and distal (B) views (FV3_3N382); C, D, Cicatricosisporites hughesii in proximal (C) and distal (D) views (FV1B2G433); E, F, Cicatricosisporites imbricatus in proximal (E) and distal (F) views (PS121R37); G, H, Cicatricosisporites sp. cf. C. imbricatus in proximal (G) and distal (H) views (PS132O343); I, J, Cicatricosisporites minutaestriatus in proximal (I) and distal (J) views (PS8bB2O532); K, L, Cicatricosisporites myrtellii in proximal (K) and distal (L) views (PS121L254); M, R, Cicatricosisporites perforatus in proximal (M) and distal (R) views (PS7c3O402); N, O, Cicatricosisporites venustus in proximal (N) and distal (O) views (PS10b3bS504); P, Q, Cicatricosisporites pseudotripartitus in proximal (I) and distal (N) views (PS10A2S383); S, T, Cicatricosisporites subrotundus in proximal (S) and distal (T) views (FV3 3_3N384); U, Cicatricosisporites pseudotripartitus in proximal view (PS10a), SEM. Scale bars: A-T, 20 μm; U...
FIG. 3. — The palynostratigraphic distribution, for the presence of the individual spores and pol... more FIG. 3. — The palynostratigraphic distribution, for the presence of the individual spores and pollen grains identified within the positive 21 samples, based on the previous available literature.
ABSTRACT Exceptionally well-preserved specimens from the middle Albian of Spain corresponding to ... more ABSTRACT Exceptionally well-preserved specimens from the middle Albian of Spain corresponding to a nearly complete fertile frond and fragments of pinnae containing soral clusters of the tree fern Weichselia reticulata have provided new data about the structure and arrangement of pinnae and their associated fertile reproductive structure. This new material has been compared with the previous studies, and recostructions of this fern and the new data indicate the segregation of vegetative and fertile fronds within the main stipe of this fern.
ABSTRACT In this paper we present new macrofossils of semi-aquatic fern Family Marsileaceae consi... more ABSTRACT In this paper we present new macrofossils of semi-aquatic fern Family Marsileaceae consisting of leaves and leaflets from the upper Hauterivian- lower Barremian Pinilla de los Moros Formation, Salas de los Infantes village, Burgos province, north of Spain. The leaves show two elliptic to flabellate-shaped leaflets that are joined to a petiole. Leaflets present entire margins and dichotomous anastomosed vein pattern. Comparison between the studied leaves and other marsileaceous leaves indicates close similarities with Regnellites nagashimae species. These leaves were deposited in a related fluvial environment near to the place where semi-aquatic plants grew. The studied macrofossils constitute at present the earliest record of leaves of Family Marsileaceae in the Mesozoic of Eurasia at the moment.
ABSTRACT A well-preserved macroflora and rich palynological assemblages corresponding to the Cama... more ABSTRACT A well-preserved macroflora and rich palynological assemblages corresponding to the Camarillas Formation (early-middle Barremian) in the San Cristo´bal and Galve Mine sites from the Galve sub-basin in northeastern Spain are presented here. These remains represent the first fossil plant evidence from these deposits. Within plant macroremains, the cheirolepidiacean Pseudofrenelopsis aff. varians (Fontaine) Watson has been found. The palynological assemblage yielded well-preserved spores and pollen grains, mainly dominated by the genus Classopollis. Spores are also abundantly represented by schizaeacean spores (Cicatricosisporites and Plicatella). This spore assemblage supports an early-middle Barremian age for these localities. It is noteworthy that small basal angiosperm pollen grains of the genera Crassipollis and Retimonocolpites, together with other indeterminate grains, have been reported here. This flora constitutes the primary food producer for dinosaur at that time, and thus the information of the flora is important for the understanding of the ecological background for the dinosaur evolution during the Early Cretaceous.
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