... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
Global Environmental Change-human and Policy Dimensions, 1999
Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2... more Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2100, with IS92a greenhouse gas forcing, together with predicted patterns of N deposition and increasing CO2, were input (offline) to the dynamic vegetation model, Hybrid v4.1 (Friend et al., 1997; Friend and White, 1999). This model represents biogeochemical, biophysical and biogeographical processes, coupling the carbon, nitrogen and water cycles on a sub-daily timestep, simulating potential vegetation and transient changes in annual growth and competition between eight generalized plant types in response to climate.Global vegetation carbon was predicted to rise from about 600 to 800 PgC (or to 650 PgC for HadCM3) while the soil carbon pool of about 1100 PgC decreased by about 8%. By the 2080s, climate change caused a partial loss of Amazonian rainforest, C4 grasslands and temperate forest in areas of southern Europe and eastern USA, but an expansion in the boreal forest area. These changes were accompanied by a decrease in net primary productivity (NPP) of vegetation in many tropical areas, southern Europe and eastern USA (in response to warming and a decrease in rainfall), but an increase in NPP of boreal forests. Global NPP increased from 45 to 50 PgC y−1 in the 1990s to about 65 PgC y−1 in the 2080s (about 58 PgC y−1 for HadCM3). Global net ecosystem productivity (NEP) increased from about 1.3 PgC y−1 in the 1990s to about 3.6 PgC y−1 in the 2030s and then declined to zero by 2100 owing to a loss of carbon from declining forests in the tropics and at warm temperate latitudes — despite strengthening of the carbon sink at northern high latitudes. HadCM3 gave a more erratic temporal evolution of NEP than HadCM2, with a dramatic collapse in NEP in the 2050s.
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
Global Environmental Change-human and Policy Dimensions, 1999
Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2... more Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2100, with IS92a greenhouse gas forcing, together with predicted patterns of N deposition and increasing CO2, were input (offline) to the dynamic vegetation model, Hybrid v4.1 (Friend et al., 1997; Friend and White, 1999). This model represents biogeochemical, biophysical and biogeographical processes, coupling the carbon, nitrogen and water cycles on a sub-daily timestep, simulating potential vegetation and transient changes in annual growth and competition between eight generalized plant types in response to climate.Global vegetation carbon was predicted to rise from about 600 to 800 PgC (or to 650 PgC for HadCM3) while the soil carbon pool of about 1100 PgC decreased by about 8%. By the 2080s, climate change caused a partial loss of Amazonian rainforest, C4 grasslands and temperate forest in areas of southern Europe and eastern USA, but an expansion in the boreal forest area. These changes were accompanied by a decrease in net primary productivity (NPP) of vegetation in many tropical areas, southern Europe and eastern USA (in response to warming and a decrease in rainfall), but an increase in NPP of boreal forests. Global NPP increased from 45 to 50 PgC y−1 in the 1990s to about 65 PgC y−1 in the 2080s (about 58 PgC y−1 for HadCM3). Global net ecosystem productivity (NEP) increased from about 1.3 PgC y−1 in the 1990s to about 3.6 PgC y−1 in the 2030s and then declined to zero by 2100 owing to a loss of carbon from declining forests in the tropics and at warm temperate latitudes — despite strengthening of the carbon sink at northern high latitudes. HadCM3 gave a more erratic temporal evolution of NEP than HadCM2, with a dramatic collapse in NEP in the 2050s.
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
This paper reports the main results of an assessment of the global-scale implications of the stab... more This paper reports the main results of an assessment of the global-scale implications of the stabilisation of atmospheric CO2 concentrations at 750 ppm (by 2250) and 550 ppm (by 2150), in relationto a scenario of unmitigated emissions. The climate change scenarios were derived from simulation experiments conducted with the HadCM2 global climate model and forced with the IPCC IS92a, S750 and S550 emissions scenarios. The simulated changes in climate were applied to an observed global baseline climatology, and applied with impacts models to estimate impacts on natural vegetation, water resources, coastal flood risk and wetland loss, crop yield and food security, and malaria. The studies used a single set of population and socio-economic scenarios about the future that are similar to those adopted in the IS92a emissions scenario.An emissions pathway which stabilises CO2 concentrations at 750 ppmby the 2230s delays the 2050 temperature increase under unmitigated emissions by around 50 years. The loss of tropical forest and grassland which occurs by the 2050s under unmitigated emissions is delayed to the 22nd century, and the switch from carbon sink to carbon source is delayed from the 2050s to the 2170s. Coastal wetland loss is slowed. Stabilisation at 750 ppm generally has relatively little effect on the impacts of climate change on water resource stress, and populations at risk of hunger or falciparum malaria until the 2080s.A pathway which stabilises CO2 concentrations at 550 ppm by the 2170s delays the 2050 temperature increase under unmitigated emissions by around 100 years. There is no substantial loss of tropical forest or grassland, even by the 2230s, although the terrestrial carbon store ceases to act as a net carbon sink by around 2170 (this time because the vegetation has reached a new equilibrium with the atmosphere). Coastal wetland loss is slowed considerably, and the increase in coastal flood risk is considerably lower than under unmitigated emissions. CO2 stabilisation at 550 ppm reduces substantially water resource stress, relative to unmitigated emissions, but has relatively little impact on populations at risk of falciparum malaria, and may even cause more people to be at risk of hunger. While this study shows that mitigation avoids many impacts, particularly in the longer-term (beyond the 2080s), stabilisation at 550 ppm appears to be necessary to avoid or significantly reduce most of the projected impacts in the unmitigated case.
This paper reports the main results of an assessment of the global-scale implications of the stab... more This paper reports the main results of an assessment of the global-scale implications of the stabilisation of atmospheric CO2 concentrations at 750 ppm (by 2250) and 550 ppm (by 2150), in relationto a scenario of unmitigated emissions. The climate change scenarios were derived from simulation experiments conducted with the HadCM2 global climate model and forced with the IPCC IS92a, S750 and S550 emissions scenarios. The simulated changes in climate were applied to an observed global baseline climatology, and applied with impacts models to estimate impacts on natural vegetation, water resources, coastal flood risk and wetland loss, crop yield and food security, and malaria. The studies used a single set of population and socio-economic scenarios about the future that are similar to those adopted in the IS92a emissions scenario.An emissions pathway which stabilises CO2 concentrations at 750 ppmby the 2230s delays the 2050 temperature increase under unmitigated emissions by around 50 years. The loss of tropical forest and grassland which occurs by the 2050s under unmitigated emissions is delayed to the 22nd century, and the switch from carbon sink to carbon source is delayed from the 2050s to the 2170s. Coastal wetland loss is slowed. Stabilisation at 750 ppm generally has relatively little effect on the impacts of climate change on water resource stress, and populations at risk of hunger or falciparum malaria until the 2080s.A pathway which stabilises CO2 concentrations at 550 ppm by the 2170s delays the 2050 temperature increase under unmitigated emissions by around 100 years. There is no substantial loss of tropical forest or grassland, even by the 2230s, although the terrestrial carbon store ceases to act as a net carbon sink by around 2170 (this time because the vegetation has reached a new equilibrium with the atmosphere). Coastal wetland loss is slowed considerably, and the increase in coastal flood risk is considerably lower than under unmitigated emissions. CO2 stabilisation at 550 ppm reduces substantially water resource stress, relative to unmitigated emissions, but has relatively little impact on populations at risk of falciparum malaria, and may even cause more people to be at risk of hunger. While this study shows that mitigation avoids many impacts, particularly in the longer-term (beyond the 2080s), stabilisation at 550 ppm appears to be necessary to avoid or significantly reduce most of the projected impacts in the unmitigated case.
Global Environmental Change-human and Policy Dimensions, 1999
Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2... more Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2100, with IS92a greenhouse gas forcing, together with predicted patterns of N deposition and increasing CO2, were input (offline) to the dynamic vegetation model, Hybrid v4.1 (Friend et al., 1997; Friend and White, 1999). This model represents biogeochemical, biophysical and biogeographical processes, coupling the carbon, nitrogen and water cycles on a sub-daily timestep, simulating potential vegetation and transient changes in annual growth and competition between eight generalized plant types in response to climate.Global vegetation carbon was predicted to rise from about 600 to 800 PgC (or to 650 PgC for HadCM3) while the soil carbon pool of about 1100 PgC decreased by about 8%. By the 2080s, climate change caused a partial loss of Amazonian rainforest, C4 grasslands and temperate forest in areas of southern Europe and eastern USA, but an expansion in the boreal forest area. These changes were accompanied by a decrease in net primary productivity (NPP) of vegetation in many tropical areas, southern Europe and eastern USA (in response to warming and a decrease in rainfall), but an increase in NPP of boreal forests. Global NPP increased from 45 to 50 PgC y−1 in the 1990s to about 65 PgC y−1 in the 2080s (about 58 PgC y−1 for HadCM3). Global net ecosystem productivity (NEP) increased from about 1.3 PgC y−1 in the 1990s to about 3.6 PgC y−1 in the 2030s and then declined to zero by 2100 owing to a loss of carbon from declining forests in the tropics and at warm temperate latitudes — despite strengthening of the carbon sink at northern high latitudes. HadCM3 gave a more erratic temporal evolution of NEP than HadCM2, with a dramatic collapse in NEP in the 2050s.
The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate... more The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate change are illustrated using six dynamic global vegetation models that explicitly represent the interactions of ecosystem carbon and water exchanges with vegetation dynamics. The models are driven by the IPCC IS92a scenario of rising CO2 (Wigley et al. 1991), and by climate changes resulting from effective CO2 concentrations corresponding to IS92a, simulated by the coupled ocean atmosphere model HadCM2-SUL. Simulations with changing CO2 alone show a widely distributed terrestrial carbon sink of 1.4–3.8 Pg C y−1 during the 1990s, rising to 3.7–8.6 Pg C y−1 a century later. Simulations including climate change show a reduced sink both today (0.6–3.0 Pg C y−1) and a century later (0.3–6.6 Pg C y−1) as a result of the impacts of climate change on NEP of tropical and southern hemisphere ecosystems. In all models, the rate of increase of NEP begins to level off around 2030 as a consequence of the ‘diminishing return’ of physiological CO2 effects at high CO2 concentrations. Four out of the six models show a further, climate-induced decline in NEP resulting from increased heterotrophic respiration and declining tropical NPP after 2050. Changes in vegetation structure influence the magnitude and spatial pattern of the carbon sink and, in combination with changing climate, also freshwater availability (runoff). It is shown that these changes, once set in motion, would continue to evolve for at least a century even if atmospheric CO2 concentration and climate could be instantaneously stabilized. The results should be considered illustrative in the sense that the choice of CO2 concentration scenario was arbitrary and only one climate model scenario was used. However, the results serve to indicate a range of possible biospheric responses to CO2 and climate change. They reveal major uncertainties about the response of NEP to climate change resulting, primarily, from differences in the way that modelled global NPP responds to a changing climate. The simulations illustrate, however, that the magnitude of possible biospheric influences on the carbon balance requires that this factor is taken into account for future scenarios of atmospheric CO2 and climate change.
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
Global Environmental Change-human and Policy Dimensions, 1999
Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2... more Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2100, with IS92a greenhouse gas forcing, together with predicted patterns of N deposition and increasing CO2, were input (offline) to the dynamic vegetation model, Hybrid v4.1 (Friend et al., 1997; Friend and White, 1999). This model represents biogeochemical, biophysical and biogeographical processes, coupling the carbon, nitrogen and water cycles on a sub-daily timestep, simulating potential vegetation and transient changes in annual growth and competition between eight generalized plant types in response to climate.Global vegetation carbon was predicted to rise from about 600 to 800 PgC (or to 650 PgC for HadCM3) while the soil carbon pool of about 1100 PgC decreased by about 8%. By the 2080s, climate change caused a partial loss of Amazonian rainforest, C4 grasslands and temperate forest in areas of southern Europe and eastern USA, but an expansion in the boreal forest area. These changes were accompanied by a decrease in net primary productivity (NPP) of vegetation in many tropical areas, southern Europe and eastern USA (in response to warming and a decrease in rainfall), but an increase in NPP of boreal forests. Global NPP increased from 45 to 50 PgC y−1 in the 1990s to about 65 PgC y−1 in the 2080s (about 58 PgC y−1 for HadCM3). Global net ecosystem productivity (NEP) increased from about 1.3 PgC y−1 in the 1990s to about 3.6 PgC y−1 in the 2030s and then declined to zero by 2100 owing to a loss of carbon from declining forests in the tropics and at warm temperate latitudes — despite strengthening of the carbon sink at northern high latitudes. HadCM3 gave a more erratic temporal evolution of NEP than HadCM2, with a dramatic collapse in NEP in the 2050s.
The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate... more The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate change are illustrated using six dynamic global vegetation models that explicitly represent the interactions of ecosystem carbon and water exchanges with vegetation dynamics. The models are driven by the IPCC IS92a scenario of rising CO2 (Wigley et al. 1991), and by climate changes resulting from effective CO2 concentrations corresponding to IS92a, simulated by the coupled ocean atmosphere model HadCM2-SUL. Simulations with changing CO2 alone show a widely distributed terrestrial carbon sink of 1.4–3.8 Pg C y−1 during the 1990s, rising to 3.7–8.6 Pg C y−1 a century later. Simulations including climate change show a reduced sink both today (0.6–3.0 Pg C y−1) and a century later (0.3–6.6 Pg C y−1) as a result of the impacts of climate change on NEP of tropical and southern hemisphere ecosystems. In all models, the rate of increase of NEP begins to level off around 2030 as a consequence of the ‘diminishing return’ of physiological CO2 effects at high CO2 concentrations. Four out of the six models show a further, climate-induced decline in NEP resulting from increased heterotrophic respiration and declining tropical NPP after 2050. Changes in vegetation structure influence the magnitude and spatial pattern of the carbon sink and, in combination with changing climate, also freshwater availability (runoff). It is shown that these changes, once set in motion, would continue to evolve for at least a century even if atmospheric CO2 concentration and climate could be instantaneously stabilized. The results should be considered illustrative in the sense that the choice of CO2 concentration scenario was arbitrary and only one climate model scenario was used. However, the results serve to indicate a range of possible biospheric responses to CO2 and climate change. They reveal major uncertainties about the response of NEP to climate change resulting, primarily, from differences in the way that modelled global NPP responds to a changing climate. The simulations illustrate, however, that the magnitude of possible biospheric influences on the carbon balance requires that this factor is taken into account for future scenarios of atmospheric CO2 and climate change.
This paper reports the main results of an assessment of the global-scale implications of the stab... more This paper reports the main results of an assessment of the global-scale implications of the stabilisation of atmospheric CO2 concentrations at 750 ppm (by 2250) and 550 ppm (by 2150), in relationto a scenario of unmitigated emissions. The climate change scenarios were derived from simulation experiments conducted with the HadCM2 global climate model and forced with the IPCC IS92a, S750 and S550 emissions scenarios. The simulated changes in climate were applied to an observed global baseline climatology, and applied with impacts models to estimate impacts on natural vegetation, water resources, coastal flood risk and wetland loss, crop yield and food security, and malaria. The studies used a single set of population and socio-economic scenarios about the future that are similar to those adopted in the IS92a emissions scenario.An emissions pathway which stabilises CO2 concentrations at 750 ppmby the 2230s delays the 2050 temperature increase under unmitigated emissions by around 50 years. The loss of tropical forest and grassland which occurs by the 2050s under unmitigated emissions is delayed to the 22nd century, and the switch from carbon sink to carbon source is delayed from the 2050s to the 2170s. Coastal wetland loss is slowed. Stabilisation at 750 ppm generally has relatively little effect on the impacts of climate change on water resource stress, and populations at risk of hunger or falciparum malaria until the 2080s.A pathway which stabilises CO2 concentrations at 550 ppm by the 2170s delays the 2050 temperature increase under unmitigated emissions by around 100 years. There is no substantial loss of tropical forest or grassland, even by the 2230s, although the terrestrial carbon store ceases to act as a net carbon sink by around 2170 (this time because the vegetation has reached a new equilibrium with the atmosphere). Coastal wetland loss is slowed considerably, and the increase in coastal flood risk is considerably lower than under unmitigated emissions. CO2 stabilisation at 550 ppm reduces substantially water resource stress, relative to unmitigated emissions, but has relatively little impact on populations at risk of falciparum malaria, and may even cause more people to be at risk of hunger. While this study shows that mitigation avoids many impacts, particularly in the longer-term (beyond the 2080s), stabilisation at 550 ppm appears to be necessary to avoid or significantly reduce most of the projected impacts in the unmitigated case.
The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate... more The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate change are illustrated using six dynamic global vegetation models that explicitly represent the interactions of ecosystem carbon and water exchanges with vegetation dynamics. The models are driven by the IPCC IS92a scenario of rising CO2 (Wigley et al. 1991), and by climate changes resulting from effective CO2 concentrations corresponding to IS92a, simulated by the coupled ocean atmosphere model HadCM2-SUL. Simulations with changing CO2 alone show a widely distributed terrestrial carbon sink of 1.4–3.8 Pg C y−1 during the 1990s, rising to 3.7–8.6 Pg C y−1 a century later. Simulations including climate change show a reduced sink both today (0.6–3.0 Pg C y−1) and a century later (0.3–6.6 Pg C y−1) as a result of the impacts of climate change on NEP of tropical and southern hemisphere ecosystems. In all models, the rate of increase of NEP begins to level off around 2030 as a consequence of the ‘diminishing return’ of physiological CO2 effects at high CO2 concentrations. Four out of the six models show a further, climate-induced decline in NEP resulting from increased heterotrophic respiration and declining tropical NPP after 2050. Changes in vegetation structure influence the magnitude and spatial pattern of the carbon sink and, in combination with changing climate, also freshwater availability (runoff). It is shown that these changes, once set in motion, would continue to evolve for at least a century even if atmospheric CO2 concentration and climate could be instantaneously stabilized. The results should be considered illustrative in the sense that the choice of CO2 concentration scenario was arbitrary and only one climate model scenario was used. However, the results serve to indicate a range of possible biospheric responses to CO2 and climate change. They reveal major uncertainties about the response of NEP to climate change resulting, primarily, from differences in the way that modelled global NPP responds to a changing climate. The simulations illustrate, however, that the magnitude of possible biospheric influences on the carbon balance requires that this factor is taken into account for future scenarios of atmospheric CO2 and climate change.
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
Global Environmental Change-human and Policy Dimensions, 1999
Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2... more Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2100, with IS92a greenhouse gas forcing, together with predicted patterns of N deposition and increasing CO2, were input (offline) to the dynamic vegetation model, Hybrid v4.1 (Friend et al., 1997; Friend and White, 1999). This model represents biogeochemical, biophysical and biogeographical processes, coupling the carbon, nitrogen and water cycles on a sub-daily timestep, simulating potential vegetation and transient changes in annual growth and competition between eight generalized plant types in response to climate.Global vegetation carbon was predicted to rise from about 600 to 800 PgC (or to 650 PgC for HadCM3) while the soil carbon pool of about 1100 PgC decreased by about 8%. By the 2080s, climate change caused a partial loss of Amazonian rainforest, C4 grasslands and temperate forest in areas of southern Europe and eastern USA, but an expansion in the boreal forest area. These changes were accompanied by a decrease in net primary productivity (NPP) of vegetation in many tropical areas, southern Europe and eastern USA (in response to warming and a decrease in rainfall), but an increase in NPP of boreal forests. Global NPP increased from 45 to 50 PgC y−1 in the 1990s to about 65 PgC y−1 in the 2080s (about 58 PgC y−1 for HadCM3). Global net ecosystem productivity (NEP) increased from about 1.3 PgC y−1 in the 1990s to about 3.6 PgC y−1 in the 2030s and then declined to zero by 2100 owing to a loss of carbon from declining forests in the tropics and at warm temperate latitudes — despite strengthening of the carbon sink at northern high latitudes. HadCM3 gave a more erratic temporal evolution of NEP than HadCM2, with a dramatic collapse in NEP in the 2050s.
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
Global Environmental Change-human and Policy Dimensions, 1999
Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2... more Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2100, with IS92a greenhouse gas forcing, together with predicted patterns of N deposition and increasing CO2, were input (offline) to the dynamic vegetation model, Hybrid v4.1 (Friend et al., 1997; Friend and White, 1999). This model represents biogeochemical, biophysical and biogeographical processes, coupling the carbon, nitrogen and water cycles on a sub-daily timestep, simulating potential vegetation and transient changes in annual growth and competition between eight generalized plant types in response to climate.Global vegetation carbon was predicted to rise from about 600 to 800 PgC (or to 650 PgC for HadCM3) while the soil carbon pool of about 1100 PgC decreased by about 8%. By the 2080s, climate change caused a partial loss of Amazonian rainforest, C4 grasslands and temperate forest in areas of southern Europe and eastern USA, but an expansion in the boreal forest area. These changes were accompanied by a decrease in net primary productivity (NPP) of vegetation in many tropical areas, southern Europe and eastern USA (in response to warming and a decrease in rainfall), but an increase in NPP of boreal forests. Global NPP increased from 45 to 50 PgC y−1 in the 1990s to about 65 PgC y−1 in the 2080s (about 58 PgC y−1 for HadCM3). Global net ecosystem productivity (NEP) increased from about 1.3 PgC y−1 in the 1990s to about 3.6 PgC y−1 in the 2030s and then declined to zero by 2100 owing to a loss of carbon from declining forests in the tropics and at warm temperate latitudes — despite strengthening of the carbon sink at northern high latitudes. HadCM3 gave a more erratic temporal evolution of NEP than HadCM2, with a dramatic collapse in NEP in the 2050s.
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
This paper reports the main results of an assessment of the global-scale implications of the stab... more This paper reports the main results of an assessment of the global-scale implications of the stabilisation of atmospheric CO2 concentrations at 750 ppm (by 2250) and 550 ppm (by 2150), in relationto a scenario of unmitigated emissions. The climate change scenarios were derived from simulation experiments conducted with the HadCM2 global climate model and forced with the IPCC IS92a, S750 and S550 emissions scenarios. The simulated changes in climate were applied to an observed global baseline climatology, and applied with impacts models to estimate impacts on natural vegetation, water resources, coastal flood risk and wetland loss, crop yield and food security, and malaria. The studies used a single set of population and socio-economic scenarios about the future that are similar to those adopted in the IS92a emissions scenario.An emissions pathway which stabilises CO2 concentrations at 750 ppmby the 2230s delays the 2050 temperature increase under unmitigated emissions by around 50 years. The loss of tropical forest and grassland which occurs by the 2050s under unmitigated emissions is delayed to the 22nd century, and the switch from carbon sink to carbon source is delayed from the 2050s to the 2170s. Coastal wetland loss is slowed. Stabilisation at 750 ppm generally has relatively little effect on the impacts of climate change on water resource stress, and populations at risk of hunger or falciparum malaria until the 2080s.A pathway which stabilises CO2 concentrations at 550 ppm by the 2170s delays the 2050 temperature increase under unmitigated emissions by around 100 years. There is no substantial loss of tropical forest or grassland, even by the 2230s, although the terrestrial carbon store ceases to act as a net carbon sink by around 2170 (this time because the vegetation has reached a new equilibrium with the atmosphere). Coastal wetland loss is slowed considerably, and the increase in coastal flood risk is considerably lower than under unmitigated emissions. CO2 stabilisation at 550 ppm reduces substantially water resource stress, relative to unmitigated emissions, but has relatively little impact on populations at risk of falciparum malaria, and may even cause more people to be at risk of hunger. While this study shows that mitigation avoids many impacts, particularly in the longer-term (beyond the 2080s), stabilisation at 550 ppm appears to be necessary to avoid or significantly reduce most of the projected impacts in the unmitigated case.
This paper reports the main results of an assessment of the global-scale implications of the stab... more This paper reports the main results of an assessment of the global-scale implications of the stabilisation of atmospheric CO2 concentrations at 750 ppm (by 2250) and 550 ppm (by 2150), in relationto a scenario of unmitigated emissions. The climate change scenarios were derived from simulation experiments conducted with the HadCM2 global climate model and forced with the IPCC IS92a, S750 and S550 emissions scenarios. The simulated changes in climate were applied to an observed global baseline climatology, and applied with impacts models to estimate impacts on natural vegetation, water resources, coastal flood risk and wetland loss, crop yield and food security, and malaria. The studies used a single set of population and socio-economic scenarios about the future that are similar to those adopted in the IS92a emissions scenario.An emissions pathway which stabilises CO2 concentrations at 750 ppmby the 2230s delays the 2050 temperature increase under unmitigated emissions by around 50 years. The loss of tropical forest and grassland which occurs by the 2050s under unmitigated emissions is delayed to the 22nd century, and the switch from carbon sink to carbon source is delayed from the 2050s to the 2170s. Coastal wetland loss is slowed. Stabilisation at 750 ppm generally has relatively little effect on the impacts of climate change on water resource stress, and populations at risk of hunger or falciparum malaria until the 2080s.A pathway which stabilises CO2 concentrations at 550 ppm by the 2170s delays the 2050 temperature increase under unmitigated emissions by around 100 years. There is no substantial loss of tropical forest or grassland, even by the 2230s, although the terrestrial carbon store ceases to act as a net carbon sink by around 2170 (this time because the vegetation has reached a new equilibrium with the atmosphere). Coastal wetland loss is slowed considerably, and the increase in coastal flood risk is considerably lower than under unmitigated emissions. CO2 stabilisation at 550 ppm reduces substantially water resource stress, relative to unmitigated emissions, but has relatively little impact on populations at risk of falciparum malaria, and may even cause more people to be at risk of hunger. While this study shows that mitigation avoids many impacts, particularly in the longer-term (beyond the 2080s), stabilisation at 550 ppm appears to be necessary to avoid or significantly reduce most of the projected impacts in the unmitigated case.
Global Environmental Change-human and Policy Dimensions, 1999
Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2... more Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2100, with IS92a greenhouse gas forcing, together with predicted patterns of N deposition and increasing CO2, were input (offline) to the dynamic vegetation model, Hybrid v4.1 (Friend et al., 1997; Friend and White, 1999). This model represents biogeochemical, biophysical and biogeographical processes, coupling the carbon, nitrogen and water cycles on a sub-daily timestep, simulating potential vegetation and transient changes in annual growth and competition between eight generalized plant types in response to climate.Global vegetation carbon was predicted to rise from about 600 to 800 PgC (or to 650 PgC for HadCM3) while the soil carbon pool of about 1100 PgC decreased by about 8%. By the 2080s, climate change caused a partial loss of Amazonian rainforest, C4 grasslands and temperate forest in areas of southern Europe and eastern USA, but an expansion in the boreal forest area. These changes were accompanied by a decrease in net primary productivity (NPP) of vegetation in many tropical areas, southern Europe and eastern USA (in response to warming and a decrease in rainfall), but an increase in NPP of boreal forests. Global NPP increased from 45 to 50 PgC y−1 in the 1990s to about 65 PgC y−1 in the 2080s (about 58 PgC y−1 for HadCM3). Global net ecosystem productivity (NEP) increased from about 1.3 PgC y−1 in the 1990s to about 3.6 PgC y−1 in the 2030s and then declined to zero by 2100 owing to a loss of carbon from declining forests in the tropics and at warm temperate latitudes — despite strengthening of the carbon sink at northern high latitudes. HadCM3 gave a more erratic temporal evolution of NEP than HadCM2, with a dramatic collapse in NEP in the 2050s.
The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate... more The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate change are illustrated using six dynamic global vegetation models that explicitly represent the interactions of ecosystem carbon and water exchanges with vegetation dynamics. The models are driven by the IPCC IS92a scenario of rising CO2 (Wigley et al. 1991), and by climate changes resulting from effective CO2 concentrations corresponding to IS92a, simulated by the coupled ocean atmosphere model HadCM2-SUL. Simulations with changing CO2 alone show a widely distributed terrestrial carbon sink of 1.4–3.8 Pg C y−1 during the 1990s, rising to 3.7–8.6 Pg C y−1 a century later. Simulations including climate change show a reduced sink both today (0.6–3.0 Pg C y−1) and a century later (0.3–6.6 Pg C y−1) as a result of the impacts of climate change on NEP of tropical and southern hemisphere ecosystems. In all models, the rate of increase of NEP begins to level off around 2030 as a consequence of the ‘diminishing return’ of physiological CO2 effects at high CO2 concentrations. Four out of the six models show a further, climate-induced decline in NEP resulting from increased heterotrophic respiration and declining tropical NPP after 2050. Changes in vegetation structure influence the magnitude and spatial pattern of the carbon sink and, in combination with changing climate, also freshwater availability (runoff). It is shown that these changes, once set in motion, would continue to evolve for at least a century even if atmospheric CO2 concentration and climate could be instantaneously stabilized. The results should be considered illustrative in the sense that the choice of CO2 concentration scenario was arbitrary and only one climate model scenario was used. However, the results serve to indicate a range of possible biospheric responses to CO2 and climate change. They reveal major uncertainties about the response of NEP to climate change resulting, primarily, from differences in the way that modelled global NPP responds to a changing climate. The simulations illustrate, however, that the magnitude of possible biospheric influences on the carbon balance requires that this factor is taken into account for future scenarios of atmospheric CO2 and climate change.
... The proteins of tissues have also received attention as a possible source of plasma protein, ... more ... The proteins of tissues have also received attention as a possible source of plasma protein, and Whipple and his colleagues (Madden and Whipple, 1940; Whipple, 1938, 1942; Whipple andMadden, 1944) have suggested the existence of a dynamic equilibrium between blood ...
Global Environmental Change-human and Policy Dimensions, 1999
Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2... more Climate output from the UK Hadley Centre's HadCM2 and HadCM3 experiments for the period 1860 to 2100, with IS92a greenhouse gas forcing, together with predicted patterns of N deposition and increasing CO2, were input (offline) to the dynamic vegetation model, Hybrid v4.1 (Friend et al., 1997; Friend and White, 1999). This model represents biogeochemical, biophysical and biogeographical processes, coupling the carbon, nitrogen and water cycles on a sub-daily timestep, simulating potential vegetation and transient changes in annual growth and competition between eight generalized plant types in response to climate.Global vegetation carbon was predicted to rise from about 600 to 800 PgC (or to 650 PgC for HadCM3) while the soil carbon pool of about 1100 PgC decreased by about 8%. By the 2080s, climate change caused a partial loss of Amazonian rainforest, C4 grasslands and temperate forest in areas of southern Europe and eastern USA, but an expansion in the boreal forest area. These changes were accompanied by a decrease in net primary productivity (NPP) of vegetation in many tropical areas, southern Europe and eastern USA (in response to warming and a decrease in rainfall), but an increase in NPP of boreal forests. Global NPP increased from 45 to 50 PgC y−1 in the 1990s to about 65 PgC y−1 in the 2080s (about 58 PgC y−1 for HadCM3). Global net ecosystem productivity (NEP) increased from about 1.3 PgC y−1 in the 1990s to about 3.6 PgC y−1 in the 2030s and then declined to zero by 2100 owing to a loss of carbon from declining forests in the tropics and at warm temperate latitudes — despite strengthening of the carbon sink at northern high latitudes. HadCM3 gave a more erratic temporal evolution of NEP than HadCM2, with a dramatic collapse in NEP in the 2050s.
The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate... more The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate change are illustrated using six dynamic global vegetation models that explicitly represent the interactions of ecosystem carbon and water exchanges with vegetation dynamics. The models are driven by the IPCC IS92a scenario of rising CO2 (Wigley et al. 1991), and by climate changes resulting from effective CO2 concentrations corresponding to IS92a, simulated by the coupled ocean atmosphere model HadCM2-SUL. Simulations with changing CO2 alone show a widely distributed terrestrial carbon sink of 1.4–3.8 Pg C y−1 during the 1990s, rising to 3.7–8.6 Pg C y−1 a century later. Simulations including climate change show a reduced sink both today (0.6–3.0 Pg C y−1) and a century later (0.3–6.6 Pg C y−1) as a result of the impacts of climate change on NEP of tropical and southern hemisphere ecosystems. In all models, the rate of increase of NEP begins to level off around 2030 as a consequence of the ‘diminishing return’ of physiological CO2 effects at high CO2 concentrations. Four out of the six models show a further, climate-induced decline in NEP resulting from increased heterotrophic respiration and declining tropical NPP after 2050. Changes in vegetation structure influence the magnitude and spatial pattern of the carbon sink and, in combination with changing climate, also freshwater availability (runoff). It is shown that these changes, once set in motion, would continue to evolve for at least a century even if atmospheric CO2 concentration and climate could be instantaneously stabilized. The results should be considered illustrative in the sense that the choice of CO2 concentration scenario was arbitrary and only one climate model scenario was used. However, the results serve to indicate a range of possible biospheric responses to CO2 and climate change. They reveal major uncertainties about the response of NEP to climate change resulting, primarily, from differences in the way that modelled global NPP responds to a changing climate. The simulations illustrate, however, that the magnitude of possible biospheric influences on the carbon balance requires that this factor is taken into account for future scenarios of atmospheric CO2 and climate change.
This paper reports the main results of an assessment of the global-scale implications of the stab... more This paper reports the main results of an assessment of the global-scale implications of the stabilisation of atmospheric CO2 concentrations at 750 ppm (by 2250) and 550 ppm (by 2150), in relationto a scenario of unmitigated emissions. The climate change scenarios were derived from simulation experiments conducted with the HadCM2 global climate model and forced with the IPCC IS92a, S750 and S550 emissions scenarios. The simulated changes in climate were applied to an observed global baseline climatology, and applied with impacts models to estimate impacts on natural vegetation, water resources, coastal flood risk and wetland loss, crop yield and food security, and malaria. The studies used a single set of population and socio-economic scenarios about the future that are similar to those adopted in the IS92a emissions scenario.An emissions pathway which stabilises CO2 concentrations at 750 ppmby the 2230s delays the 2050 temperature increase under unmitigated emissions by around 50 years. The loss of tropical forest and grassland which occurs by the 2050s under unmitigated emissions is delayed to the 22nd century, and the switch from carbon sink to carbon source is delayed from the 2050s to the 2170s. Coastal wetland loss is slowed. Stabilisation at 750 ppm generally has relatively little effect on the impacts of climate change on water resource stress, and populations at risk of hunger or falciparum malaria until the 2080s.A pathway which stabilises CO2 concentrations at 550 ppm by the 2170s delays the 2050 temperature increase under unmitigated emissions by around 100 years. There is no substantial loss of tropical forest or grassland, even by the 2230s, although the terrestrial carbon store ceases to act as a net carbon sink by around 2170 (this time because the vegetation has reached a new equilibrium with the atmosphere). Coastal wetland loss is slowed considerably, and the increase in coastal flood risk is considerably lower than under unmitigated emissions. CO2 stabilisation at 550 ppm reduces substantially water resource stress, relative to unmitigated emissions, but has relatively little impact on populations at risk of falciparum malaria, and may even cause more people to be at risk of hunger. While this study shows that mitigation avoids many impacts, particularly in the longer-term (beyond the 2080s), stabilisation at 550 ppm appears to be necessary to avoid or significantly reduce most of the projected impacts in the unmitigated case.
The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate... more The possible responses of ecosystem processes to rising atmospheric CO2 concentration and climate change are illustrated using six dynamic global vegetation models that explicitly represent the interactions of ecosystem carbon and water exchanges with vegetation dynamics. The models are driven by the IPCC IS92a scenario of rising CO2 (Wigley et al. 1991), and by climate changes resulting from effective CO2 concentrations corresponding to IS92a, simulated by the coupled ocean atmosphere model HadCM2-SUL. Simulations with changing CO2 alone show a widely distributed terrestrial carbon sink of 1.4–3.8 Pg C y−1 during the 1990s, rising to 3.7–8.6 Pg C y−1 a century later. Simulations including climate change show a reduced sink both today (0.6–3.0 Pg C y−1) and a century later (0.3–6.6 Pg C y−1) as a result of the impacts of climate change on NEP of tropical and southern hemisphere ecosystems. In all models, the rate of increase of NEP begins to level off around 2030 as a consequence of the ‘diminishing return’ of physiological CO2 effects at high CO2 concentrations. Four out of the six models show a further, climate-induced decline in NEP resulting from increased heterotrophic respiration and declining tropical NPP after 2050. Changes in vegetation structure influence the magnitude and spatial pattern of the carbon sink and, in combination with changing climate, also freshwater availability (runoff). It is shown that these changes, once set in motion, would continue to evolve for at least a century even if atmospheric CO2 concentration and climate could be instantaneously stabilized. The results should be considered illustrative in the sense that the choice of CO2 concentration scenario was arbitrary and only one climate model scenario was used. However, the results serve to indicate a range of possible biospheric responses to CO2 and climate change. They reveal major uncertainties about the response of NEP to climate change resulting, primarily, from differences in the way that modelled global NPP responds to a changing climate. The simulations illustrate, however, that the magnitude of possible biospheric influences on the carbon balance requires that this factor is taken into account for future scenarios of atmospheric CO2 and climate change.
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